[1] To any one who has attentively read my 'Origin of Species' this Introduction will be superfluous. As I stated that work that I should soon publish the facts on which the conclusions given in it were founded, I here beg permission to remark that the great delay in publishing this first work has been caused by continued ill-health.

[2] M. Pouchet has recently ('Plurality of Races,' Eng. Translat., 1864, p. 83, &c.) insisted that variation under domestication throws no light on the natural modification of species. I cannot perceive the force of his arguments, or, to speak more accurately, of his assertions to this effect.

[3] Léon Dufour in 'Annales des Scienc. Nat.' (3rd series, Zoolog.), tom. v. p. 6.

[4] In treating the several subjects included in the present and succeeding works I have continually been led to ask for information from many zoologists, botanists, geologists, breeders of animals, and horticulturists, and I have invariably received from them the most generous assistance. Without such aid I could have effected little. I have repeatedly applied for information and specimens to foreigners, and to British merchants and officers of the Government residing in distant lands, and, with the rarest exceptions, I have received prompt, open-handed, and valuable assistance. I cannot express too strongly my obligations to the many persons who have assisted me, and who, I am convinced, would be equally willing to assist others in any scientific investigation.

[5] Owen, 'British Fossil Mammals,' p. 123 to 133. Pictet's 'Traité de Pal.,' 1853, tom. i. p. 202. De Blainville, in his 'Ostéographie, Canidæ,' p. 142, has largely discussed the whole subject, and concludes that the extinct parent of all domesticated dogs came nearest to the wolf in organization, and to the jackal in habits.

[6] Pallas, I believe, originated this doctrine in 'Act. Acad. St. Petersburgh,' 1780, Part ii. Ehrenberg has advocated it, as may be seen in De Blainville's 'Ostéographie,' p. 79. It has been carried to an extreme extent by Col. Hamilton Smith in the 'Naturalist Library,' vol. ix. and x. Mr. W. C. Martin adopts it in his excellent 'History of the Dog,' 1845; as does Dr. Morton, as well as Nott and Gliddon, in the United States. Prof. Low, in his 'Domesticated Animals,' 1845, p. 666, comes to this same conclusion. No one has argued on this side with more clearness and force than the late James Wilson, of Edinburgh, in various papers read before the Highland Agricultural and Wernerian Societies. Isidore Geoffroy Saint Hilaire ('Hist. Nat. Gén.,' 1860, tom. iii. p. 107), though he believes that most dogs have descended from the jackal, yet inclines to the belief that some are descended from the wolf. Prof. Gervais ('Hist. Nat. Mamm.,' 1855, tom. ii. p. 69), referring to the view that all the domestic races are the modified descendants of a single species, after a long discussion, says, "Cette opinion est, suivant nous du moins, la moins probable."

[7] Berjeau, 'The Varieties of the Dog; in old Sculptures and Pictures,' 1863. 'Der Hund,' von Dr. F. L. Walther, s. 48, Giessen, 1817: this author seems carefully to have studied all classical works on the subject. See also 'Volz, Beiträge zur Kultur-geschichte,' Leipzig, 1852, s. 115. 'Youatt on the Dog,' 1845, p. 6. A very full history is given by De Blainville in his 'Ostéographie, Canidæ.'

[8] I have seen drawings of this dog from the tomb of the son of Esar Haddon, and clay models in the British Museum. Nott and Gliddon, in their 'Types of Mankind,' 1854, p. 393, give a copy of these drawings. This dog has been called a Thibetan mastiff, but Mr. H. A. Oldfield, who is familiar with the so-called Thibet mastiff, and has examined the drawings in the British Museum, informs me that he considers them different.

[9] 'Proc. Zoolog. Soc.,' July 12th, 1831.

[10] 'Sporting in Algeria,' p. 51.

[11] Berjeau gives fac-similes of the Egyptian drawings. Mr. C. L. Martin, in his 'History of the Dog,' 1845, copies several figures from the Egyptian monuments, and speaks with much confidence with respect to their identity with still living dogs. Messrs. Nott and Gliddon ('Types of Mankind,' 1854, p. 388) give still more numerous figures. Mr. Gliddon asserts that a curl-tailed greyhound, like that represented on the most ancient monuments, is common in Borneo; but the Rajah, Sir J. Brooke, informs me that no such dog exists there.

[12] These, and the following facts on the Danish remains, are taken from M. Morlot's most interesting memoir in 'Soc. Vaudoise des Sc. Nat.,' tom. vi., 1860, pp. 281, 299, 320.

[13] 'Die Fauna der Pfahlbauten,' 1861, s. 117, 162.

[14] De Blainville, 'Ostéographie, Canidæ.'

[15] Sir R. Schomburgk has given me information on this head. See also 'Journal of R. Geograph. Soc.,' vol. xiii., 1843, p. 65.

[16] 'Domestication of Animals:' Ethnological Soc., Dec. 22nd, 1863.

[17] 'Journal of Researches,' &c., 1845, p. 393. With respect to Canis antarcticus, see p. 193. For the case of the antelope, see 'Journal Royal Geograph. Soc.,' vol. xxiii. p. 94.

[18] The authorities for the foregoing statements are as follow:—Richardson, in 'Fauna Boreali-Americana,' 1829, pp. 64, 75; Dr. Kane, 'Arctic Explorations,' 1856, vol. i. pp. 398, 455; Dr. Hayes, 'Arctic Boat Journey,' 1860, p. 167. Franklin's 'Narrative,' vol. i. p. 269, gives the case of three whelps of a black wolf being carried away by the Indians. Parry, Richardson, and others, give accounts of wolves and dogs naturally crossing in the eastern parts of North America. Seeman, in his 'Voyage of H.M.S. Herald,' 1853, vol. ii. p. 26, says the wolf is often caught by the Esqimaux for the purpose of crossing with their dogs, and thus adding to their size and strength. M. Lamare-Picquot, in 'Bull. de la Soc. d'Acclimat.,' tom. vii., 1860, p. 148, gives a good account of the half-bred Esquimaux dogs.

