It is all-important to remember that naturalists have no golden rule by which to distinguish species and varieties; they grant some little variability to each species, but, when they meet with a somewhat greater amount of difference between any two forms, they rank both as species, unless they are enabled to connect them together by the closest intermediate gradations; and this, from the reasons just assigned, we can seldom hope to effect in any one geological section. Supposing B and C to be two species, and a third, A, to be found in an older and underlying bed; even if A were strictly intermediate between B and C, it would simply be ranked as a third and distinct species, unless at the same time it could be closely connected by intermediate varieties with either one or both forms. Nor should it be forgotten, as before explained, that A might be the actual progenitor of B and C, and yet would not necessarily be strictly intermediate between them in all respects. So that we might obtain the parent-species and its several modified descendants from the lower and upper beds of the same formation, and, unless we obtained numerous transitional gradations, we should not recognize their blood-relationship, and should consequently rank them as distinct species.
SUDDEN APPEARANCE OF GROUPS OF ALLIED SPECIES.
page 282.
The abrupt manner in which whole groups of species suddenly appear in certain formations has been urged by several paleontologists—for instance, by Agassiz, Pictet, and Sedgwick—as a fatal objection to the belief in the transmutation of species. If numerous species, belonging to the same genera or families, have really started into life at once, the fact would be fatal to the theory of evolution through natural selection. For the development by this means of a group of forms, all of which are descended from some one progenitor, must have been an extremely slow process; and the progenitors must have lived long before their modified descendants. But we continually overrate the perfection of the geological record, and falsely infer, because certain genera or families have not been found beneath a certain stage, that they did not exist before that stage. In all cases positive paleontological evidence may be implicitly trusted; negative evidence is worthless, as experience has so often shown. We continually forget how large the world is, compared with the area over which our geological formations have been carefully examined; we forget that groups of species may elsewhere have long existed, and have slowly multiplied, before they invaded the ancient archipelagoes of Europe and the United States. We do not make due allowance for the intervals of time which have elapsed between our consecutive formations—longer, perhaps, in many cases than the time required for the accumulation of each formation. These intervals will have given time for the multiplication of species from some one parent-form; and, in the succeeding formation, such groups or species will appear as if suddenly created.
HOW LITTLE WE KNOW OF FORMER INHABITANTS OF THE WORLD.
Even in so short an interval as that between the first and second edition of Pictet’s great work on Paleontology, published in 1844–’46 and in 1853–’57, the conclusions on the first appearance and disappearance of several groups of animals have been considerably modified; and a third edition would require still further changes. I may recall the well-known fact that in geological treatises, published not many years ago, mammals were always spoken of as having abruptly come in at the commencement of the tertiaryB series. And now one of the richest known accumulations of fossil mammals belongs to the middle of the secondary series; and true mammals have been discovered in the new red sandstone at nearly the commencement of this great series. Cuvier used to urge that no monkey occurred in any tertiary stratum; but now extinct species have been discovered in India, South America, and in Europe, as far back as the Miocene stage. Had it not been for the rare accident of the preservation of footsteps in the new red sandstone of the United States, who would have ventured to suppose that no less than at least thirty different bird-like animals, some of gigantic size, existed during that period? Not a fragment of bone has been discovered in these beds. Not long ago, paleontologists maintained that the whole class of birds came suddenly into existence during the Eocene period; but now we know, on the authority of Professor Owen, that a bird certainly lived during the deposition of the upper greensand; and still more recently, that strange bird, the archeopteryx, with a long, lizard-like tail, bearing a pair of feathers on each joint, and with its wings furnished with two free claws, has been discovered in the oölitic slates of Solenhofen. Hardly any recent discovery shows more forcibly than this, how little we as yet know of the former inhabitants of the world.
B Tertiary.—The latest geological epoch, immediately preceding the establishment of the present order of things.
THE EXTINCTION OF SPECIES INVOLVED IN MYSTERY.
page 294.
