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Fossil plants, Vol. 2

Chapter 2: PREFACE
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About This Book

A comprehensive palaeobotanical textbook surveying fossil representatives of non-flowering plant groups, with detailed treatments of Sphenophyllales, Psilotales, Lycopodiales and arborescent lycopods. It compares living and fossil morphology and anatomy, illustrates stem and reproductive structures, and discusses taxonomic distinctions such as Lepidodendron and related genera. Chapters integrate microscopic sections, casts, and impressions, and consider palaeogeographic distribution and recent research developments. Numerous drawings and plates support descriptions and anatomical interpretations for students of botany and geology.

PREFACE

I REGRET that pressure of other work has prevented the completion of this Volume within a reasonable time since the publication of Volume I. Had Volume II been written ten years ago, the discoveries made in the course of the last decade would have given an out-of-date character to much of the subject-matter. It is more especially in regard to the Ferns and the extinct members of the Gymnosperms that our outlook has been materially altered by recent contributions to Palaeobotany. It is, however, some satisfaction to be able to add that recent progress has been relatively slight in that part of the subject dealt with in the first volume.

The original intention was to complete the whole work in two volumes. Soon after the second volume was begun, it became evident that the remaining divisions of the plant-kingdom could not be included within the compass of a single volume. I decided, therefore, to take the consequences of having embarked on too ambitious a plan of treatment, and to preserve uniformity of proportion by reserving the seed-bearing plants for a third volume. The third volume will include the Pteridosperms, other than those briefly described in the final chapter of the present volume, and other classes of Gymnosperms. I propose also to devote such space as is available within the limits of a text-book to the neglected subject of the geographical distribution of plants at different stages in the history of the earth. It is my intention to complete Volume III with as little delay as possible. As I have written elsewhere, the past history of the Flowering plants needs special treatment, and anything more than a mere compilation can be adequately attempted only after considerable research and with the assistance of botanists possessing a special knowledge of different families of Angiosperms. The need of a critical examination of available data in regard to the geological history of this dominant group will not be lost sight of.

I am well aware that while certain genera have received an undue share of attention in the present volume, others have been ignored or treated with scant consideration. For this inconsistency I have no excuse to offer, beyond the statement that the subject is a large one, and selection is necessary even though the work consists of three volumes.

The publication in 1909 of a collection of excellent photographs of Palaeozoic Plants, with brief descriptive notes, by Mr Newell Arber, as one of a series of popular “Nature Books,” bears striking testimony to the remarkable spread of interest in the study of the vegetation of the past, which is one of the outstanding features in the recent history of botanical science.

In the list of illustrations I have mentioned the source of all figures which have been previously published. I would, however, supplement the statement of fact with an expression of thanks to corporate bodies and to individuals who have allowed me to make use of blocks, drawings, or photographs.

I wish to thank my colleague, Mr A. G. Tansley, for placing at my disposal several blocks originally published in the pages of the New Phytologist. To Professor Bertrand of Lille and to his son Dr Paul Bertrand I am indebted for several prints and descriptive notes of specimens in their possession. My friends Dr Nathorst of Stockholm and Dr Zeiller of Paris have generously responded to my requests for information on various points. I wish especially to thank Dr Kidston for several excellent prints of specimens in his collection and for the loan of sections. I have profited by more than one examination of his splendid collection at Stirling. Professor Weiss has generously allowed me to borrow sections from the Manchester University collections, more especially several which have been reproduced in the chapter devoted to the genus Lepidodendron. To Professor F. W. Oliver my thanks are due for the loan of sections from the collection under his charge at University College. I have pleasure also in thanking Dr Scott, not only for lending me sections of a Lepidodendron and for allowing me to use some drawings of Miadesmia originally made by Mrs Scott for reproduction in his invaluable book, Studies in Fossil Botany, but for kindly undertaking the laborious task of reading the proofs of this volume. It would be unfair to express my gratitude to Dr Scott for many helpful suggestions and criticisms, without explicitly stating that thanks to a friend for reading proofs must not be interpreted as an attempt to claim his support for all statements or views expressed. The General Editor of the Series, Mr A. E. Shipley, has also kindly read the proofs. I am under obligations also for assistance of various kinds to Prof. Thomas of Auckland, New Zealand, to Mr Boodle of Kew, to Mr D. M. S. Watson of Manchester, to Mr T. G. Hill of University College, and to Mr Gordon of Emmanuel College, Cambridge. I am indebted to the kind offices of Miss M. C. Knowles for the photograph of the specimen of Archaeopteris hibernica in the Irish National Museum, Dublin, reproduced on page 561.