[19] 'Fauna Boreali-Americana,' 1829, pp. 73, 78, 80. Nott and Gliddon, 'Types of Mankind,' p. 383. The naturalist and traveller Bartram is quoted by Hamilton Smith, in 'Nat. Hist. Lib.,' vol. x. p. 156. A Mexican domestic dog seems also to resemble a wild dog of the same country; but this may be the prairie-wolf. Another capable judge, Mr. J. K. Lord ('The Naturalist in Vancouver Island,' 1866, vol. ii. p. 218), says that the Indian dog of the Spokans, near the Rocky Mountains, "is beyond all question nothing more than a tamed Cayote or prairie-wolf," or Canis latrans.

[20] I quote this from Mr. R. Hill's excellent account of the Alco or domestic dog of Mexico, in Gosse's 'Naturalist's Sojourn in Jamaica,' 1851, p. 329.

[21] 'Naturgeschichte der Saeugethiere von Paraguay,' 1830, s. 151.

[22] Quoted in Humboldt's 'Aspects of Nature' (Eng. transl.), vol. i. p. 108.

[23] Paget's 'Travels in Hungary and Transylvania,' vol. i. p. 501. Jeitteles, 'Fauna Hungariæ Superioris,' 1862, s. 13. See Pliny, 'Hist. of the World' (Eng. transl.), 8th book, ch. xl., about the Gauls crossing their dogs. See also 'Hist. Animal.' lib. viii. c. 28. For good evidence about wolves and dogs naturally crossing near the Pyrenees, see M. Mauduyt, 'Du Loup et de ses Races,' Poitiers, 1851; also Pallas, in 'Acta Acad. St. Petersburgh,' 1780, part ii. p. 94.

[24] I give this on excellent authority, namely, Mr. Blyth (under the signature of Zoophilus), in the 'Indian Sporting Review,' Oct. 1856, p. 134. Mr. Blyth states that he was struck with the resemblance between a brush-tailed race of pariah-dogs, north-west of Cawnpore, and the Indian wolf. He gives corroborative evidence with respect to the dogs of the valley of the Nerbudda.

[25] For numerous and interesting details on the resemblance of dogs and jackals, see Isid. Geoffroy St. Hilaire, 'Hist. Nat. Gén.,' 1860, tom. iii. p. 101. See also 'Hist. Nat. des Mammifères,' par Prof. Gervais, 1855, tom. ii. p. 60.

[26] Güldenstädt, 'Nov. Comment. Acad. Petrop.,' tom. xx., pro anno 1775, p. 449.

[27] Quoted by De Blainville in his 'Ostéographie, Canidæ,' pp. 79, 98.

[28] See Pallas, in 'Act. Acad. St. Petersburgh,' 1780, part ii. p. 91. For Algeria, see Isid. Geoffroy St. Hilaire, 'Hist. Nat. Gén.,' tom. iii. p. 177. In both countries it is the male jackal which pairs with female domestic dogs.

[29] John Barbut's 'Description of the Coast of Guinea in 1746.'

[30] 'Travels in South Africa,' vol. ii. p. 272.

[31] Selwyn, Geology of Victoria; 'Journal of Geolog. Soc.,' vol. xiv., 1858, p. 536, and vol. xvi., 1860, p. 148; and Prof M^cCoy, in 'Annals and Mag. of Nat. Hist.' (3rd series), vol. ix., 1862, p. 147. The Dingo differs from the dogs of the central Polynesian islands. Dieffenbach remarks ('Travels,' vol. ii. p. 45) that the native New Zealand dog also differs from the Dingo.

[32] 'Proceedings Zoolog. Soc.,' 1833, p. 112. See, also, on the taming of the common wolf, L. Lloyd, 'Scandinavian Adventures,' vol. i. p. 460, 1854. With respect to the jackal, see Prof. Gervais, 'Hist. Nat. Mamm.,' tom. ii. p. 61. With respect to the aguara of Paraguay, see Rengger's work.

[33] Roulin, in 'Mém. présent. par divers Savans,' tom. vi. p. 341.

[34] Martin, 'History of the Dog,' p. 14.

[35] Quoted by L. Lloyd in 'Field Sports of North of Europe,' vol. i. p. 387.

[36] Quatrefages, 'Soc. d'Acclimat.,' May 11th, 1863, p. 7.

[37] 'Annals and Mag. of Nat. Hist.,' vol. xv., 1845, p. 140.

[38] Azara, 'Voyages dans l'Amér. Mérid.,' tom. i. p. 381; his account is fully confirmed by Rengger. Quatrefages gives an account of a bitch brought from Jerusalem to France which burrowed a hole and littered in it. See 'Discours, Exposition des Races Canines,' 1865, p. 3.

[39] With respect to wolves burrowing holes, see Richardson, Fauna Boreali-Americana,' p. 64; and Bechstein, 'Naturgesch. Deutschlands,' b. i. s. 617.

[40] See Poeppig, 'Reise in Chile,' b. i. s. 290; Mr. G. Clarke, as above; and Rengger, s. 155.

[41] Dogs, 'Nat. Library,' vol. x. p. 121: an endemic South American dog seems also to have become feral in this island. See Gosse's 'Jamaica,' p. 340.

[42] Low, 'Domesticated Animals,' p. 650.

[43] 'The Naturalist Library,' Dogs, vol. x. pp. 4, 19.

[44] Quoted by Prof. Gervais, 'Hist. Nat. Mamm.,' tom. ii. p. 66.