The extinction of species has been involved in the most gratuitous mystery. Some authors have even supposed that as the individual has a definite length of life, so have species a definite duration. No one can have marveled more than I have done at the extinction of species. When I found in La Plata the tooth of a horse imbedded with the remains of mastodon, megatherium, toxodon, and other extinct monsters, which all co-existed with still living shells at a very late geological period, I was filled with astonishment; for, seeing that the horse, since its introduction by the Spaniards into South America, has run wild over the whole country and has increased in numbers at an unparalleled rate, I asked myself what could so recently have exterminated the former horse under conditions of life apparently so favorable. But my astonishment was groundless. Professor Owen soon perceived that the tooth, though so like that of the existing horse, belonged to an extinct species. Had this horse been still living, but in some degree rare, no naturalist would have felt the least surprise at its rarity; for rarity is the attribute of a vast number of species of all classes, in all countries. If we ask ourselves why this or that species is rare, we answer that something is unfavorable in its conditions of life; but what that something is we can hardly ever tell. On the supposition of the fossil horse still existing as a rare species, we might have felt certain, from the analogy of all other mammals, even of the slow-breeding elephant, and from the history of the naturalization of the domestic horse in South America, that under more favorable conditions it would in a very few years have stocked the whole continent. But we could not have told what the unfavorable conditions were which checked its increase, whether some one or several contingencies, and at what period of the horse’s life, and in what degree, they severally acted. If the conditions had gone on, however slowly, becoming less and less favorable, we assuredly should not have perceived the fact, yet the fossil horse would certainly have become rarer and rarer, and finally extinct;—its place being seized on by some more successful competitor.
It is most difficult always to remember that the increase of every creature is constantly being checked by unperceived hostile agencies; and that these same unperceived agencies are amply sufficient to cause rarity, and finally extinction. So little is this subject understood that I have heard surprise repeatedly expressed at such great monsters as the mastodon and the more ancient dinosaurians having become extinct; as if mere bodily strength gave victory in the battle of life. Mere size, on the contrary, would in some cases determine, as has been remarked by Owen, quicker extermination from the greater amount of requisite food. Before man inhabited India or Africa, some cause must have checked the continued increase of the existing elephant. A highly capable judge, Dr. Falconer, believes that it is chiefly insects which, from incessantly harassing and weakening the elephant in India, check its increase; and this was Bruce’s conclusion with respect to the African elephant in Abyssinia. It is certain that insects and blood-sucking bats determine the existence of the larger naturalized quadrupeds in several parts of South America.
I may repeat what I published in 1845, namely, that to admit that species generally become rare before they become extinct—to feel no surprise at the rarity of a species, and yet to marvel greatly when the species ceases to exist, is much the same as to admit that sickness in the individual is the forerunner of death—to feel no surprise at sickness, but, when the sick man dies, to wonder and to suspect that he died by some deed of violence.
DEAD LINKS BETWEEN LIVING SPECIES.
No one will deny that the Hipparion is intermediate between the existing horse and certain older ungulate forms. What a wonderful connecting link in the chain of mammals is the Typotherium from South America, as the name given to it by Professor Gervais expresses, and which can not be placed in any existing order! The Sirenia form a very distinct group of mammals, and one of the most remarkable peculiarities in the existing dugong and lamentin is the entire absence of hind limbs, without even a rudiment being left; but the extinct Halitherium had, according to Professor Flower, an ossified thigh-bone “articulated to a well-defined acetabulum in the pelvis,” and it thus makes some approach to ordinary hoofed quadrupeds, to which the Sirenia are in other respects allied. The cetaceans or whales are widely different from all other mammals, but the tertiary Zeuglodon and Squalodon, which have been placed by some naturalists in an order by themselves, are considered by Professor Huxley to be undoubtedly cetaceans, “and to constitute connecting links with the aquatic carnivora.”
Even the wide interval between birds and reptiles has been shown by the naturalist just quoted to be partially bridged over in the most unexpected manner, on the one hand, by the ostrich and extinct Archeopteryx, and on the other hand, by the Compsognathus, one of the dinosaurians—that group which includes the most gigantic of all terrestrial reptiles. Turning to the Invertebrata, Barrande asserts, and a higher authority could not be named, that he is every day taught that, although palæozoic animals can certainly be classed under existing groups, yet that at this ancient period the groups were not so distinctly separated from each other as they now are.
