CHAPTER XVI

THE CAMBRIAN

THE PALEOZOIC ERA. The second volume of the geological record, called the Paleozoic (Greek, PALAIOS, ancient; ZOE, life), has come down to us far less mutilated and defaced than has the first volume, which contains the traces of the most ancient life of the globe. Fossils are far more abundant in the Paleozoic than in the earlier strata, while the sediments in which they were entombed have suffered far less from metamorphism and other causes, and have been less widely buried from view, than the strata of the pre-Cambrian groups. By means of their fossils we can correlate the formations of widely separated regions from the beginning of the Paleozoic on, and can therefore trace some outline of the history of the continents.

Paleozoic time, although shorter than the pre-Cambrian as measured by the thickness of the strata, must still be reckoned in millions of years. During this vast reach of time the changes in organisms were very great. It is according to the successive stages in the advance of life that the Paleozoic formations are arranged in five systems,—the CAMBRIAN, the ORDOVICIAN, the SILURIAN, the DEVONIAN, and the CARBONIFEROUS. On the same basis the first three systems are grouped together as the older Paleozoic, because they alike are characterized by the dominance of the invertebrates; while the last two systems are united in the later Paleozoic, and are characterized, the one by the dominance of fishes, and the other by the appearance of amphibians and reptiles.

Each of these systems is world-wide in its distribution, and may be recognized on any continent by its own peculiar fauna. The names first given them in Great Britain have therefore come into general use, while their subdivisions, which often cannot be correlated in different countries and different regions, are usually given local names.

The first three systems were named from the fact that their strata are well displayed in Wales. The Cambrian carries the Roman name of Wales, and the Ordovician and Silurian the names of tribes of ancient Britons which inhabited the same country. The Devonian is named from the English county Devon, where its rocks were early studied. The Carboniferous was so called from the large amount of coal which it was found to contain in Great Britain and continental Europe.

THE CAMBRIAN

DISTRIBUTION OF STRATA. The Cambrian rocks outcrop in narrow belts about the pre-Cambrian areas of eastern Canada and the Lake Superior region, the Adirondacks and the Green Mountains. Strips of Cambrian formations occupy troughs in the pre-Cambrian rocks of New England and the maritime provinces of Canada; a long belt borders on the west the crystalline rocks of the Blue Ridge; and on the opposite side of the continent the Cambrian reappears in the mountains of the Great Basin and the Canadian Rockies. In the Mississippi valley it is exposed in small districts where uplift has permitted the stripping off of younger rocks. Although the areas of outcrop are small, we may infer that Cambrian rocks were widely deposited over the continent of North America.

PHYSICAL GEOGRAPHY. The Cambrian system of North America comprises three distinct series, the LOWER CAMBRIAN, the MIDDLE CAMBRIAN, and the UPPER CAMBRIAN, each of which is characterized by its own peculiar fauna. In sketching the outlines of the continent as it was at the beginning of the Paleozoic, it must be remembered that wherever the Lower Cambrian formations now are found was certainly then sea bottom, and wherever the Lower Cambrian are wanting, and the next formations rest directly on pre-Cambrian rocks, was probably then land.

EARLY CAMBRIAN GEOGRAPHY. In this way we know that at the opening of the Cambrian two long, narrow mediterranean seas stretched from north to south across the continent. The eastern sea extended from the Gulf of St. Lawrence down the Champlain-Hudson valley and thence along the western base of the Blue Ridge south at least to Alabama. The western sea stretched from the Canadian Rockies over the Great Basin and at least as far south as the Grand Canyon of the Colorado in Arizona.

Between these mediterraneans lay a great central land which included the pre-Cambrian U-shaped area of the Laurentian peneplain, and probably extended southward to the latitude of New Orleans. To the east lay a land which we may designate as APPALACHIA, whose western shore line was drawn along the site of the present Blue Ridge, but whose other limits are quite unknown. The land of Appalachia must have been large, for it furnished a great amount of waste during the entire Paleozoic era, and its eastern coast may possibly have lain even beyond the edge of the present continental shelf. On the western side of the continent a narrow land occupied the site of the Sierra Nevada Mountains.

Thus, even at the beginning of the Paleozoic, the continental plateau of North America had already been left by crustal movements in relief above the abysses of the great oceans on either side. The mediterraneans which lay upon it were shallow, as their sediments prove. They were EPICONTINENTAL SEAS; that is, they rested UPON (Greek, EPI) the submerged portion of the continental plateau. We have no proof that the deep ocean ever occupied any part of where North America now is.

The Middle and Upper Cambrian strata are found together with the Lower Cambrian over the area of both the eastern and the western mediterraneans, so that here the sea continued during the entire period. The sediments throughout are those of shoal water. Coarse cross-bedded sandstones record the action of strong shifting currents which spread coarse waste near shore and winnowed it of finer stuff. Frequent ripple marks on the bedding planes of the strata prove that the loose sands of the sea floor were near enough to the surface to be agitated by waves and tidal currents. Sun cracks show that often the outgoing tide exposed large muddy flats to the drying action of the sun. The fossils, also, of the strata are of kinds related to those which now live in shallow waters near the shore.