[45] J. Hunter shows that the long period of seventy-three days given by Buffon is easily explained by the bitch having received the dog many times during a period of sixteen days ('Phil. Transact.,' 1787, p. 253). Hunter found that the gestation of a mongrel from wolf and dog ('Phil. Transact.,' 1759, p. 160) apparently was sixty-three days, for she received the dog more than once. The period of a mongrel dog and jackal was fifty-nine days. Fred. Cuvier found the period of gestation of the wolf to be ('Dict. Class. d'Hist. Nat.,' tom. iv. p. 8) two months and a few days, which agrees with the dog. Isid. G. St. Hilaire, who has discussed the whole subject, and from whom I quote Bellingeri, states ('Hist. Nat. Gén.,' tom. iii. p. 112) that in the Jardin des Plantes the period of the jackal has been found to be from sixty to sixty-three days, exactly as with the dog.

[46] See Isid. Geoffroy St. Hilaire, 'Hist. Nat. Gén.,' tom. iii. p. 112, on the odour of jackals. Col. Ham. Smith, in 'Nat. Hist. Lib.,' vol. x. p. 289.

[47] Quoted by Quatrefages in 'Bull. Soc. d'Acclimat.,' May 11th, 1863.

[48] 'Journal de la Physiologie,' tom. ii. p. 385.

[49] See Mr. R. Hill's excellent account of this breed in Gosse's 'Jamaica,' p. 338; Rengger's 'Saeugethiere von Paraguay,' s. 153. With respect to Spitz dogs, see Bechstein's 'Naturgesch. Deutschlands,' 1801, b. i. s. 638. With respect to Dr. Hodgkin's statement made before Brit. Assoc., see 'The Zoologist,' vol. iv., for 1845-46, p. 1097.

[50] 'Acta Acad. St. Petersburgh,' 1780, part ii. pp. 84, 100.

[51] M. Broca has shown ('Journal de Physiologie,' tom. ii. p. 353) that Buffon's experiments have been often misrepresented. Broca has collected (pp. 390-395) many facts on the fertility of crossed dogs, wolves, and jackals.

[52] 'De la Longévité Humaine,' par M. Flourens, 1855, p. 143. Mr. Blyth says ('Indian Sporting Review,' vol. ii. p. 137) that he has seen in India several hybrids from the pariah-dog and jackal; and between one of these hybrids and a terrier. The experiments of Hunter on the jackal are well known. See also Isid. Geoffroy St. Hilaire, 'Hist. Nat. Gén.,' tom. iii, p. 217, who speaks of the hybrid offspring of the jackal as perfectly fertile for three generations.

[53] On authority of F. Cuvier, quoted in Bronn's 'Geschichte der Natur,' B. ii. s. 164.

[54] W. C. L. Martin, 'History of the Dog,' 1845, p. 203. Mr. Philip P. King, after ample opportunities of observation, informs me that the Dingo and European dogs often cross in Australia.

[55] Rüppel, 'Neue Wirbelthiere von Abyssinien,' 1835-40; 'Mammif.,' s. 39, pl. xiv. There is a specimen of this fine animal in the British Museum.

[56] Even Pallas admits this: see 'Act. Acad. St. Petersburgh,' 1780, p. 93.

[57] Quoted by I. Geoffroy, 'Hist. Nat. Gén.,' tom. iii. p. 453.

[58] F. Cuvier, in 'Annales du Muséum,' tom. xviii. p. 337; Godron, 'De l'Espèce,' tom. i. p. 342; and Col. Ham. Smith, in 'Naturalist's Library,' vol. ix. p. 101.

[59] Isid. Geoffroy Saint Hilaire, 'Hist. des Anomalies,' 1832, tom. i. p. 660. Gervais, 'Hist. Nat. des Mammifères,' tom. ii., 1855, p. 66. De Blainville ('Ostéographie, Canidæ,' p. 137) has also seen an extra molar on both sides.

[60] 'Ostéographie, Canidæ,' p. 137.

[61] Würzburger, 'Medecin, Zeitschrift,' 1860, B. i. s. 265.

[62] Mr. Yarell, in 'Proc. Zoolog. Soc.,' Oct. 8th, 1833. Mr. Waterhouse showed me a skull of one of these dogs, which had only a single molar on each side and some imperfect incisors.

[63] Quoted in 'The Veterinary,' London, vol. viii. p. 415.

[64] 'Hist Nat. Général,' tom. iii. p. 448.

[65] W. Scrope, 'Art of Deer-Stalking,' p. 354.

[66] Quoted by Col. Ham. Smith in 'Naturalist's Library,' vol. x. p. 79.

[67] De Blainville, 'Ostéographie, Canidæ,' p. 134. F. Cuvier, 'Annales du Muséum,' tom. xviii. p. 342. In regard to mastiffs, see Col. Ham. Smith, 'Nat Lib.,' vol. x. p. 218. For the Thibet mastiff, see Mr. Hodgson in 'Journal of As. Soc. of Bengal,' vol. i., 1832, p. 342.

[68] 'The Dog,' 1845, p. 186. With respect to diseases, Youatt asserts (p. 167) that the Italian greyhound is "strongly subject" to polypi in the matrix or vagina. The spaniel and pug (p. 182) are most liable to bronchocele. The liability to distemper (p. 232) is extremely different in different breeds. On the distemper, see also Col. Hutchinson on 'Dog Breaking,' 1850, p. 279.

[69] See Youatt on the Dog, p. 15; 'The Veterinary,' London, vol. xi. p. 235.

[70] 'Journal of As. Soc. of Bengal,' vol. iii. p. 19.

[71] 'Travels,' vol. ii. p. 15.

[72] Hodgson, in 'Journal of As. Soc. of Bengal,' vol. i. p. 342.

[73] 'Field Sports of the North of Europe,' vol. ii. p. 165.