Many of the illustrations are reproduced from drawings by my wife: those made from the actual specimens are distinguished by the addition of the initials M. S. I am grateful to her also for some improvements in the letter-press. For the drawings made from sections and for some of the outline sketches I am responsible. I have availed myself freely of the facilities afforded by Professor McKenny Hughes in the Sedgwick Museum of Geology for the examination of specimens under the charge of Mr Newell Arber, the University Demonstrator in Palaeobotany. It is a pleasure to add that, as on former occasions, I am indebted to the vigilance of the Readers of the University Press for the detection of several errors which escaped my notice in the revision of the proofs.

A. C. SEWARD.
Botany School, Cambridge.
March 12, 1910.

TABLE OF CONTENTS

———————
SPHENOPHYLLALES (continued from Volume I.).  Pp. 1–16.
 
PAGE
Sphenophyllum
1–7
Cheirostrobus
7–12
Sphenophyllales and Psilotaceae
12–16
PSILOTALES.  Pp. 17–29.
Psilotum and Tmesipteris
17–24
Fossils described by authors as being closely allied to Psilotum
24–26
Psilophyton
26–29
LYCOPODIALES.  Pp. 30–91.
Recent Lycopodiales (General)
30–33
Lycopodiaceae (Recent)
33–49
Selaginellaceae (Recent)
49–58
Isoetaceae (Recent)
58–66
Fossil Lycopodiales
66–91
Isoetaceae (Fossil)
66–68
Pleuromeia
68–73
Herbaceous fossil species of Lycopodiales
73–91
Lycopodites
76–84
Selaginellites
85–88
Lycostrobus
88–91
Poecilitostachys
91
ARBORESCENT LYCOPODIALES.  Pp. 92–195.
Lepidodendron
93–181
i.
General
93–97
ii.
Leaves and Leaf-cushions
97–105
iii.
Lepidophloios
105–109
iv.
The anatomy of Lepidodendron vasculare
109–123
v.
Lepidodendron stems as represented by casts and impressions of partially decorticated specimens
123–128
  a. Knorria 124–126; b. Bergeria 126, 127; c. Aspidiaria 127, 128.  
vi.
Lepidodendroid axes known as Ulodendron and Halonia
128–139
  a. Ulodendron 128–135; b. Halonia 135–139.  
vii.
Anatomical characters of vegetative Lepidodendron shoots
139–181
  1. Lepidodendron esnostense 139, 140; 2. L. rhodumnense 140; 3. L. saalfeldense 141; 4. L. fuliginosum 141–160; 5. L. Harcourtii 160–163; 6. L. Wünschianum 163–171; 7. L. macrophyllum 171; 8. L. Veltheimianum 171–177; 9. L. Pedroanum 177, 178; 10. L. australe 178–181.  
viii.
Fertile shoots of Lepidodendreae
181–195
  A. Lepidostrobus 181–191. i. Lepidostrobus variabilis 187, 188; ii. L. oldhamius 188–190; iii. L. Brownii, etc. 190, 191.  
  B. Spencerites 192–195.  
SIGILLARIA.  Pp. 196–226.
i.
General 196–210; ii. Leaves 210–215; iii. Fertile shoots 215–218; iv. The structure of Sigillarian stems 218–224; v. Sigillaria Brardi 224–226.  
STIGMARIA. Pp. 227–247.
BOTHRODENDREAE.  Pp. 248–270.
Bothrodendron 248–264. a. B. minutifolium 251–253; b. B. punctatum 254, 255; c. B. kiltorkense 255–259. Anatomy of vegetative shoots of Bothrodendron 260–262; Cones of Bothrodendron 262–264.  
Pinakodendron
264
Omphalophloios
264–266
General considerations
266–270
SEED-BEARING PLANTS CLOSELY ALLIED TO
MEMBERS OF THE LYCOPODIALES.
  Pp. 271–279.
  i. Lepidocarpon 271–275; ii. Miadesmia 275–279.  
FILICALES.  Pp. 280–323.
I.
Leptosporangiate Filicales
283–316
  Eufilicineae 284–316. Osmundaceae 285, 286; Schizaeaceae 286–291; Matonineae 291–293; Loxsomaceae 293; Hymenophyllaceae 294; Cyatheaceae 294–296; Dennstaedtiinae 296; Polypodiaceae 296; Parkeriaceae 297; Dipteridinae 298.