The sediments which gathered in the mediterranean seas were very thick, reaching in places the enormous depth of ten thousand feet. Hence the bottoms of these seas were sinking troughs, ever filling with waste from the adjacent land as fast as they subsided.

LATE CAMBRIAN GEOGRAPHY. The formations of the Middle and Upper Cambrian are found resting unconformably on the pre-Cambrian rocks from New York westward into Minnesota and at various points in the interior, as in Missouri and in Texas. Hence after earlier Cambrian time the central land subsided, with much the same effect as if the Mississippi valley were now to lower gradually, and the Gulf of Mexico to spread northward until it entered Lake Superior. The Cambrian seas transgressed the central land and strewed far and wide behind their advancing beaches the sediments of the later Cambrian upon an eroded surface of pre-Cambrian rocks.

The succession of the Cambrian formations in North America records many minor oscillations and varying conditions of physical geography; yet on the whole it tells of widening seas and lowering lands. Basal conglomerates and coarse sandstones which must have been laid near shore are succeeded by shaly sandstones, sandy shales, and shales. Toward the top of the series heavy beds of limestone, extending from the Blue Ridge to Missouri, speak of clear water, and either of more distant shores or of neighboring lands which were worn or sunk so low that for the most part their waste was carried to the sea in solution.

In brief, the Cambrian was a period of submergence. It began with the larger part of North America emerged as great land masses. It closed with most of the interior of the continental plateau covered with a shallow sea.

THE LIFE OF THE CAMBRIAN PERIOD

It is now for the first time that we find preserved in the offshore deposits of the Cambrian seas enough remains of animal life to be properly called a fauna. Doubtless these remains are only the most fragmentary representation of the life of the time, for the Cambrian rocks are very old and have been widely metamorphosed. Yet the five hundred and more species already discovered embrace all the leading types of invertebrate life, and are so varied that we must believe that their lines of descent stretch far back into the pre-Cambrian past.

PLANTS. No remains of plants have been found in Cambrian strata, except some doubtful markings, as of seaweed.

SPONGES. The sponges, the lowest of the multicellular animals, were represented by several orders. Their fossils are recognized by the siliceous spicules, which, as in modern sponges, either were scattered through a mass of horny fibers or were connected in a flinty framework.

COELENTERATES. This subkingdom includes two classes of interest to the geologist,—the HYDROZOA, such as the fresh-water hydra and the jellyfish, and the CORALS. Both classes existed in the Cambrian.

The Hydrozoa were represented not only by jellyfish but also by the GRAPTOLITE, which takes its name from a fancied resemblance of some of its forms to a quill pen. It was a composite animal with a horny framework, the individuals of the colony living in cells strung on one or both sides along a hollow stem, and communicating by means of a common flesh in this central tube. Some graptolites were straight, and some curved or spiral; some were single stemmed, and others consisted of several radial stems united. Graptolites occur but rarely in the Upper Cambrian. In the Ordovician and Silurian they are very plentiful, and at the close of the Silurian they pass out of existence, never to return.

CORALS are very rarely found in the Cambrian, and the description of their primitive types is postponed to later chapters treating of periods when they became more numerous.

ECHINODERMS. This subkingdom comprises at present such familiar forms as the crinoid, the starfish, and the sea urchin. The structure of echinoderms is radiate. Their integument is hardened with plates or particles of carbonate of lime.

Of the free echinoderms, such as the starfish and the sea urchin, the former has been found in the Cambrian rocks of Europe, but neither have so far been discovered in the strata of this period in North America. The stemmed and lower division of the echinoderms was represented by a primitive type, the CYSTOID, so called from its saclike form, A small globular or ovate "calyx" of calcareous plates, with an aperture at the top for the mouth, inclosed the body of the animal, and was attached to the sea bottom by a short flexible stalk consisting of disks of carbonate of lime held together by a central ligament.

ARTHOPODS. These segmented animals with "jointed feet," as their name suggests, may be divided in a general way into water breathers and air breathers. The first-named and lower division comprises the class of the CRUSTACEA,—arthropods protected by a hard exterior skeleton, or "crust,"—of which crabs, crayfish, and lobsters are familiar examples. The higher division, that of the air breathers, includes the following classes: spiders, scorpions, centipedes, and insects.

THE TRILOBITE. The aquatic arthropods, the Crustacea, culminated before the air breathers; and while none of the latter are found in the Cambrian, the former were the dominant life of the time in numbers, in size, and in the variety of their forms. The leading crustacean type is the TRILOBITE, which takes its name from the three lobes into which its shell is divided longitudinally. There are also three cross divisions,—the head shield, the tail shield, and between the two the thorax, consisting of a number of distinct and unconsolidated segments. The head shield carries a pair of large, crescentic, compound eyes, like those of the insect. The eye varies greatly in the number of its lenses, ranging from fourteen in some species to fifteen thousand in others. Figure 268, C, is a restoration of the trilobite, and shows the appendages, which are found preserved only in the rarest cases.