Some writers have objected to any extinct species, or group of species, being considered as intermediate between any two living species or groups of species. If by this term it is meant that an extinct form is directly intermediate in all its characters between two living forms or groups, the objection is probably valid. But in a natural classification many fossil species certainly stand between living species, and some extinct genera between living genera, even between genera belonging to distinct families. The most common case, especially with respect to very distinct groups, such as fish and reptiles, seems to be that, supposing them to be distinguished at the present day by a score of characters, the ancient members are separated by a somewhat lesser number of characters; so that the two groups formerly made a somewhat nearer approach to each other than they now do.
LIVING DESCENDANTS OF FOSSIL SPECIES.
It may be asked in ridicule, whether I suppose that the megatherium and other allied huge monsters, which formerly lived in South America, have left behind them the sloth, armadillo, and ant-eater, as their degenerate descendants. This can not for an instant be admitted. These huge animals have become wholly extinct, and have left no progeny. But in the caves of Brazil there are many extinct species which are closely allied in size and in all other characters to the species still living in South America; and some of these fossils may have been the actual progenitors of the living species. It must not be forgotten that, on our theory, all the species of the same genus are the descendants of some one species; so that, if six genera, each having eight species, be found in one geological formation, and in a succeeding formation there be six other allied or representative genera each with the same number of species, then we may conclude that generally only one species of each of the older genera has left modified descendants, which constitute the new genera containing the several species; the other seven species of each old genus having died out and left no progeny. Or, and this will be a far commoner case, two or three species in two or three alone of the six older genera will be the parents of the new genera: the other species and the other whole genera having become utterly extinct. In failing orders, with the genera and species decreasing in numbers as is the case with the Edentata of South America, still fewer genera and species will leave modified blood-descendants.
UNNECESSARY TO EXPLAIN THE CAUSE OF EACH INDIVIDUAL DIFFERENCE.
vol. ii, page 425.
In accordance with the views maintained by me in this work and elsewhere, not only various domestic races, but the most distinct genera and orders within the same great class—for instance, mammals, birds, reptiles, and fishes—are all the descendants of one common progenitor, and we must admit that the whole vast amount of difference between these forms has primarily arisen from simple variability. To consider the subject under this point of view is enough to strike one dumb with amazement. But our amazement ought to be lessened when we reflect that beings almost infinite in number, during an almost infinite lapse of time, have often had their whole organization rendered in some degree plastic, and that each slight modification of structure which was in any way beneficial under excessively complex conditions of life has been preserved, while each which was in any way injurious has been rigorously destroyed. And the long-continued accumulation of beneficial variations will infallibly have led to structures as diversified, as beautifully adapted for various purposes and as excellently co-ordinated, as we see in the animals and plants around us. Hence I have spoken of selection as the paramount power, whether applied by man to the formation of domestic breeds, or by nature to the production of species.
If an architect were to rear a noble and commodious edifice, without the use of cut stone, by selecting from the fragments at the base of a precipice wedge-formed stones for his arches, elongated stones for his lintels, and flat stones for his roof, we should admire his skill and regard him as the paramount power. Now, the fragments of stone, though indispensable to the architect, bear to the edifice built by him the same relation which the fluctuating variations of organic beings bear to the varied and admirable structures ultimately acquired by their modified descendants.
Some authors have declared that natural selection explains nothing, unless the precise cause of each slight individual difference be made clear. If it were explained to a savage utterly ignorant of the art of building, how the edifice had been raised stone upon stone, and why wedge-formed fragments were used for the arches, flat stones for the roof, etc., and if the use of each part and of the whole building were pointed out, it would be unreasonable if he declared that nothing had been made clear to him, because the precise cause of the shape of each fragment could not be told. But this is a nearly parallel case with the objection that selection explains nothing, because we know not the cause of each individual difference in the structure of each being.
The shape of the fragments of stone at the base of our precipice may be called accidental, but this is not strictly correct; for the shape of each depends on a long sequence of events, all obeying natural laws; on the nature of the rock, on the lines of deposition or cleavage, on the form of the mountain, which depends on its upheaval and subsequent denudation, and lastly on the storm or earthquake which throws down the fragments. But in regard to the use to which the fragments may be put, their shape may be strictly said to be accidental.