The habit, leaf-form, and distribution of ferns 300–309; The anatomy of ferns 309–316.

 
II.
Marattiales
316–321
III.
Ophioglossales
321–323
FOSSIL FERNS.  Pp. 324–394.
Osmundaceae 324–346; Schizaeaceae 346–351; Gleicheniaceae 351–355; Matonineae 355–363; Hymenophyllaceae 363–365; Cyatheaceae 365–375; Polypodiaceae 375–380; Dipteridinae 380–394.
MARATTIALES (FOSSIL).  Pp. 395–411.
Ptychocarpus 397; Danaeites 398; Parapecopteris 398; Asterotheca 398–400; Hawlea 400; Scolecopteris 401, 402; Discopteris 402–404; Dactylotheca 404–406; Renaultia 406; Zeilleria 407; Urnatopteris 407; Marattiopsis 407–409; Danaeopsis 409; Nathorstia 410, 411.
PSARONIEAE.  Pp. 412–426.
OPHIOGLOSSALES (FOSSIL).  Pp. 427–431.
COENOPTERIDEAE.  Pp. 432–472.
I.
Botryoptereae 434–443  
II.
Zygoptereae 443–470  
HYDROPTERIDEAE AND SAGENOPTERIS.  Pp. 473–483.
Marsiliaceae 473–475; Salviniaceae 475–477; Sagenopteris 477–483.
GENERA OF PTERIDOSPERMS, FERNS, AND PLANTAE INCERTAE SEDIS.  Pp. 484–580.
Taeniopteris 485–494; Weichselia 494–496; Glossopteris 496–512; Gangamopteris 512–517; Lesleya 517–519; Neuropteridium 519–523; Cardiopteris 523–525; Aphlebia 525–529; Sphenopteris 529–532; Mariopteris, Diplotmema, Palmatopteris 532–537; Cephalotheca 537; Thinnfeldia 537–544; Lomatopteris 544–546; Cycadopteris 546; Ptilozamites 546–548; Ctenopteris 548–550; Dichopteris 550–552; Odontopteris 552–556; Callipteris 557–559; Callipteridium 560; Archaeopteris 552–565; Neuropteris 565–571; Cyclopteris 571, 572; Linopteris 572, 573; Alethopteris 573–576; Lonchopteris 576; Pecopteris 576–580.
INDEX
Pp. 609–624

LIST OF ILLUSTRATIONS

Several of the illustrations are printed from blocks for which I am indebted to learned societies or to individuals. The sources from which clichés were obtained are mentioned within square brackets.

Frontispiece. Thamnopteris Schlechtendalii (Eich.). From a photograph given to me by Dr Kidston and Mr Gwynne-Vaughan. (page 329.)
Fig.
 
