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Territory in Bird Life

Chapter 22: INDEX
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About This Book

The author investigates territorial behavior across a wide range of bird species, using field observation and theoretical interpretation to show how individuals acquire and defend breeding areas, how song and display relate to boundary marking, and how territory affects mating, interspecific conflicts, and migration. Detailed case studies and maps illustrate aggressive encounters, joint defense by both sexes, and competition between species for nesting sites. The work traces probable evolutionary functions linking territory to reproductive success, presents mechanistic and ecological factors that shape territoriality, and acknowledges where explanation remains tentative or speculative.

1 June 1915, R. M. Barrington.

2 Dictionary of Birds, p. 556.

3 Social Psychology.

4 Manual of Psychology.

5 Ibis, April 1918.

6 Zoologist, 1912, p. 327.


INDEX

  • Acquired experience, 300
  • Adjustments, transitory, of distribution, 275
  • Alarm notes, 119
  • Arrival, advantages and disadvantages of late, 33-44
  • Assemblies in winter, 262, 263
  • Assembly grounds, 173
  • Attainment of reproduction, 171
  •  
  • Barrington, R. M., on the sex of migrants, 25
  • Battle between two male Cuckoos, 82
  • —— between two Moor-Hens, 86, 92, 93, 94
  • —— —— Pied Wagtails, 86
  • —— —— Raven and Buzzard, 217
  • —— —— Raven and Peregrine, 216 law of, 13,19
  • Behaviour routine, 262
  • —— sexual, 3
  • Bickerings, 96
  • Birthplace, return to, 43, 50
  • Blackbird, 87, 182, 222, 244
  • Blackcap, 81, 156, 224, 230
  • Black Grouse, 63
  • Black-tailed Godwit, 53
  • Boundaries, 1, 5
  • —— conflicts for retention of, 7, 62
  • —— disputes as to, 1
  • Brambling, 124
  • Breeding ground, search for appropriate, 270, 271
  • —— range, extension of, 291-92
  • —— site, acquirement of, 3
  • —— stations, evolution of, 15-19
  • —— —— repeatedly visited long before nesting-time, 64
  • —— territory, 2, 3, 7
  • —— —— evolution of, 18
  • —— —— foundation of, 7
  • —— —— innate capacity to return to former, 279-81
  • Bridled Guillemot, 64
  • Brooding, 180
  • —— impulse, 191
  • Bunting, Cirl, 28, 140
  • —— Corn, 28
  • —— Reed, 28, 68, 69, 85, 104, 132, 158, 160, 244
  • —— Yellow, 28, 30, 47, 64, 140, 159, 162, 183, 187, 188, 189, 235, 236, 286
  • Buzzard, 217
  •  
  • Capacity, innate, to return to former breeding territory, 279-81
  • Carrion Crow, 226
  • Chaffinch, 28, 31, 32, 33, 45, 87, 103, 156, 159, 235, 236, 244
  • —— Donegal, 160 Change of breeding quarters owing to unsuitableness, 50
  • Chiffchaff, 49, 51, 80, 139, 140, 221, 224, 244
  • Cirl Bunting, 28, 140
  • Clarke, W. Eagle, Studies in Bird Migration, 24
  • Cleanliness of nest, 180
  • Cliff-breeding species, 63
  • Climatic changes, alteration of routine, due to, 284
  • —— changes, food dependent on, 283
  • —— conditions, influence of, 20
  • Communities, 202
  • —— birds after breeding-season remain in, 265-67
  • Competition, female, for males, 13
  • Complexity of strife, 84-85
  • Conflicts between males during the mating period, 74, 86
  • —— between males during the nesting period, 87
  • —— for areas, 10, 11, 13, 62
  • —— of Ruff, 54
  • —— sexual, 10
  • Congenital disposition, 135
  • Contests between males for possession of females, 80
  • Coot, 61
  • Corn-Bunting, 28
  • Corncrake, 39
  • Crow, Carrion, 226, 227
  • Crow, Hooded, 202
  • Cuckoo, 52, 82, 144, 296
  • —— restricted breeding area, 52
  • Curlew, 119, 138, 140, 250, 262, 263, 265, 273
  •  
  • Danger warnings, 269
  • Darwin, C., Descent of Man, 35
  • —— on the arrival of males before females, 35