“FACE TO FACE WITH AN INSOLUBLE DIFFICULTY.”
And here we are led to face a great difficulty, in alluding to which I am aware that I am traveling beyond my proper province. An omniscient Creator must have foreseen every consequence which results from the laws imposed by him. But can it be reasonably maintained that the Creator intentionally ordered, if we use the words in any ordinary sense, that certain fragments of rock should assume certain shapes so that the builder might erect his edifice? If the various laws which have determined the shape of each fragment were not predetermined for the builder’s sake, can it be maintained with any greater probability that he specially ordained for the sake of the breeder each of the innumerable variations in our domestic animals and plants—many of these variations being of no service to man, and not beneficial, far more often injurious, to the creatures themselves? Did he ordain that the crop and tail-feathers of the pigeon should vary in order that the fancier might make his grotesque pouter and fantail breeds? Did he cause the frame and mental qualities of the dog to vary in order that a breed might be formed of indomitable ferocity, with jaws fitted to pin down the bull for man’s brutal sport? But if we give up the principle in one case—if we do not admit that the variations of the primeval dog were intentionally guided in order that the greyhound, for instance, that perfect image of symmetry and vigor, might be formed—no shadow of reason can be assigned for the belief that variations, alike in nature and the result of the same general laws, which have been the groundwork through natural selection of the formation of the most perfectly adapted animals in the world, man included, were intentionally and specially guided. However much we may wish it, we can hardly follow Professor Asa Gray in his belief “that variation has been led along certain beneficial lines,” like a stream “along definite and useful lines of irrigation.” If we assume that each particular variation was from the beginning of all time preordained, then that plasticity of organization, which leads to many injurious deviations of structure, as well as the redundant power of reproduction which inevitably leads to a struggle for existence, and, as a consequence, to the natural selection or survival of the fittest, must appear to us superfluous laws of nature. On the other hand, an omnipotent and omniscient Creator ordains everything and foresees everything. Thus we are brought face to face with a difficulty as insoluble as is that of free-will and predestination.
WHY DISTASTEFUL?
page 618.
The main conclusion arrived at in this work, namely, that man is descended from some lowly organized form, will, I regret to think, be highly distasteful to many. But there can hardly be a doubt that we are descended from barbarians. The astonishment which I felt on first seeing a party of Fuegians on a wild and broken shore will never be forgotten by me, for the reflection at once rushed into my mind—such were our ancestors. These men were absolutely naked and bedaubed with paint, their long hair was tangled, their mouths frothed with excitement, and their expression was wild, startled, and distrustful. They possessed hardly any arts, and like wild animals lived on what they could catch; they had no government, and were merciless to every one not of their own small tribe. He who has seen a savage in his native land will not feel much shame, if forced to acknowledge that the blood of some more humble creature flows in his veins. For my own part I would as soon be descended from that heroic little monkey, who braved his dreaded enemy in order to save the life of his keeper, or from that old baboon, who, descending from the mountains, carried away in triumph his young comrade from a crowd of astonished dogs—as from a savage who delights to torture his enemies, offers up bloody sacrifices, practices infanticide without remorse, treats his wives like slaves, knows no decency, and is haunted by the grossest superstitions.
Man may be excused for feeling some pride at having risen, though not through his own exertions, to the very summit of the organic scale; and the fact of his having thus risen, instead of having been aboriginally placed there, may give him hope for a still higher destiny in the distant future. But we are not here concerned with hopes or fears, only with the truth as far as our reason permits us to discover it; and I have given the evidence to the best of my ability. We must, however, acknowledge, as it seems to me, that man with all his noble qualities, with sympathy which feels for the most debased, with benevolence which extends not only to other men but to the humblest living creature, with his godlike intellect which has penetrated into the movements and constitution of the solar system—with all these exalted powers—man still bears in his bodily frame the indelible stamp of his lowly origin.
“ACCORDS BETTER WITH WHAT WE KNOW OF THE CREATOR’S LAWS.”
page 428.