During the long ages of the Cambrian the trilobite varied greatly. Again and again new species and genera appeared, while the older types became extinct. For this reason and because of their abundance, trilobites are used in the classification of the Cambrian system. The Lower Cambrian is characterized by the presence of a trilobitic fauna in which the genus Olenellus is predominant. This, the OLENELLUS ZONE, is one of the most important platforms in the entire geological series; for, the world over, it marks the beginning of Paleozoic time, while all underlying strata are classified as pre-Cambrian. The Middle Cambrian is marked by the genus Paradoxides, and the Upper Cambrian by the genus Olenus. Some of the Cambrian trilobites were giants, measuring as much as two feet long, while others were the smallest of their kind, a fraction of an inch in length.

Another type of crustacean which lived in the Cambrian and whose order is still living is illustrated in Figure 269.

WORMS. Trails and burrows of worms have been left on the sea beaches and mud flats of all geological times from the Algonkian to the present.

BRACHIOPODS. These soft-bodied animals, with bivalve shells and two interior armlike processes which served for breathing, appeared in the Algonkian, and had now become very abundant. The two valves of the brachiopod shell are unequal in size, and in each valve a line drawn from the beak to the base divides the valve into two equal parts. It may thus be told from the pelecypod mollusk, such as the clam, whose two valves are not far from equal in size, each being divided into unequal parts by a line dropped from the beak.

Brachiopods include two orders. In the most primitive order—that of the INARTICULATE brachiopods—the two valves are held together only by muscles of the animal, and the shell is horny or is composed of phosphate of lime. The DISCINA, which began in the Algonkian, is of this type, as is also the LINGULELLA of the Cambrian. Both of these genera have lived on during the millions of years of geological time since their introduction, handing down from generation to generation with hardly any change to their descendants now living off our shores the characters impressed upon them at the beginning.

The more highly organized ARTICULATE brachiopods have valves of carbonate of lime more securely joined by a hinge with teeth and sockets (Fig. 270). In the Cambrian the inarticulates predominate, though the articulates grow common toward the end of the period.

MOLLUSKS. The three chief classes of mollusks—the PELECYPODS (represented by the oyster and clam of to-day), the GASTROPODS (represented now by snails, conches, and periwinkles), and the CEPHALOPODS (such as the nautilus, cuttlefish, and squids)—were all represented in the Cambrian, although very sparingly.

Pteropods, a suborder of the gastropods, appeared in this age. Their papery shells of carbonate of lime are found in great numbers from this time on.

Cephalopods, the most highly organized of the mollusks, started into existence, so far as the record shows, toward, the end of the Cambrian, with the long extinct ORTHOCERAS (STRAIGHTHORN) and the allied genera of its family. The Orthoceras had a long, straight, and tapering shell, divided by cross partitions into chambers. The animal lived in the "body chamber" at the larger end, and walled off the other chambers from it in succession during the growth of the shell. A central tube, the SIPHUNCLE, passed through from the body chamber to the closed tip of the cone.

The seashells, both brachiopods and mollusks, are in some respects the most important to the geologist of all fossils. They have been so numerous, so widely distributed, and so well preserved because of their durable shells and their station in growing sediments, that better than any other group of organisms they can be used to correlate the strata of different regions and to mark by their slow changes the advance of geological time.

CLIMATE. The life of Cambrian times in different countries contains no suggestion of any marked climatic zones, and as in later periods a warm climate probably reached to the polar regions.

CHAPTER XVII

THE ORDOVICIAN AND SILURIAN
[Footnote: Often known as the Lower Silurian.]

THE ORDOVICIAN

In North America the Ordovician rocks lie conformably on the Cambrian. The two periods, therefore, were not parted by any deformation, either of mountain making or of continental uplift. The general submergence which marked the Cambrian continued into the succeeding period with little interruption.

SUBDIVISIONS AND DISTRIBUTION OF STRATA. The Ordovician series, as they have been made out in New York, are given for reference in the following table, with the rocks of which they are chiefly composed:

5 Hudson . . . . . . . . shales 4 Utica . . . . . . . . shales 3 Trenton . . . . . . . limestones 2 Chazy . . . . . . . . limestones 1 Calciferous . . . . . sandy limestones

These marine formations of the Ordovician outcrop about the Cambrian and pre-Cambrian areas, and, as borings show, extend far and wide over the interior of the continent beneath more recent strata. The Ordovician sea stretched from Appalachia across the Mississippi valley. It seems to have extended to California, although broken probably by several mountainous islands in the west.