[74] 'Hist. Nat. des Mammif., 1855, tom. ii. pp. 66, 67.

[75] 'History of Quadrupeds,' 1793, vol. i. p. 238.

[76] 'Oriental Field Sports,' quoted by Youatt, 'The Dog,' p. 15.

[77] Quoted by Mr. Galton, 'Domestication of Animals,' p. 13.

[78] 'Hist. Nat. Gén.,' tom. iii. p. 450.

[79] Mr. Greenhow on the Canadian Dog, in Loudon's 'Mag. of Nat. Hist.,' vol. vi., 1833, p. 511.

[80] See Mr. C. O. Groom-Napier on the webbing of the hind feet of Otter-hounds, in 'Land and Water,' Oct. 13th, 1866, p. 270.

[81] 'Fauna Boreali-Americana,' 1829, p. 62.

[82] 'The Horse in all his Varieties,' &c., 1829, pp. 230, 234.

[83] 'The Dog,' 1845, pp. 31, 35; with respect to King Charles's spaniel, p. 45; for the setter, p. 90.

[84] In the 'Encyclop. of Rural Sports,' p. 557.

[85] 'The Farrier,' 1828, vol. i. p. 337.

[86] See Col. Hamilton Smith on the antiquity of the Pointer, in 'Naturalist's Library,' vol. x. p. 195.

[87] The Newfoundland dog is believed to have originated from a cross between the Esquimaux dog and a large French hound. See Dr. Hodgkin, 'Brit. Assoc.,' 1844; Bechstein's 'Naturgesch. Deutschlands,' Band i. s. 574; 'Naturalist's Library,' vol. x. p. 132; also Mr. Jukes' 'Excursion in and about Newfoundland.'

[88] De Blainville, 'Ostéographie, Felis,' p. 65, on the character of F. caligulata; pp. 85, 89, 90, 175, on the other mummied species. He quotes Ehrenberg on F. maniculata being mummied.

[89] Asiatic Soc. of Calcutta; Curator's Report, Aug. 1856. The passage from Sir W. Jardine is quoted from this Report. Mr. Blyth, who has especially attended to the wild and domestic cats of India, has given in this Report a very interesting discussion on their origin.

[90] 'Fauna Hungariæ Sup.,' 1862, s. 12.

[91] Isid. Geoffrey Saint Hilaire, 'Hist. Nat. Gen.,' tom. iii. p. 177.

[92] 'Proc. Zoolog. Soc.,' 1863, p. 184.

[93] 'Saeugethiere von Paraguay,' 1830, s. 212.

[94] 'Mem. présentés par divers Savans: Acad. Roy. des Sciences,' tom. vi. p. 346. Gomara first noticed this fact in 1554.

[95] 'Narrative of Voyages,' vol. ii. p. 180.

[96] J. Crawfurd, 'Descript. Dict. of the Indian Islands,' p. 255. The Madagascar cat is said to have a twisted tail: see Desmarest, in 'Encyclop. Nat. Mamm.,' 1820, p. 233, for some of the other breeds.

[97] Admiral Lutké's Voyage, vol. iii. p. 308.

[98] 'Zoology of the Voyage of the Beagle, Mammalia,' p. 20. Dieffenbach, 'Travels in New Zealand,' vol. ii. p. 185. Ch. St. John, 'Wild Sports of the Highlands,' 1846, p. 49.

[99] Quoted by Isid. Geoffroy, 'Hist. Nat. Gén.,' tom. iii. p. 427.

[100] Rütimeyer, 'Fauna der Pfalbauten,' 1861, s. 122.

[101] See Youatt on the Horse: J. Lawrence on the Horse, 1829: W. C. L. Martin, 'History of the Horse,' 1845: Col. Ham. Smith, in 'Naturalist's Library, Horses,' 1841, vol. xii.: Prof. Veith, 'Die Naturgesch. Haussäugethiere,' 1856.

[102] Crawfurd, 'Descript. Dict. of Indian Islands,' 1856, p. 153. "There are many different breeds, every island having at least one peculiar to it." Thus in Sumatra there are at least two breeds; in Achin and Batubara one; in Java several breeds; one in Bali, Lomboc, Sumbawa (one of the best breeds), Tambora, Bima, Gunung-api, Celebes, Sumba, and Philippines. Other breeds are specified by Zollinger in the 'Journal of the Indian Archipelago,' vol. v. p. 343, &c.

[103] 'The Horse,' &c., by John Lawrence, 1829, p. 14.

[104] 'The Veterinary,' London, vol. v. p. 543.

[105] Proc. Veterinary Assoc., in 'The Veterinary,' vol. xiii. p. 42.

[106] 'Bulletin de la Soc. Géolog.,' tom. xxii., 1866, p. 22.

[107] Mr. Percival, of the Enniskillen Dragoons, in 'The Veterinary,' vol. i. p. 224: see Azara, 'Des Quadrupèdes du Paraguay,' tom. ii. p. 313. The French translator of Azara refers to other cases mentioned by Huzard as occurring in Spain.

[108] Godron, 'De l'Espèce,' tom i. p. 378.

[109] 'Ueber die Eigenschaften,' &c., 1828, s. 10.

[110] 'Domesticated Animals of the British Islands,' pp. 527, 532. In all the veterinary treatises and papers which I have read, the writers insist in the strongest terms on the inheritance by the horse of all good and bad tendencies and qualities. Perhaps the principle of inheritance is not really stronger in the horse than in any other animal; but, from its value, the tendency has been more carefully observed.

[111] Andrew Knight crossed breeds so different in size as a dray-horse and Norwegian pony: see A. Walker on 'Intermarriage,' 1838, p. 205.

[112] 'Naturalist's Library,' Horses, vol. xii. p. 208.

[113] Gervais, 'Hist Nat. Mamm.,' tom. ii. p. 143. Owen, 'British Fossil Mammals,' p. 383.