Page
Sphenophyllostachys
2
Sphenophyllostachys Römeri
Sphenophyllum trichomatosum
S. majus
3
Sphenophyllostachys fertilis
[Council of the Royal Society of London.]
4, 5
Sphenophyllostachys Dawsoni
[Mr A. G. Tansley, Editor of the New Phytologist.]
6
Cheirostrobus pettycurensis
Pseudobornia ursina
8
Psilotum triquetrum
18
Psilotum triquetrum (anatomy)
20
Tmesipteris tannensis
22
Lycopodium (seven species)
35
Lycopodium squarrosum
36
Lycopodium cernuum
37
Lycopodium obscurum
38
Lycopodium (anatomy of stem)
41
Lycopodium (anatomy of cones)
45
Lycopodium cernuum (cone)
[Council of the Royal Society of Edinburgh.]
47–49
Selaginella grandis
50
Selaginella (anatomy)
52
Isoetes echinospora
I. lacustris
59
Isoetes lacustris (anatomy)
62
Pleuromeia Sternbergi
70
Selaginellites and Lycopodites
80
Lycopodites lanceolatus
[Council of the Geological Society of London.]
81
Lycopodites falcatus
83
Selaginellites primaevus
86
Lycostrobus Scotti
89
Picea excelsa
94
Lepidodendron Sternbergii
97
Sigillaria (leaves)
98
Lepidodendron (leaves)
99
Lepidodendron Veltheimianum
101
Lepidodendron leaf-cushion
102
Lepidodendron and Lepidophloios leaf-cushions
104
Lepidophloios leaf-cushion
108
Lepidodendron vasculare
112–122
Knorria mirabilis
125
Lepidodendron Veltheimianum (Ulodendron)
129
Diagrammatic section illustrating the branch-theory of the Ulodendroid scar
[Council of the Manchester Literary and Philosophical Society.]
132
Pinus clausa
134
Lepidophloios scoticus
135
Halonia tortuosa
136
Lepidodendron fuliginosum
[Council of the Cambridge Philosophical Society.]
143–147
Lepidodendron vasculare and L. fuliginosum
148
Lepidodendron fuliginosum
149
L. fuliginosum
150–152
Lepidodendron obovatum
154
Lepidodendron aculeatum
[Oxford University Press: Annals of Botany.]
155, 156
Stigmaria radiculosa
157
Stigmarian rootlet
158
Lepidodendron Harcourtii and L. fuliginosum
162
Lepidodendron Wünschianum
163
L. Wünschianum
165, 166
L. Wünschianum
[Editor of the New Phytologist.]
168, 169
Lepidodendron Veltheimianum
173
L. Veltheimianum and L. macrophyllum
176
Lepidodendron australe
[Dr H. Woodward, Editor of the Geological Magazine.]
179
Lepidostrobus
183, 184
Lepidodendron and Lepidostrobi
186
Lepidostrobus
188
Spencerites insignis
[Oxford University Press: Annals of Botany.]
193
Sigillaria elegans, S. rugosa, S. tessellata, Omphalophloios anglicus
197
Sigillaria McMurtriei
199
Sigillaria mammillaris
199
Sigillaria Brardi, S. laevigata, and Lepidodendron Wortheni
200
Carica sp.
202
Sigillaria
205, 206
Sigillaria Brardi
212
Sigillariostrobus
216
Sigillaria elegans and S. elongata
220
Sigillaria Brardi
225
Stigmaria ficoides
227, 228
Cyperus papyrus
230
Stages in the development of Sigillaria
236
Stigmariopsis
237
Stigmaria
241
Bothrodendron punctatum
250
Bothrodendron minutifolium, B. punctatum, B. kiltorkense and Lepidostrobus Olryi
252
Bothrodendron minutifolium
254
Bothrodendron Leslei
[Trustees of the British Museum.]
258
Bothrodendron mundum
259
Bothrostrobus
[Council of the Manchester Literary and Philosophical Society.]
263
Omphalophloios
265
Lepidocarpon Lomaxi
273
Miadesmia and Bothrodendron
276
Angiopteris evecta and Cycas revoluta
283
Osmunda cinnamomea, O. regalis, and Todea barbara
286
Schizaea elegans
287
Aneimia rotundifolia
288
Aneimia flexuosa, A. phyllitidis, Hymenophyllum, Matonia pectinata, Thyrsopteris elegans, Gleichenia
289
Gleichenia dicarpa
290
Gleichenites Rostafinskii, Gleichenia dicarpa, G. dichotoma
290
Matonia pectinata
[Council of the Royal Society.]
292
Matonia pectinata
293