PHYSICAL GEOGRAPHY. The physical history of the period is recorded in the succession of its formations. The sandstones of the Upper Cambrian, as we have learned, tell of a transgressing sea which gradually came to occupy the Mississippi valley and the interior of North America. The limestones of the early and middle Ordovician show that now the shore had become remote and the lands had become more low. The waters now had cleared. Colonies of brachiopods and other lime-secreting animals occupied the sea bottom, and their debris mantled it with sheets of limy ooze. The sandy limestones of the Calciferous record the transition stage from the Cambrian when some sand was still brought in from shore. The highly fossiliferous limestones of the Trenton tell of clear water and abundant life. We need not regard this epicontinental sea as deep. No abysmal deposits have been found, and the limestones of the period are those which would be laid in clear, warm water of moderate depth like that of modern coral seas.

The shales of the Utica and Hudson show that the waters of the sea now became clouded with mud washed in from land. Either the land was gradually uplifted, or perhaps there had arrived one of those periodic crises which, as we may imagine, have taken place whenever the crust of the shrinking earth has slowly given way over its great depressions, and the ocean has withdrawn its waters into deepening abysses. The land was thus left relatively higher and bordered with new coastal plains. The epicontinental sea was shoaled and narrowed, and muds were washed in from the adjacent lands.

THE TACONIC DEFORMATION. The Ordovician was closed by a deformation whose extent and severity are not yet known. From the St. Lawrence River to New York Bay, along the northwestern and western border of New England, lies a belt of Cambrian-Ordovician rocks more than a mile in total thickness, which accumulated during the long ages of those periods in a gradually subsiding trough between the Adirondacks and a pre-Cambrian range lying west of the Connecticut River. But since their deposition these ancient sediments have been crumpled and crushed, broken with great faults, and extensively metamorphosed. The limestones have recrystallized into marbles, among them the famous marbles of Vermont; the Cambrian sandstones have become quartzites, and the Hudson shale has been changed to a schist exposed on Manhattan Island and northward.

In part these changes occurred at the close of the Ordovician, for in several places beds of Silurian age rest unconformably on the upturned Ordovician strata; but recent investigations have made it probable that the crustal movements recurred at later times, and it was perhaps in the Devonian and at the close of the Carboniferous that the greater part of the deformation and metamorphism was accomplished. As a result of these movements,— perhaps several times repeated,—a great mountain range was upridged, which has been long since leveled by erosion, but whose roots are now visible in the Taconic Mountains of western New England.

THE CINCINNATI ANTICLINE. Over an oval area in Ohio, Indiana, and Kentucky, whose longer axis extends from north to south through Cincinnati, the Ordovician strata rise in a very low, broad swell, called the Cincinnati anticline. The Silurian and Devonian strata thin out as they approach this area and seem never to have deposited upon it. We may regard it, therefore, as an island upwarped from the sea at the close of the Ordovician or shortly after.

PETROLEUM AND NATURAL GAS. These valuable illuminants and fuels are considered here because, although they are found in traces in older strata, it is in the Ordovician that they occur for the first time in large quantities. They range throughout later formations down to the most recent.

The oil horizons of California and Texas are Tertiary; those of Colorado, Cretaceous; those of West Virginia, Carboniferous; those of Pennsylvania, Kentucky, and Canada, Devonian; and the large field of Ohio and Indiana belongs to the Ordovician and higher systems.

Petroleum and natural gas, wherever found, have probably originated from the decay of organic matter when buried in sedimentary deposits, just as at present in swampy places the hydrogen and carbon of decaying vegetation combine to form marsh gas. The light and heat of these hydrocarbons we may think of, therefore, as a gift to the civilized life of our race from the humble organisms, both animal and vegetable, of the remote past, whose remains were entombed in the sediments of the Ordovician and later geological ages.

Petroleum is very widely disseminated throughout the stratified rocks. Certain limestones are visibly greasy with it, and others give off its characteristic fetid odor when struck with a hammer. Many shales are bituminous, and some are so highly charged that small flakes may be lighted like tapers, and several gallons of oil to the ton may be obtained by distillation.

But oil and gas are found in paying quantities only when certain conditions meet:

1. A SOURCE below, usually a bituminous shale, from whose organic matter they have been derived by slow change.

2. A RESERVOIR above, in which they have gathered. This is either a porous sandstone or a porous or creviced limestone.

3. Oil and gas are lighter than water, and are usually under pressure owing to artesian water. Hence, in order to hold them from escaping to the surface, the reservoir must have the shape of an ANTICLINE, DOME, or LENS.

4. It must also have an IMPERVIOUS COVER, usually a shale. In these reservoirs gas is under a pressure which is often enormous, reaching in extreme cases as high as a thousand five hundred pounds to the square inch. When tapped it rushes out with a deafening roar, sometimes flinging the heavy drill high in air. In accounting for this pressure we must remember that the gas has been compressed within the pores of the reservoir rock by artesian water, and in some cases also by its own expansive force. It is not uncommon for artesian water to rise in wells after the exhaustion of gas and oil.

LIFE OF THE ORDOVICIAN

During the ages of the Ordovician, life made great advances. Types already present branched widely into new genera and species, and new and higher types appeared.

Sponges continued from the Cambrian. Graptolites now reached their climax.