[114] 'Kenntniss der fossilen Pferde,' 1863, s. 131.

[115] Mr. W. C. L. Martin ('The Horse,' 1845, p. 34), in arguing against the belief that the wild Eastern horses are merely feral, has remarked on the improbability of man in ancient times having extirpated a species in a region where it can now exist in numbers.

[116] 'Transact. Maryland Academy,' vol. i. part i. p. 28.

[117] Mr. Mackinnon on 'The Falkland Islands,' p. 25. The average height of the Falkland horses is said to be 14 hands 2 inches. See also my 'Journal of Researches.'

[118] Pallas, 'Act. Acad. St. Petersburgh,' 1777, part ii. p. 265. With respect to the tarpans scraping away the snow, see Col. Hamilton Smith in 'Nat. Lib.,' vol. xii. p. 165.

[119] Franklin's 'Narrative,' vol. i. p. 87; note by Sir J. Richardson.

[120] Mr. J. H. Moor, 'Notices of the Indian Archipelago:' Singapore, 1837, p. 189. A pony from Java was sent ('Athenæum,' 1842, p. 718) to the Queen only 28 inches in height. For the Loo Choo Islands, see Beechey's 'Voyage,' 4th edit., vol. i. p. 499.

[121] J. Crawford, 'History of the Horse;' 'Journal of Royal United Service Institution,' vol. iv.

[122] 'Essays on Natural History,' 2nd series, p. 161.

[123] 'Quadrupèdes du Paraguay,' tom. ii. p. 333.

[124] Prof. Low, 'Domesticated Animals,' p. 546. With respect to the writer in India, see 'India Sporting Review,' vol. ii. p. 181. As Lawrence has remarked ('The Horse,' p. 9), "perhaps no instance has ever occurred of a three-part bred horse (i.e. a horse, one of whose grand-parents was of impure blood) saving his distance in running two miles with thoroughbred racers." Some few instances are on record of seven-eighths racers having been successful.

[125] Prof. Gervais (in his 'Hist. Nat. Mamm.,' tom. ii. p. 144) has collected many facts on this head. For instance, Solomon (Kings, b. i. ch. x. v. 28) bought horses in Egypt at a high price.

[126] 'The Field,' July 13th, 1861, p. 42.

[127] E. Vernon Harcourt, 'Sporting in Algeria,' p. 26.

[128] I state this from my own observations made during several years on the colours of horses. I have seen cream-coloured, light-dun and mouse-dun horses dappled, which I mention because it has been stated (Martin, 'History of the Horse,' p. 134) that duns are never dappled. Martin (p. 205) refers to dappled asses. In 'The Farrier' (London, 1828, pp. 453, 455) there are some good remarks on the dappling of horses; and likewise in Col. Hamilton Smith on 'The Horse.'

[129] Some details are given in 'The Farrier,' 1828, pp. 452, 455. One of the least ponies I ever saw, of the colour of a mouse, had a conspicuous spinal stripe. A small Indian chesnut pony had the same stripe, as had a remarkably heavy chesnut cart-horse. Race-horses often have the spinal stripe.

[130] I have received information, through the kindness of the Consul-General, Mr. J. R. Crowe, from Prof. Boeck, Rasck, and Esmarck, on the colours of the Norwegian ponies. See, also, 'The Field,' 1861, p. 431.

[131] Col. Ham. Smith, 'Nat. Lib.,' vol. xii. p. 275.

[132] Mr. G. Clark, in 'Annal and Mag. of Nat. History,' 2nd series, vol. ii., 1848, p. 363. Mr. Wallace informs me that he saw in Java a dun and clay-coloured horse with spinal and leg stripes.

[133] See, also, on this point, 'The Field,' July 27th, 1861, p. 91.

[134] 'The Field,' 1861, pp. 431, 493, 545.

[135] 'Ueber die Eigenschaften,' &c, 1828, s. 13, 14.

[136] 'Naturalist's Library,' vol. xii. (1841), pp. 109, 156 to 163, 280, 281. Cream-colour, passing into Isabella (i.e. the colour of the dirty linen of Queen Isabella), seems to have been common in ancient times. See also Pallas's account of the wild horses of the East, who speaks of dun and brown as the prevalent colours.

[137] Azara, 'Quadrupèdes du Paraguay,' tom. ii. p. 307; for the colour of mules, see p. 350. In North America, Catlin (vol. ii. p. 57) describes the wild horses, believed to have descended from the Spanish horses of Mexico, as of all colours, black, grey, roan, and roan pied with sorrel. F. Michaux ('Travels in North America,' Eng. translat., p. 235) describes two wild horses from Mexico as roan. In the Falkland Islands, where the horse has been feral only between 60 and 70 years, I was told that roans and iron-greys were the prevalent colours. These several facts show that horses do not generally revert to any uniform colour.

[138] Dr. Sclater, in 'Proc. Zoolog. Soc.,' 1862, p. 164.

[139] W. C. Martin, 'History of the Horse,' 1845, p. 207.

[140] Col. Sykes' Cat. of Mammalia, 'Proc. Zoolog. Soc.,' July 12th, 1831. Williamson, 'Oriental Field Sports,' vol. ii., quoted by Martin, p. 206.

[141] Blyth, in 'Charlesworth's Mag. of Nat. Hist.,' vol. iv., 1840, p. 83. I have also been assured by a breeder that this is the case.

[142] One case is given by Martin, 'The Horse,' p. 205.

[143] 'Journal As. Soc. of Bengal,' vol. xxviii. 1860, p. 231. Martin on the Horse, p. 205.

[144] Hermann von Nathusius, 'Die Racen des Schweines,' Berlin, 1860; and 'Vorstudien fur Geschichte,' &c., 'Schweineschädel,' Berlin, 1864. Rütimeyer, 'Die Fauna der Pfahlbauten,' Basel, 1861.