  • Defence of territory, 6
  • Development, sexual, 6
  • Disposition, congenital, 135
  • —— functioning of, 74
  • —— inherited, 5
  • —— to defend the territory, 73-118
  • —— to mate, 27
  • —— to remain in a particular place in a particular environment, 6
  • —— to secure a territory, 6, 20-72
  • Distribution, adjustment of, 275
  • Dove, Turtle, 126, 232
  • Dunlin, 250
  •  
  • Emotional behaviour, 53, 82, 114
  • —— manifestation, 90, 283
  • —— response, 26
  • Enemies, 282
  • Energy, waste of, 219
  • Environment, 6
  • —— and food, 56
  • —— changes of, 283
  • —— external, conditions in, 279, 282
  • Equipment, hereditary, 6
  • Evolution of breeding stations or territory, 15, 19
  • —— of the territory, 176
  • —— of the voice, 163
  • Existence, struggle for, 294
  • Experience, acquired, 300
  • Experiments, removal of nests for, 181, 185, 190, 213, 214
  • Exposure, its effect on nestlings, 180
  • External environment, conditions in, 279, 282
  •  
  • Falcon, 48, 71
  • Feeding grounds, neutral, 125 in communities, 70 Females, fighting amongst, 109-118
  • —— sexual impulse of, 13
  • Fieldfare, 124
  • Fighting instinct, 79, 82
  • Flight, emotional behaviour of Godwit during, 53
  • Flocks, in winter, birds collect together in, 262
  • Flycatcher, 244
  • Food, procuring of, 5
  • —— abundance, or scarcity of, its relation to prosperity of young, 15, 16
  • —— its bearing, on the movement of flocks, 262
  • —— rearing of young dependent on rapid and regular, 179, 195
  • —— supply, proximity to, necessary for rearing young, 179, 195
  • Fortuitous mating, 174
  • Fowler, Ward, on the value of communities, Rooks, 202
  • Fulmar, 121, 247
  • Functional activity, 259
  • —— instinct of Reeve, 173
  • Functioning of the disposition, 275
  • —— of the primary dispositions, 100
  •  
  • Garden Warbler, 223, 225, 230
  • Gätke, H., Birds of Heligoland, 24
  • —— on the absence of song in birds on Heligoland, 124
  • —— on the early arrival of Guillemots on Heligoland, 64
  • Godwit, emotional behaviour of, during flight, 53
  • —— Black-tailed, 53
  • Grasshopper Warbler, 39, 131, 139, 153, 155, 187, 244
  • Greenfinch, 28, 33, 140, 235, 236
  • Gregarious instinct, 20, 61, 141, 265-66, 269, 276, 289, 290, 291, 296, 300
  • Grouse, Black, 63
  • Guillemot, Bridled, 64
  • —— Common, 63, 64, 121, 192, 195, 206, 211, 247
  • —— Ringed, 64
  • Gull, Common, 119
  • —— Herring, 210, 278
  •  
  • Habit formation, law of, 8, 62, 65, 66, 67, 205
  • Hawfinch, 28
  • Headquarters, 176, 206, 207, 274
  • —— restricted, 8, 9, 30, 50, 58, 64, 127
  • Hedge-Sparrow, 213, 221, 244
  • Hereditary equipment, 6
  • Herring-Gull, 210, 278
  • Hooded-Crow, 202
  • Hostility and territory, relationship between, 242
  • House-Sparrow, 218
  •  
  • Imitation, vocal, powers of, 156, 157, 161
  • Impulse, internal, 279
  • —— to brood, 191
  • Inherited disposition, 5
  • Instinct, fighting, 79-82
  • —— gregarious, 20, 61, 141, 265, 266, 269, 276, 289, 290, 291, 296, 300
  • —— migratory, 37
  • —— of song related to establishment of territory, 125
  • —— sexual, reawakening of, 4, 18
  • Instinctive response, 180
  • Instincts susceptible to stimulation, 259
  • Internal impulse, 279 Internal stimulation, 62, 123
  • Interpretation of battles, 75
  • Intolerance of other birds, 218, 219
  • Intrusion resented, 274
  • Isolation, impulse to seek, 288
  • —— of male, 12, 62, 65, 73, 81
  • —— of male during breeding season, 267, 272, 273, 275, 281
  •  
  • Jay, 87, 156, 283
  •  
  • Kestrel, 228
  • Kittiwake, 116, 200, 247
  •  
  • Lapwing, 58, 59, 61, 62, 64, 84, 103, 104, 126, 189, 190, 220, 251, 284
  • Lapwing, life-history of, 58-61
  • Late arrival, advantages and disadvantages of, 33-44
  • Law of battle, 74, 75, 86