STROMATOPORA—colonies of minute hydrozoans allied to corals—grew in places on the sea floor, secreting stony masses composed of thin, close, concentric layers, connected by vertical rods. The Stromatopora are among the chief limestone builders of the Silurian and Devonian periods.

CORALS developed along several distinct lines, like modern corals they secreted a calcareous framework, in whose outer portions the polyps lived. In the Ordovician, corals were represented chiefly by the family of the CHOETETES, all species of which are long since extinct. The description of other types of corals will be given under the Silurian, where they first became abundant.

ECHINODERMS. The cystoid reaches its climax, but there appear now two higher types of echinoderms,—the crinoid and the starfish. The CRINOID, named from its resemblance to the lily, is like the cystoid in many respects, but has a longer stem and supports a crown of plumose arms. Stirring the water with these arms, it creates currents by which particles of food are wafted to its mouth. Crinoids are rare at the present time, but they grew in the greatest profusion in the warm Ordovician seas and for long ages thereafter. In many places the sea floor was beautiful with these graceful, flowerlike forms, as with fields of long-stemmed lilies. Of the higher, free-moving classes of the echinoderms, starfish are more numerous than in the Cambrian, and sea urchins make their appearance in rare archaic forms.

CRUSTACEANS. Trilobites now reach their greatest development and more than eleven hundred species have been described from the rocks of this period. It is interesting to note that in many species the segments of the thorax have now come to be so shaped that they move freely on one another. Unlike their Cambrian ancestors, many of the Ordovician trilobites could roll themselves into balls at the approach of danger. It is in this attitude, taken at the approach of death, that trilobites are often found in the Ordovician and later rocks. The gigantic crustaceans called the EURYPTERIDS were also present in this period.

The arthropods had now seized upon the land. Centipedes and insects of a low type, the earliest known land animals, have been discovered in strata of this system.

BRYOZOANS. No fossils are more common in the limestones of the time than the small branching stems and lacelike mats of the bryozoans,—the skeletons of colonies of a minute animal allied in structure to the brachiopod.

BRACHIOPODS. These multiplied greatly, and in places their shells formed thick beds of coquina. They still greatly surpassed the mollusks in numbers.

CEPHALOPODS. Among the mollusks we must note the evolution of the cephalopods. The primitive straight Orthoceras has now become abundant. But in addition to this ancestral type there appears a succession of forms more and more curved and closely coiled, as illustrated in Figure 285. The nautilus, which began its course in this period, crawls on the bottom of our present seas.

VERTEBRATES. The most important record of the Ordovician is that of the appearance of a new and higher type, with possibilities of development lying hidden in its structure that the mollusk and the insect could never hope to reach. Scales and plates of minute fishes found in the Ordovician rocks near Canon City, Colorado, show that the humblest of the vertebrates had already made its appearance. But it is probable that vertebrates had been on the earth for ages before this in lowly types, which, being destitute of hard parts, would leave no record.

THE SILURIAN

The narrowing of the seas and the emergence of the lands which characterized the closing epoch of the Ordovician in eastern North America continue into the succeeding period of the Silurian. New species appear and many old species now become extinct.

THE APPALACHIAN REGION. Where the Silurian system is most fully developed, from New York southward along the Appalachian Mountains, it comprises four series:

4 Salina . . . shales, impure limestones, gypsum, salt 3 Niagara . . . chiefly limestones 2 Clinton . . . sandstones, shales, with some limestones 1 Medina . . . conglomerates, sandstones

The rocks of these series are shallow-water deposits and reach the total thickness of some five thousand feet. Evidently they were laid over an area which was on the whole gradually subsiding, although with various gentle oscillations which are recorded in the different formations. The coarse sands of the heavy Medina formations record a period of uplift of the oldland of Appalachia, when erosion went on rapidly and coarse waste in abundance was brought down from the hills by swift streams and spread by the waves in wide, sandy flats. As the lands were worn lower the waste became finer, and during an epoch of transition—the Clinton— there were deposited various formations of sandstones, shales, and limestones. The Niagara limestones testify to a long epoch of repose, when low-lying lands sent little waste down to the sea.

The gypsum and salt deposits of the Salina show that toward the close of the Silurian period a slight oscillation brought the sea floor nearer to the surface, and at the north cut off extensive tracts from the interior sea. In these wide lagoons, which now and then regained access to the open sea and obtained new supplies of salt water, beds of salt and gypsum were deposited as the briny waters became concentrated by evaporation under a desert climate. Along with these beds there were also laid shales and impure limestones.

In New York the "salt pans" of the Salina extended over an area one hundred and fifty miles long from east to west and sixty miles wide, and similar salt marshes occurred as far west as Cleveland, Ohio, and Goderich on Lake Huron. At Ithaca, New York, the series is fifteen hundred feet thick, and is buried beneath an equal thickness of later strata. It includes two hundred and fifty feet of solid salt, in several distinct beds, each sealed within the shales of the series.

Would you expect to find ancient beds of rock salt inclosed in beds of pervious sandstone?

The salt beds of the Salina are of great value. They are reached by well borings, and their brines are evaporated by solar heat and by boiling. The rock salt is also mined from deep shafts.