[145] Nathusius, 'Die Racen des Schweines,' Berlin, 1860. An excellent appendix is given with references to published and trustworthy drawings of the breeds of each country.

[146] For Europe, see Bechstein, 'Naturgesch. Deutschlands,' 1801, b. i., s. 505. Several accounts have been published on the fertility of the offspring from wild and tame swine. See Burdach's 'Physiology,' and Godron, 'De l'Espèce,' tom. i. p. 370. For Africa, 'Bull. de la Soc. d'Acclimat.,' tom. iv. p. 389. For India, see Nathusius, 'Schweineschädel,' s. 148.

[147] Sir W. Elliot, Catalogue of Mammalia, 'Madras Journal of Lit. and Science,' vol. x. p. 219.

[148] 'Pfahlbauten,' s. 163 et passim.

[149] See Rütimeyer's Neue Beitrage, ... Torfschweine, Verh. Naturfor. Gesell. in Basel, iv. i., 1865, s. 139.

[150] Stan. Julien, quoted by De Blainville, 'Ostéographie,' p. 163.

[151] Richardson, 'Pigs, their Origin,' &c., p. 26.

[152] 'Die Racen des Schweines,' s. 47, 64.

[153] 'Proc. Zoolog. Soc.,' 1861, p. 263.

[154] Sclater, in 'Proc. Zoolog. Soc.,' Feb. 26th, 1861.

[155] 'Proc. Zoolog. Soc.,' 1862, p. 13.

[156] 'Journal of Voyages and Travels from 1821 to 1829,' vol. i. p. 300.

[157] Rev. G. Low, 'Fauna Orcadensis,' p. 10. See also Dr. Hibbert's account of the pig of the Shetland Islands.

[158] 'Die Racen des Schweines,' s. 70.

[159] These woodcuts are copied from engravings given in Mr. S. Sidney's excellent edition of 'The Pig,' by Youatt, 1860. See pp. 1, 16, 19.

[160] 'Schweineschädel,' s. 74, 135.

[161] Nathusius, 'Die Racen des Schweines,' s. 71.

[162] 'Die Racen des Schweines,' s. 47. 'Schweineschädel,' s. 104. Compare, also, the figures of the old Irish and the improved Irish breeds in Richardson on 'The Pig,' 1847.

[163] Quoted by Isid. Geoffroy, 'Hist. Nat. Gén.,' tom. iii. p. 441.

[164] S. Sidney, 'The Pig,' p. 61.

[165] 'Schweineschädel,' s. 2, 20.

[166] 'Proc. Zoolog. Soc.,' 1837, p. 23. I have not given the caudal vertebræ, as Mr. Eyton says some might possibly have been lost. I have added together the dorsal and lumbar vertebræ, owing to Prof. Owen's remarks ('Journal Linn. Soc.,' vol. ii. p. 28) on the difference between dorsal and lumbar vertebræ depending only on the development of the ribs. Nevertheless the difference in the number of the ribs in pigs deserves notice.

[167] 'Edinburgh New Philosoph. Journal,' April 1863. See also De Blainville's 'Ostéographie,' p. 128, for various authorities on this subject.

[168] Eudes-Deslongchamps, 'Mémoires de la Soc. Linn. de Normandie,' vol. vii., 1842, p. 41. Richardson, 'Pigs, their Origin, &c.,' 1847, p. 30. Nathusius, 'Die Racen des Schweines,' 1860, s. 54.

[169] D. Johnson's 'Sketches of Indian Field Sports,' p. 272. Mr. Crawfurd informs me that the same fact holds good with the wild pigs of the Malay peninsula.

[170] For Turkish pigs, see Desmarest, 'Mammalogie,' 1820, p. 391. For those of Westphalia, see Richardson's 'Pigs, their Origin,' &c., 1847, p. 41.

[171] With respect to the several foregoing and following statements on feral pigs, see Roulin, in 'Mém. présentés par divers Savans à l'Acad.,' &c., Paris, tom. vi., 1835, p. 326. It should be observed that his account does not apply to truly feral pigs; but to pigs long introduced into the country and living in a half-wild state. For the truly feral pigs of Jamaica, see Gosse's 'Sojourn in Jamaica,' 1851, p. 386; and Col. Hamilton Smith, in 'Nat. Library,' vol. ix. p. 93. With respect to Africa, see Livingstone's 'Expedition to the Zambesi,' 1865, p. 153. The most precise statement with respect to the tusks of the West Indian feral boars is by P. Labat (quoted by Roulin); but this author attributes the state of these pigs to descent from a domestic stock which he saw in Spain. Admiral Sulivan, R.N., had ample opportunities of observing the wild pigs on Eagle Islet in the Falklands; and he informs me that they resembled wild boars with bristly ridged backs and large tusks. The pigs which have run wild in the province of Buenos Ayres (Rengger, 'Säugethiere,' s. 331) have not reverted to the wild type. De Blainville ('Ostéographie,' p. 132) refers to two skulls of domestic pigs sent from Patagonia by Al. d'Orbigny, and he states that they have the occipital elevation of the wild European boar, but that the head altogether is "plus courte et plus ramassée." He refers, also, to the skin of a feral pig from North America, and says, "il ressemble tout à fait à un petit sanglier, mais il est presque tout noir, et peut-être un peu plus ramassé dans ses formes."

[172] Gosse's 'Jamaica,' p. 386, with a quotation from Williamson's 'Oriental Field Sports.' Also Col. Hamilton Smith, in 'Naturalist's Library,' vol. ix. p. 94.

[173] S. Sidney's edition of 'Youatt on the Pig,' 1860, pp. 7, 26, 27, 29, 30.

[174] 'Schweineschädel,' s. 140.