Thyrsopteris elegans, Cyathea spinulosa, Dicksonia coniifolia, D. culcita, Davallia concinna, Alsophila excelsa
294
Dicksonia Bertervana
[Trustees of the British Museum.]
295
Dipteris quinquefurcata, D. conjugata, D. Wallichii, and Polypodium quercifolium
297
Davallia aculeata
299
Polypodium Billardieri
302
Polypodium quercifolium
303
Hemitelia capensis
304
Pteris aquilina
[Council of the Linnean Society of London.]
305, 306
Matonia pectinata, Matonidium, Gleichenia dicarpa, and
Trichomanes reniforme (anatomy)
310
Trichomanes scandens
[Editor of the New Phytologist.]
311
Platyzoma microphylla
[Editor of the New Phytologist.]
312
Cyathea Imrayana
[Editor of the New Phytologist.].
313
Angiopteris evecta and Marattia fraxinea
317
Angiopteris evecta and Danaea
318
Angiopteris evecta
319
Marattia fraxinea, M. Kaulfussii, Kaulfussia, and Marattiopsis Münsteri
320
Ophioglossum vulgatum
322
Botrychium virginianum
322
Zalesskya gracilis
327
Zalesskya diploxylon
328
Thamnopteris Schlechtendalii
330
Lonchopteris virginiensis
331
Osmundites Dunlopi
333
Osmundites Kolbei
[Editor of the Geological Magazine.]
334, 335
O. Kolbei
336
Cladophlebis denticulata, Todites Williamsoni, Discopteris Rallii, Kidstonia heracleensis, and Todeopsis primaeva
340
Cladophlebis denticulata
342, 345
Klukia exilis
[Council of the Cambridge Philosophical Society.]
348
Ruffordia Goepperti
349
Chrysodium lanzaeanum, Lygodium Kaulfussi, Marattia Hookeri
350
Gleichenites longipennis, G. delicatula, G. Nordenskioldi and G. Zippei
354
Gleichenites hantonensis
[Council of the Palaeontographical Society.]
356
Laccopteris elegans
[Council of the Royal Society.]
357
Matonidium Wiesneri, Marattiopsis marantacea, Gleichenites gracilis, Laccopteris Goepperti, and L. Muensteri
358
Laccopteris polypodioides
[Trustees of the British Museum.]
359
Laccopteris
[Trustees of the British Museum.]
359
? Laccopteris polypodioides
[Trustees of the British Museum.]
360
Matonidium Goepperti
[Editor of the Encyclopaedia Britannica.]
362
Senftenbergia elegans, Oligocarpia Brongniartii, Trichomanes sp., Hymenophyllum tunbridgense, Sphenopteris (Hymenophyllites) quadridactylites
364
Coniopteris hymenophylloides
[Council of the Manchester Literary and Philosophical Society.]
368
C. hymenophylloides
369
Coniopteris quinqueloba
370
Coniopteris arguta
371
Coniopteris arguta and C. hymenophylloides
372
Oncopteris Nettvalli
373
Protopteris punctata
373
Laccopteris polypodioides, L. Muensteri, Dicksonia, Onychiopsis Mantelli, Hausmannia Sewardi, H. Kohlmanni, and Protopteris Witteana
374
Adiantides antiquus and A. lindsayoides
376
Onychiopsis Mantelli
379
Dictyophyllum exile
381
Dictyophyllum Nilssoni, Rhizomopteris Schenki, Camptopteris spiralis, and D. exile
382
Dictyophyllum rugosum
[Trustees of the British Museum.]
384
Thaumatopteris Münsteri
386
Clathropteris meniscoides
387
Clathropteris egyptiaca
[Editor of the Geological Magazine.]
388
Camptopteris spiralis
389
Hausmannia dichotoma
391
Hausmannia sp.
393
Alethopteris lonchitica, Lonchopteris rugosa, Sphenopteris Hoeninghausi, Parapecopteris neuropteroides,and Pecopteris (Dactylotheca) plumosa
399
Ptychocarpus unita, Asterotheca Sternbergii, Danaeites sarepontanus, Hawlea Miltoni, H. pulcherrima, Scolecopteris elegans
400
Dactylotheca plumosa
405
D. plumosa
406
Nathorstia angustifolia and N. latifolia
410
Psaronius
414
Psaronius infarctus, P. coalescens, P. musaeformis, and P. asterolithus
416
Pecopteris Sterzeli
419
Caulopteris peltigera and Megaphyton insigne
421
Ptychopteris
423
Dicksonia antarctica
424