Similar deposits of salt, formed under like conditions, occur in
the rocks of later systems down to the present. The salt beds of
Texas are Permian, those of Kansas are Permian, and those of
Louisiana are Tertiary.

THE MISSISSIPPI VALLEY. The heavy near-shore formations of the Silurian in the Appalachian region thin out toward the west. The Medina and the Clinton sandstones are not found west of Ohio, where the first passes into a shale and the second into a limestone. The Niagara limestone, however, spreads from the Hudson River to beyond the Mississippi, a distance of more than a thousand miles. During the Silurian period the Mississippi valley region was covered with a quiet, shallow, limestone-making sea, which received little waste from the low lands which bordered it.

The probable distribution of land and sea in eastern North America and western Europe is shown in Figure 287. The fauna of the interior region and of eastern Canada are closely allied with that of western Europe, and several species are identical. We can hardly account for this except by a shallow-water connection between the two ancient epicontinental seas. It was perhaps along the coastal shelves of a northern land connecting America and Europe by way of Greenland and Iceland that the migration took place, so that the same species came to live in Iowa and in Sweden.

THE WESTERN UNITED STATES. So little is found of the rocks of the system west of the Missouri River that it is quite probable that the western part of the United States had for the most part emerged from the sea at the close of the Ordovician and remained land during the Silurian. At the same time the western land was perhaps connected with the eastern land of Appalachia across Arkansas and Mississippi; for toward the south the Silurian sediments indicate an approach to shore.

LIFE OF THE SILURIAN

In this brief sketch it is quite impossible to relate the many changes of species and genera during the Silurian.

CORALS. Some of the more common types are familiarly known as cup corals, honeycomb corals, and chain corals. In the CUP CORALS the most important feature is the development of radiating vertical partitions, or SEPTA, in the cell of the polyp. Some of the cup corals grew in hemispherical colonies (Fig. 288), while many were separate individuals (Fig. 289), building a single conical, or horn-shaped cell, which sometimes reached the extreme size of a foot in length and two or three inches in diameter.

HONEYCOMB CORALS consist of masses of small, close-set prismatic cells, each crossed by horizontal partitions, or TABULAE, while the septa are rudimentary, being represented by faintly projecting ridges or rows of spines.

CHAIN CORALS are also marked by tabulae. Their cells form elliptical tubes, touching each other at the edges, and appearing in cross section like the links of a chain. They became extinct at the end of the Silurian.

The corals of the SYRINGOPORA family are similar in structure to chain corals, but the tubular columns are connected only in places.

To the echinoderms there is now added the BLASTOID (bud-shaped). The blastoid is stemmed and armless, and its globular "head" or "calyx," with its five petal-like divisions, resembles a flower bud. The blastoids became more abundant in the Devonian, culminated in the Carboniferous, and disappeared at the end of the Paleozoic.

The great eurypterids—some of which were five or six feet in length—and the cephalopods were still masters of the seas. Fishes were as yet few and small; trilobites and graptolites had now passed their prime and had diminished greatly in numbers. Scorpions are found in this period both in Europe and in America. The limestone-making seas of the Silurian swarmed with corals, crinoids, and brachiopods.

With the end of the Silurian period the AGE OF INVERTEBRATES comes to a close, giving place to the Devonian, the AGE OF FISHES.

CHAPTER XVIII

THE DEVONIAN

In America the Silurian is not separated from the Devonian by any mountain-making deformation or continental uplift. The one period passed quietly into the other. Their conformable systems are so closely related, and the change in their faunas is so gradual, that geologists are not agreed as to the precise horizon which divides them.

SUBDIVISIONS AND PHYSICAL GEOGRAPHY. The Devonian is represented in New York and southward by the following five series. We add the rocks of which they are chiefly composed.

5 Chemung . . . . . . sandstones and sandy shales 4 Hamilton . . . . . . shales and sandstones 3 Corniferous . . . . . . limestones 2 Oriskany . . . . . . sandstones 1 Helderberg . . . . . . limestones

The Helderberg is a transition epoch referred by some geologists to the Silurian. The thin sandstones of the Oriskany mark an epoch when waves worked over the deposits of former coastal plains. The limestones of the Corniferous testify to a warm and clear wide sea which extended from the Hudson to beyond the Mississippi. Corals throve luxuriantly, and their remains, with those of mollusks and other lime-secreting animals, built up great beds of limestone. The bordering continents, as during the later Silurian, must now have been monotonous lowlands which sent down little of even the finest waste to the sea.

In the Hamilton the clear seas of the previous epoch became clouded with mud. The immense deposits of coarse sandstones and sandy shales of the Chemung, which are found off what was at the time the west coast of Appalachia, prove an uplift of that ancient continent.