[175] 'Die Fauna der Pfahlbauten,' 1861, s. 109, 149, 222. See also Geoffroy Saint Hilaire, in 'Mém. du Mus. d'Hist. Nat.,' tom. x. p. 172; and his son Isidore, in 'Hist. Nat. Gén.,' tom. iii. p. 69. Vasey, in his 'Delineations of the Ox Tribe,' 1851, p. 127, says the zebu has four, and the common ox five, sacral vertebræ. Mr. Hodgson found the ribs either thirteen or fourteen in number; see a note in 'Indian Field,' 1858, p. 62.

[176] 'The Indian Field,' 1858, p. 74, where Mr. Blyth gives his authorities with respect to the feral humped cattle. Pickering, also, in his 'Races of Man,' 1850, p. 274, notices the peculiar character of the grunt-like voice of the humped cattle.

[177] Mr. H. E. Marquand, in 'The Times,' June 23rd, 1856.

[178] Vasey, 'Delineations of the Ox-Tribe,' p. 124. Brace's 'Hungary,' 1851, p. 94. The Hungarian cattle descend, according to Rütimeyer ('Zahmen. Europ. Rindes,' 1866, s. 13), from Bos primigenius.

[179] Moll and Gayot, 'La Connaissance Gén. du Bœuf,' Paris, 1860. Fig 82 is that of the Podolian breed.

[180] A translation appeared in three parts in the 'Annals and Mag. of Nat. Hist.,' 2nd series, vol. iv., 1849.

[181] See, also, Rütimeyer's 'Beitrage pal. Gesch. der Wiederkauer,' Basel, 1865, s. 54.

[182] Pictet's 'Paléontologie,' tom. i. p. 365 (2nd edit.). With respect to B. trochoceros, see Rütimeyer's 'Zahmen Europ. Rindes,' 1866, s. 26.

[183] Owen, 'British Fossil Mammals,' 1846, p. 510.

[184] 'British Pleistocene Mammalia,' by W. B. Dawkins and W. A. Sandford, 1866. p. xv.

[185] W. R. Wilde, 'An Essay on the Animal Remains, &c., Royal Irish Academy,' 1860, p. 29. Also 'Proc. of R. Irish Academy,' 1858, p. 48.

[186] 'Lecture: Royal Institution of G. Britain,' May 2nd, 1856, p. 4. 'British Fossil Mammals,' p. 513.

[187] Nilsson, in 'Annals and Mag. of Nat. Hist.,' 1849, vol. iv. p. 354.

[188] See W. R. Wilde, ut supra; and Mr. Blyth, in 'Proc. Irish Academy,' March 5th, 1864.

[189] Laing's 'Tour in Norway,' p. 110.

[190] Isid. Geoffroy St. Hilaire, 'Hist. Nat. Gén.,' tom. iii. p. 96.

[191] Idem, tom. iii. pp. 82, 91.

[192] 'Quadrupèdes du Paraguay,' tom. ii. p. 360.

[193] Walther, 'Das Rindvieh,' 1817, s. 30.

[194] I am much indebted to the present Earl of Tankerville for information about his wild cattle; and for the skull which was sent to Prof. Rütimeyer. The fullest account of the Chillingham cattle is given by Mr. Hindmarsh, together with a letter by the late Lord Tankerville, in 'Annals and Mag. of Nat. Hist.,' vol. ii., 1839, p. 274. See Bewick, 'Quadrupeds,' 2nd edit., 1791, p. 35, note. With respect to those of Duke of Queensberry, see Pennant's 'Tour in Scotland,' p. 109. For those of Chartley, see Low's 'Domesticated Animals of Britain,' 1845, p. 238. For those of Gisburne, see Bewick's 'Quadrupeds, and Encyclop. of Rural Sports,' p. 101.

[195] Boethius was born in 1470; 'Annals and Mag. of Nat. Hist.,' vol. ii., 1839, p. 281; and vol. iv. 1849, p. 424.

[196] Youatt on Cattle, 1834, p. 48: See also p. 242, on short-horn cattle. Bell, in his 'British Quadrupeds,' p. 423, states that, after long attending to the subject, he has found that white cattle invariably have coloured ears.

[197] Azara, 'Des Quadrupèdes du Paraguay,' tom. ii. p. 361. Azara quotes Buffon for the feral cattle of Africa. For Texas, see 'Times,' Feb. 18th, 1846.

[198] Anson's Voyage. See Kerr and Porter's 'Collection,' vol. xii. p. 103.

[199] See also Mr. Mackinnon's pamphlet on the Falkland Islands, p. 24.

[200] 'The Age of the Ox, Sheep, Pig,' &c., by Prof. James Simonds, published by order of the Royal Agricult. Soc.

[201] 'Ann. Agricult. France,' April 1897. as quoted in 'The Veterinary,' vol. xii. p. 725. I quote Tessier's observations from Youatt on Cattle, p. 527.

[202] 'The Veterinary,' vol. viii. p. 681, and vol. x. p. 268. Low's 'Domest. Animals of Great Britain,' p. 297.

[203] Mr. Ogleby, in 'Proc. Zoolog. Soc.,' 1836, p. 138, and 1840, p. 4.

[204] Leguat's Voyage, quoted by Vasey in his 'Delineations of the Ox-tribe,' p. 132.

[205] 'Travels in South Africa,' pp. 317, 336.

[206] 'Mém. de l'Institut présent. par divers Savans,' tom. vi., 1835, p. 333. For Brazil, see 'Comptes Rendus,' June 15th, 1846. See Azara, 'Quadrupèdes du Paraguay,' tom. ii. pp. 359, 361.