Rhacopteris sp.
427
Noeggerathia foliosa
429
Chiropteris Zeilleri
[Annals of the South African Museum.]
430
Tubicaulis solenites
[Editor of the New Phytologist.]
435
Botryopteris cylindrica
439
Botryopteris ramosa
441
Botryopteris antiqua
442
Clepsydropsis antiqua, Etapteris Scotti, Diplolabis forensis, Zygopteris primaria, Stauropteris oldhamia
444
Diplolabis forensis, Botryopteris forensis, Corynepteris coralloides, Schizopteris pinnata
445
Metaclepsydropsis duplex, Stauropteris oldhamia, Ankyropteris scandens
450
Ankyropteris Grayi
451
Thamnopteris Schlechtendalii, Ankyropteris corrugata, A. bibractensis
453
Ankyropteris bibractensis
454
Ankyropteris corrugata
457
Ankyropteris corrugata
[Editor of the New Phytologist.]
458
Ankyropteris corrugata
459, 460
Etapteris Scotti
[Editor of the New Phytologist.]
462
Etapteris, Botryopteris forensis
463
Stauropteris oldhamia
[Editor of the New Phytologist.]
464
Stauropteris oldhamia
467
Stauropteris oldhamia
[Editor of the New Phytologist.]
468
Stauropteris
[Editor of the New Phytologist.]
469
Asterochlaena laxa
[Editor of the New Phytologist.]
472
Sporocarp-like bodies (? Sagenopteris)
478
Regnellidium diphyllum, Sagenopteris rhoifolia
479
Sagenopteris Phillipsi
[Trustees of the British Museum.]
480
Sagenopteris Phillipsi
[Council of the Manchester Literary and Philosophical Society.]
481
Taeniopteris multinervis, Lesleya Delafondi
487
Taeniopteris Carnoti, T. spatulata, T. coriacea
490
Taeniopteris Carruthersi
[Annals of the South African Museum.]
491
Taeniopteris vittata
493
Weichselia Mantelli, W. erratica
495
Glossopteris Browniana
[Council of the Geological Society of London.]
499
Glossopteris Browniana
[Trustees of the British Museum.]
500, 501
Vertebraria indica
502
Vertebraria indica, Onoclea struthiopteris
503
Glossopteris fronds attached to rhizome
504
Glossopteris indica, G. angustifolia
[Trustees of the British Museum.]
506, 507
Glossopteris angustifolia var. taeniopteroides
[Council of the Geological Society.]
508
Blechnoxylon talbragarense
509
Glossopteris retifera
[Trustees of the British Museum.]
511
Gangamopteris cyclopteroides
[Trustees of the British Museum.]
515
Arberia sp.
517
Lesleya simplicinervis
518
Neuropteridium validum
[Trustees of the British Museum.]
520
Neuropteridium intermedium
522
Cardiopteris frondosa
524
Gunnera manicata
527
Sphenopteris obtusiloba, Pecopteris arborescens, Sphenopteris furcata
529
Sphenopteris affinis
531
Palmatopteris, Mariopteris, Diplotmema Zeilleri, Neuropteris macrophylla, N. heterophylla, N. Scheuchzeri, Alloiopteris Essinghii
535
Cephalotheca mirabilis
536
Thinnfeldia odontopteroides, Ptilozamites
[Council of the Geological Society.]
539
Thinnfeldia odontopteroides
[Council of the Geological Society.]
540
Thinnfeldia odontopteroides
[Annals of the South African Museum.]
541
Thinnfeldia rhomboidalis
542
Lomatopteris jurensis, L. Schimperi, Thinnfeldia rhomboidalis
544
Ptilozamites Heeri
547
Ctenopteris cycadea
549
Dichopteris visianica
551
Alethopteris lonchitica, Mariopteris muricata, Odontopteris cf. alpina
553
Odontopteris minor
554
Odontopteris genuina, Callipteridium gigas, Callipteris Pellati, C. lyratifolia
557
Callipteris conferta
559
Archaeopteris hibernica
561
Archaeopteris hibernica, A. archetypus, A.fissilis, A. fimbriata
564
Neuropteris with Cyclopteris leaflets
[From a block received from Mr Carruthers.]
566
Neuropteris heterophylla
568
Neuropteris macrophylla
569
Neuropteris Scheuchzeri
570
Linopteris neuropteroides
573
Alethopteris Serlii
575
Pecopteris arborescens
578