Authors of the highest eminence seem to be fully satisfied with the view that each species has been independently created. To my mind it accords better with what we know of the laws impressed on matter by the Creator, that the production and extinction of the past and present inhabitants of the world should have been due to secondary causes, like those determining the birth and death of the individual. When I view all beings not as special creations, but as the lineal descendants of some few beings which lived long before the first bed of the Cambrian system was deposited, they seem to me to become ennobled. Judging from the past, we may safely infer that not one living species will transmit its unaltered likeness to a distant futurity. And of the species now living very few will transmit progeny of any kind to a far distant futurity; for the manner in which all organic beings are grouped, shows that the greater number of species in each genus, and all the species in many genera, have left no descendants, but have become utterly extinct. We can so far take a prophetic glance into futurity as to foretell that it will be the common and widely-spread species, belonging to the larger and dominant groups within each class, which will ultimately prevail and procreate new and dominant species. As all the living forms of life are the lineal descendants of those which lived long before the Cambrian epoch, we may feel certain that the ordinary succession by generation has never once been broken, and that no cataclysm has desolated the whole world. Hence we may look with some confidence to a secure future of great length. And as natural selection works solely by and for the good of each being, all corporeal and mental endowments will tend to progress toward perfection.
THE GRANDEUR OF THIS VIEW OF LIFE.
It is interesting to contemplate a tangled bank, clothed with many plants of many kinds, with birds singing on the bushes, with various insects flitting about, and with worms crawling through the damp earth, and to reflect that these elaborately constructed forms, so different from each other, and dependent upon each other in so complex a manner, have all been produced by laws acting around us. These laws, taken in the largest sense, being growth with reproduction; inheritance which is almost implied by reproduction; variability from the indirect and direct action of the conditions of life, and from use and disuse: a ratio of increase so high as to lead to a struggle for life, and as a consequence to natural selection, entailing divergence of character and the extinction of less-improved forms. Thus, from the war of nature, from famine and death, the most exalted object which we are capable of conceiving, namely, the production of the higher animals, directly follows. There is grandeur in this view of life, with its several powers, having been originally breathed by the Creator into a few forms or into one; and that, while this planet has gone cycling on according to the fixed law of gravity, from so simple a beginning endless forms most beautiful and most wonderful have been and are being evolved.
NOT INCOMPATIBLE WITH THE BELIEF IN IMMORTALITY.
page 612.
I am aware that the assumed instinctive belief in God has been used by many persons as a rash argument for his existence. But this is a rash argument, as we should thus be compelled to believe in the existence of many cruel and malignant spirits, only a little more powerful than man; for the belief in them is far more general than in a beneficent Deity. The idea of a universal and beneficent Creator does not seem to arise in the mind of man, until he has been elevated by long-continued culture.
He who believes in the advancement of man from some low organized form, will naturally ask, How does this bear on the belief in the immortality of the soul? The barbarous races of man, as Sir J. Lubbock has shown, possess no clear belief of this kind; but arguments derived from the primeval beliefs of savages are, as we have just seen, of little or no avail. Few persons feel any anxiety from the impossibility of determining at what precise period in the development of the individual, from the first trace of a minute germinal vesicle, man becomes an immortal being; and there is no greater cause for anxiety because the period can not possibly be determined in the gradually ascending organic scale.
I am aware that the conclusions arrived at in this work will be denounced by some as highly irreligious; but he who denounces them is bound to show why it is more irreligious to explain the origin of man as a distinct species by descent from some lower form, through the laws of variation and natural selection, than to explain the birth of the individual through the laws of ordinary reproduction. The birth both of the species and of the individual are equally parts of that grand sequence of events, which our minds refuse to accept as the result of blind chance. The understanding revolts at such a conclusion, whether or not we are able to believe that every slight variation of structure—the union of each pair in marriage—the dissemination of each seed—and other such events, have all been ordained for some special purpose.
page 503.
Among the scenes which are deeply impressed on my mind, none exceed in sublimity the primeval forests undefaced by the hand of man; whether those of Brazil, where the powers of life are predominant, or those of Tierra del Fuego, where death and decay prevail. Both are temples filled with the varied productions of the God of Nature; no one can stand in these solitudes unmoved, and not feel that there is more in man than the mere breath of his body.
THE END.