[207] 'Schweineschädel,' 1864, s. 104. Nathusius states that the form of skull characteristic of the niata cattle occasionally appears in European cattle; but he is mistaken, as we shall hereafter see, in supposing that these cattle do not form a distinct race. Prof. Wyman, of Cambridge, United States, informs me that the common cod-fish presents a similar monstrosity, called by the fishermen the "bulldog cod." Prof. Wyman also concluded, after making numerous inquiries in La Plata, that the niata cattle transmit their peculiarities or form a race.

[208] Ueber Art des Zahmen Europ. Rindes, 1866, s. 28.

[209] 'Descriptive Cat. of Ost. Collect. of College of Surgeons,' 1853, p. 624. Vasey, in his 'Delineations of the Ox-tribe,' has given a figure of this skull; and I sent a photograph of it to Prof. Rütimeyer.

[210] Loudon's 'Magazine of Nat. Hist.,' vol. i., 1829, p. 113. Separate figures are given of the animal, its hoofs, eye, and dewlap.

[211] Low, 'Domesticated Animals of the British Isles,' p. 264.

[212] 'Mém. de l'Institut présent. par divers Savans,' tom. vi., 1835, p. 332.

[213] Idem, pp. 304, 368, &c.

[214] Youatt on Cattle, p. 193. A full account of this bull is taken from Marshall.

[215] Youatt on Cattle, p. 116. Lord Spencer has written on this same subject.

[216] Blyth on the genus Ovis, in 'Annals and Mag. of Nat. History,' vol. vii., 1841, p. 261: with respect to the parentage of the breeds, see Mr. Blyth's excellent articles in 'Land and Water,' 1867, pp. 134, 156. Gervais, 'Hist. Nat. des Mammifères,' 1855, tom. ii. p. 191.

[217] Dr. L. Fitzinger, 'Ueber die Racen des Zahmen Schafes,' 1860, s. 86.

[218] J. Anderson, 'Recreations in Agriculture and Natural History,' vol. ii. p. 164.

[219] 'Pfahlbauten,' s. 127, 193.

[220] Youatt on Sheep, p. 120.

[221] 'Journal of the Asiatic Soc. of Bengal,' vol. xvi. pp. 1007, 1016.

[222] Youatt on Sheep, pp. 142-169.

[223] 'Journal Asiat. Soc. of Bengal,' vol. xvi., 1847, p. 1015.

[224] 'Hist. Nat. Gén.,' tom. iii. p. 435.

[225] Youatt on Sheep, p. 138.

[226] 'Journal Asiat. Soc. of Bengal,' vol. xvi., 1847, pp. 1015, 1016.

[227] 'Racen des Zahmen Schafes,' s. 77.

[228] 'Rural Economy of Norfolk,' vol. ii. p. 136.

[229] Youatt on Sheep, p. 312. On same subject, see excellent remarks in 'Gardener's Chronicle,' 1858, p. 868. For experiments in crossing Cheviot sheep with Leicesters, see Youatt, p. 325.

[230] Youatt on Sheep, note, p. 491.

[231] 'The Veterinary,' vol. x. p. 217.

[232] A translation of his paper is given in 'Bull. Soc. Imp. d'Acclimat.,' tom. ix., 1862, p. 723.

[233] Erman's 'Travels in Siberia' (Eng. trans.), vol. i. p. 228. For Pallas on the fat-tailed sheep, I quote from Anderson's account of the 'Sheep of Russia,' 1794, p. 34. With respect to the Crimean sheep, see Pallas' 'Travels' (Eng. trans.), vol. ii. p. 454. For the Karakool sheep, see Burnes' 'Travels in Bokhara,' vol. iii. p. 151.

[234] See Report of the Directors of the Sierra Leone Company, as quoted in White's 'Gradation of Man,' p. 95. With respect to the change which sheep undergo in the West Indies, see also Dr. Davy, in 'Edin. New. Phil. Journal,' Jan. 1852. For the statement made by Roulin, see 'Mém. de l'Institut présent. par divers Savans,' tom. vi., 1835, p. 347.

[235] Youatt on Sheep, p. 69, where Lord Somerville is quoted. See p. 117, on the presence of wool under the hair. With respect to the fleeces of Australian sheep, p. 185. On selection counteracting any tendency to change, see pp. 70, 117, 120, 168.

[236] Audubon and Bachman, 'The Quadrupeds of North America,' 1846, vol. v. p. 365.

[237] 'Journal of R. Agricult. Soc. of England,' vol. xx., part ii. W. C. Spooner on Cross-Breeding.

[238] 'Philosoph. Transactions,' London, 1813, p. 88.

[239] Isidore Geoffroy St. Hilaire, 'Hist. Nat. Générale,' tom. iii. p. 87. Mr. Blyth ('Land and Water,' 1867, p. 37) has arrived at a similar conclusion, but he thinks that certain Eastern races may perhaps be in part descended from the Asiatic markhor.

[240] Rütimeyer, 'Pfahlbauten,' s. 127.

[241] Godron, 'De l'Espèce,' tom. i. p. 402.

[242] 'Annals and Mag. of Nat. History,' vol. ii. (2nd series), 1848, p. 363.

[243] 'De l'Espèce,' tom. i. p. 406. Mr. Clark also refers to differences in the shape of the mammæ. Godron states that in the Nubian race the scrotum is divided into two lobes; and Mr. Clark gives a ludicrous proof of this fact, for he saw in the Mauritius a male goat of the Muscat breed purchased at a high price for a female in full milk. These differences in the scrotum are probably not due to descent from distinct species; for Mr. Clark states that this part varies much in form.

[244] Mr. Clark, 'Annals and Mag. of Nat. Hist.,' vol. ii. (2nd series), 1848, p. 361.

[245] Desmarest, 'Encyclop. Méthod. Mammalogie,' p. 480.

[246] 'Journal of Asiatic Soc. of Bengal,' vol. xvi., 1847, pp. 1020, 1025.