  • Lesser Whitethroat, 230, 244
  • Linnet, 156
  •  
  • M'Dougall, Dr, Social Psychology, 77
  • Magpie, 219, 283
  • Males arrive before advent of females, 24
  • Marsh-Warbler, 39, 40, 52, 81, 132, 140, 153, 155, 156, 165, 225
  • Martin, 201, 218
  • Mating, difficulties of, 172
  • —— fortuitous, 174
  • Maximum number supportable in a given locality, 49
  • Meadow-Pipit, 188
  • Meeting places for antics, 54, 63
  • Mental Image, 77
  • Merlin, 227
  • Migration, 3-4
  • —— distance no criterion, 279
  • —— its relation to territory, 259
  • Migration, origin of, 260, 277
  • —— phenomenon of, 277
  • Migratory instinct, 37
  • —— species more highly specialised than resident species, 56
  • Missel-Thrush, 21
  • Mobility of the land and stability of the water, 260
  • Moor-hen, 61, 85, 103, 218, 250, 251
  • Morgan, Professor Lloyd, on instinctive behaviour, 74
  • —— on emotional behaviour, 114
  •  
  • Nest, cleaning of, 180
  • —— construction of, 3
  • Nests, removal of, for experiments, 181, 185, 190, 213, 214
  • Neutral feeding grounds, 62, 125
  • —— ground, 98
  • Newton, E., on the arrival at breeding stations of males before female, 24, 35
  • Nightingale, 39, 156
  • Notes of alarm, 119
  • —— of anger, 119
  • —— of recognition, 139
  • —— of warning, 119, 139, 141, 145, 151, 153
  •  
  • Offspring, rearing of, 3, 4
  • Organic change, sexual, 92, 123
  • —— changes, 65
  • —— condition of Reeve, 173
  • Owl, Wood, 156
  •  
  • Paired for life, 55-56
  • Parental instinctive response, failure of, 185
  • Partridge, 87, 218
  • Persecution, Carrion Crow and Magpie, 226 Persecution, Carrion Crow and Rook, 227
  • —— House Sparrow and Martin, 218
  • —— Lapwing and Snipe, 220
  • —— Raven and Buzzard, 217
  • —— —— and Peregrine, 216
  • —— Starling and Woodpecker, 218, 237
  • Persistency to remain in territory, 68
  • Pied-Wagtail, 86, 155
  • Pigeon, Wood, 219
  • Pipit, Meadow, 188
  • —— Tree, 51, 188, 189, 222, 244, 278
  • Polyandrous females, 144
  • Predatory species, 268
  • Promiscuous pairing of Ruffs, 172
  • Proximity to food-supply necessary for rearing young, 179, 195
  • Psychology, Manual of, 1
  • Puffin, 63, 116, 200
  • Pugnacious instinct, 87-109
  • Pugnacity, 11, 62
  • —— of females to obtain mates, 109-118
  • —— of males, prior to mating-season, 77-81
  • —— of Moor-Hen, 218
  •  
  • Racial preparation, 41, 43, 46, 67, 205, 206, 266
  • Rail, Water, 218
  • Raven, 48, 202, 216
  • Razor-bill, 63, 64, 200, 247
  • Readjustment of territory, 146
  • Rearing of offspring, 3, 4
  • Red-backed Shrike, 39, 50, 51, 156
  • Redbreast, 47
  • Redshank, 139
  • Redwing, 124
  • Redstart, 230, 244
  • Reed-Bunting, 28, 68, 69, 85, 104, 132, 156, 158, 160, 244, 246, 273
  • Reed-Warbler, 49, 51, 68, 81, 132, 140, 152, 153, 211, 225
  • Reeve, 171
  • Relation of song to the territory, 119-68
  • —— of territory to migration, 259
  • —— of territory to the system of reproduction, 169-214
  • Relationship to a territory, 169
  • Reproduction, 14, 15
  • —— and territory, 169-214
  • —— attainment of, 2, 6, 37
  • —— goal of, 6
  • Ringed Guillemot, 64
  • Robbery of territory, 104-107
  • Rock-formation, suitability for Guillemots nesting on, 196
  • Rook, 202, 227
  • Routine behaviour, 262
  • Ruff, 54, 63, 172
  • —— meeting places for conflicts, 54
  • Ruffs, promiscuous pairing of, 172
  •  
  • Savi's Warbler, 139
  • Sedge-Warbler, 25, 44, 152, 226, 244
  • Selous, E., on the life-history of Ruffs and Reeves, 172
  • —— on meeting places for conflicts and antics, 54
  • —— on the meeting places of Black Grouse, 63
  • Service, Robert, on flocks of unmated Sedge-Warblers, 44, 45
  • Sexual behaviour, 3
  • —— conflicts, 10
  • —— development, 6