The Chemung series extends from the Catskill Mountains to northeastern Ohio and south to northeastern Tennessee, covering an area of not less than a hundred thousand square miles. In eastern New York it attains three thousand feet in thickness; in Pennsylvania it reaches the enormous thickness of two miles; but it rapidly thins to the west. Everywhere the Chemung is made of thin beds of rapidly alternating coarse and fine sands and clays, with an occasional pebble layer, and hence is a shallow-water deposit. The fine material has not been thoroughly winnowed from the coarse by the long action of strong waves and tides. The sands and clays have undergone little more sorting than is done by rivers. We must regard the Chemung sandstones as deposits made at the mouths of swift, turbid rivers in such great amount that they could be little sorted and distributed by waves.

Over considerable areas the Chemung sandstones bear little or no trace of the action of the sea. The Catskill Mountains, for example, have as their summit layers some three thousand feet of coarse red sandstones of this series, whose structure is that of river deposits, and whose few fossils are chiefly of fresh-water types. The Chemung is therefore composed of delta deposits, more or less worked over by the sea. The bulk of the Chemung equals that of the Sierra Nevada Mountains. To furnish this immense volume of sediment a great mountain range, or highland, must have been upheaved where the Appalachian lowland long had been. To what height the Devonian mountains of Appalachia attained cannot be told from the volume of the sediments wasted from them, for they may have risen but little faster than they were worn down by denudation. We may infer from the character of the waste which they furnished to the Chemung shores that they did not reach an Alpine height. The grains of the Chemung sandstones are not those which would result from mechanical disintegration, as by frost on high mountain peaks, but are rather those which would be left from the long chemical decay of siliceous crystalline rocks; for the more soluble minerals are largely wanting. The red color of much of the deposits points to the same conclusion. Red residual clays accumulated on the mountain sides and upland summits, and were washed as ocherous silt to mingle with the delta sands. The iron- bearing igneous rocks of the oldland also contributed by their decay iron in solution to the rivers, to be deposited in films of iron oxide about the quartz grains of the Chemung sandstones, giving them their reddish tints.

LIFE OF THE DEVONIAN

PLANTS. The lands were probably clad with verdure during Silurian times, if not still earlier; for some rare remains of ferns and other lowly types of vegetation have been found in the strata of that system. But it is in the Devonian that we discover for the first time the remains of extensive and luxuriant forests. This rich flora reached its climax in the Carboniferous, and it will be more convenient to describe its varied types in the next chapter.

RHIZOCARPS. In the shales of the Devonian are found microscopic spores of rhizocarps in such countless numbers that their weight must be reckoned in hundreds of millions of tons. It would seem that these aquatic plants culminated in this period, and in widely distant portions of the earth swampy flats and shallow lagoons were filled with vegetation of this humble type, either growing from the bottom or floating free upon the surface. It is to the resinous spores of the rhizocarps that the petroleum and natural gas from Devonian rocks are largely due. The decomposition of the spores has made the shales highly bituminous, and the oil and gas have accumulated in the reservoirs of overlying porous sandstones.

INVERTEBRATES. We must pass over the ever-changing groups of the invertebrates with the briefest notice. Chain corals became extinct at the close of the Silurian, but other corals were extremely common in the Devonian seas. At many places corals formed thin reefs, as at Louisville, Kentucky, where the hardness of the reef rock is one of the causes of the Falls of the Ohio.

Sponges, echinoderms, brachiopods, and mollusks were abundant. The cephalopods take a new departure. So far in all their various forms, whether straight, as the Orthoceras, or curved, or close- coiled as in the nautilus, the septum, or partition dividing the chambers, met the inner shell along a simple line, like that of the rim of a saucer. There now begins a growth of the septum by which its edges become sharply corrugated, and the suture, or line of juncture of the septum and the shell, is thus angled. The group in which this growth of the septum takes place is called the GONIATITE (Greek GONIA, angle).

VERTEBRATES. It is with the greatest interest that we turn now to study the backboned animals of the Devonian; for they are believed to be the ancestors of the hosts of vertebrates which have since dominated the earth. Their rudimentary structures foreshadowed what their descendants were to be, and give some clue to the earliest vertebrates from which they sprang. Like those whose remains are found in the lower Paleozoic systems, all of these Devonian vertebrates were aquatic and go under the general designation of fishes.

The lowest in grade and nearest, perhaps, to the ancestral type of vertebrates, was the problematic creature, an inch or so long, of Figure 297. Note the circular mouth not supplied with jaws, the lack of paired fins, and the symmetric tail fin, with the column of cartilaginous, ringlike vertebrae running through it to the end. The animal is probably to be placed with the jawless lampreys and hags,—a group too low to be included among true fishes.

OSTRACODERMS. This archaic group, long since extinct, is also too lowly to rank among the true fishes, for its members have neither jaws nor paired fins. These small, fishlike forms were cased in front with bony plates developed in the skin and covered in the rear with scales. The vertebrae were not ossified, for no trace of them has been found.

DEVONIAN FISHES. The TRUE FISHES of the Devonian can best be understood by reference to their descendants now living. Modern fishes are divided into several groups: SHARKS and their allies; DIPNOANS; GANOIDS, such as the sturgeon and gar; and TELEOSTS,— most common fishes, such as the perch and cod.