  • —— function, discharge of, 2, 3, 26
  • —— impulse of females, 13 Sexual instinct in the migratory male, 26
  • —— of Reeve, 173
  • —— —— reawakening of, 4, 18
  • —— life of birds, 1
  • —— maturity, males arrive at, before females, 36
  • —— organic change, 92, 123
  • —— selection, 166
  • Shag, 121
  • Shrike, Red-backed, 39, 50, 51, 156
  • Skylark, 188, 236, 244
  • Snipe, 153, 156, 219, 220
  • Sociability when not paired, 125, 126
  • Song, as an aid in searching for a mate, 12
  • —— its influence on mating, 167
  • —— origin of, 138
  • —— relation to reproduction, 123
  • —— relation to territory, 119-168
  • —— volume of, influenced by age, seasonal sexual development, or isolation, 166
  • Song-Thrush, 222, 244
  • Sparrow, House, 218
  • Sparrow-Hawk, 269
  • Spring, at approach of, birds lose their shyness, 138
  • Stability of the water and mobility of the land, 260
  • Starling, 217, 218, 237, 251
  • Stimulation, internal, 62, 123
  • —— question of, 284
  • Stonechat, 87 187, 188, 189, 222, 233, 234
  • Stout, Dr, Manual of Psychology, 1, 77
  • Struggle for existence, 294
  • Susceptibility to position, 96
  • Swallow, 21, 156, 278
  •  
  • Territory, 1, 5
  • —— adjustment of, 10
  • Territory and reproduction, 169-214
  • —— breeding, 2, 3, 7
  • —— dates of acquisition of, 33
  • —— defence of, 6
  • —— desertion of, after rearing young, 276
  • —— disposition to defend, 73-118
  • —— disposition to secure, 6, 20-72
  • —— establishment of, 74, 285
  • —— evolution of, 176
  • —— failure to secure, 286
  • —— fights for, 10, 11, 13, 62
  • —— ownership of, 189
  • —— possession of, a stimulus to song, 136
  • —— its relation to migration, 259
  • —— its relation to reproduction, 169-214
  • —— readjustment of, 147
  • —— restriction of, advantageous for mating, 172
  • —— restricted, 8, 9, 30, 50, 58, 64
  • —— separate for male and female Cuckoo, 144
  • —— song, its relation to the, 119-68
  • —— temporary desertion of, 28, 35, 58, 59
  • —— and hostility, relationship between, 242
  • Thrush, Song, 222, 244
  • Tit, Blue, 221, 226
  • —— Great, 221
  • —— Long-tailed, 226
  • Tradition, 300
  • Tree-Pipit, 51, 188, 189, 222, 232, 244, 278
  • Turtle-Dove, 126, 232
  •  
  • Union of sexes, 12
  • Ussher, H. B., on the hostility between Choughs and Hooded Crows and Choughs and Ravens, 227
  •  
  • Vocal Imitation, 156, 157, 161
  • Voice calls of Curlew, 263
  •  
  • Wagtail, Pied, 86, 155
  • —— Yellow, 278
  • Wanderings from land, Guillemots, 193
  • Warbler, Garden, 223, 225, 230
  • —— Grasshopper, 39, 131, 139, 155, 187, 244
  • —— Marsh, 39, 40, 52, 81, 132, 140, 155, 156, 225
  • —— Reed, 49, 51, 68, 81, 132, 140, 152, 153, 211, 225
  • —— Savi's, 139
  • —— Sedge, 25, 44, 152, 226, 244
  • —— Willow, 25, 47, 50, 51, 80, 91, 140, 187, 211, 232, 244, 273
  • —— Wood, 50, 51, 132, 221
  • Warfare between different species and its relation to the territory, 215-58
  • Warning notes, 119
  • —— of danger, 269
  • Water Rail, 218
  • Wheatear, 25, 51
  • Whimbrel, 140
  • Whinchat, 39, 50, 51, 81, 222, 232, 233, 234, 244
  • Whitethroat, 25, 50, 68, 69, 124, 140, 182, 187, 189, 190, 213, 230, 244
  • —— Lesser, 230, 244
  • Wild Duck, 250
  • Will, the, to fight, 102
  • Willow-Warbler, 25, 47, 50, 51, 80, 91, 140, 187, 211, 232, 244, 273
  • Winter assemblies, 262-63
  • Witherby, H. F., in British Birds, on the return to former breeding-ground, 281
  • Wood-Owl, 156
  • Wood-Pigeon, 219
  • Wood-Warbler, 50, 51, 132, 221
  • Woodpecker, Lesser Spotted, 237
  • —— Great Spotted, 237, 238
  • —— Green, 20, 71, 156, 208, 218, 237
  • Wren, 244
  •  
  • Yellow Bunting, 28, 30, 47, 64, 140, 159, 162, 183, 187, 188, 189, 235, 236, 286
  • Young die in nest from exposure, 184, 185