SHARKS. Of all groups of living fishes the sharks are the oldest and still retain most fully the embryonic characters of their Paleozoic ancestors. Such characters are the cartilaginous skeleton, and the separate gill slits with which the throat wall is pierced and which are arranged in line like the gill openings of the lamprey. The sharks of the Silurian and Devonian are known to us chiefly by their teeth and fin spines, for they were unprotected by scales or plates, and were devoid of a bony skeleton. Figure 299 is a restoration of an archaic shark from a somewhat higher horizon. Note the seven gill slits and the lappetlike paired fins. These fins seem to be remnants of the continuous fold of skin which, as embryology teaches, passed from fore to aft down each side of the primitive vertebrate.

Devonian sharks were comparatively small. They had not evolved into the ferocious monsters which were later to be masters of the seas.

DIPNOANS, OR LUNG FISHES. These are represented to-day by a few peculiar fishes and are distinguished by some high structures which ally them with amphibians. An air sac with cellular spaces is connected with the gullet and serves as a rudimentary lung. It corresponds with the swim bladder of most modern fishes, and appears to have had a common origin with it. We may conceive that the primordial fishes not only had gills used in breathing air dissolved in water, but also developed a saclike pouch off the gullet. This sac evolved along two distinct lines. On the line of the ancestry of most modern fishes its duct was closed and it became the swim bladder used in flotation and balancing. On another line of descent it was left open, air was swallowed into it, and it developed into the rudimentary lung of the dipnoans and into the more perfect lungs of the amphibians and other air- breathing vertebrates.

One of the ancient dipnoans is illustrated in Figure 300. Some of the members of this order were, like the ostracoderms, cased in armor, but their higher rank is shown by their powerful jaws and by other structures. Some of these armored fishes reached twenty- five feet in length and six feet across the head. They were the tyrants of the Devonian seas.

GANOIDS. These take their name from their enameled plates or scales of bone. The few genera now surviving are the descendants of the tribes which swarmed in the Devonian seas. A restoration of one of a leading order, the FRINGE-FINNED ganoids, is given in Figure 301. The side fins, which correspond to the limbs of the higher vertebrates, are quite unlike those of most modern fishes. Their rays, instead of radiating from a common base, fringe a central lobe which contains a cartilaginous axis. The teeth of the Devonian ganoids show a complicated folded structure.

GENERAL CHARACTERISTICS OF DEVONIAN FISHES. THE NOTOCHORD IS PERSISTENT. The notochord is a continuous rod of cartilage, or gristle, which in the embryological growth of vertebrate animals supports the spinal nerve cord before the formation of the vertebrae. In most modern fishes and in all higher vertebrates the notochord is gradually removed as the bodies of the vertebrae are formed about it; but in the Devonian fishes it persists through maturity and the vertebrae remain incomplete.

THE SKELETON IS CARTILAGINOUS. This also is an embryological characteristic. In the Devonian fishes the vertebrae, as well as the other parts of the skeleton, have not ossified, or changed to bone, but remain in their primitive cartilaginous condition.

THE TAIL FIN IS VERTEBRATED. The backbone runs through the fin and is fringed above and below with its vertical rays. In some fishes with vertebrated tail fins the fin is symmetric, and this seems to be the primitive type. In others the tail fin is unsymmetric: the backbone runs into the upper lobe, leaving the two lobes of unequal size. In most modern fishes (the teleosts) the tail fin is not vertebrated: the spinal column ends in a broad plate, to which the diverging fin rays are attached.

But along with these embryonic characters, which were common to all Devonian fishes, there were other structures in certain groups which foreshadowed the higher structures of the land vertebrates which were yet to come: air sacs which were to develop into lungs, and cartilaginous axes in the side fins which were a prophecy of limbs. The vertebrates had already advanced far enough to prove the superiority of their type of structure to all others. Their internal skeleton afforded the best attachment for muscles and enabled them to become the largest and most powerful creatures of the time. The central nervous system, with the predominance given to the ganglia at the fore end of the nerve cord,—the brain,— already endowed them with greater energy than the invertebrates; and, still more important, these structures contained the possibility of development into the more highly organized land vertebrates which were to rule the earth.

TELEOSTS. The great group of fishes called the teleosts, or those with complete bony skeletons, to which most modern fishes belong, may be mentioned here, although in the Devonian they had not yet appeared. The teleosts are a highly specialized type, adapted most perfectly to their aquatic environment. Heavy armor has been discarded, and reliance is placed instead on swiftness. The skeleton is completely ossified and the notochord removed. The vertebrae have been economically withdrawn from the tail, and the cartilaginous axis of the side fins has been fotfoid unnecessary. The air sac has become a swim bladder. In this complete specialization they have long since lost the possibility of evolving into higher types.

It is interesting to note that the modern teleosts in their embryological growth pass through the stages which characterized the maturity of their Devonian ancestors; their skeleton is cartilaginous and their tail fin vertebrated.