Dimensions.—Diameter of the outer shell 0.12, middle 0.06, inner 0.03; cortical pores 0.012, bars 0.003; length of the six spines 0.08, breadth 0.03.
Habitat.—Central Pacific, Station 266, depth 2750 fathoms.
14. Hexacontium hexaconicum, n. sp.
Cortical shell spiny, thick walled. Pores regular circular, of the same breadth as the bars, ten to twelve on the radius; between them long bristle-shaped by-spines, half as long as the six main spines. Radial proportion of the three spheres = 1 : 3 : 9. Six main spines conical, as long as the radius of the outer shell, four times as broad at the base as one pore.
Dimensions.—Diameter of the outer shell 0.18, middle 0.06, inner 0.02; cortical pores and bars 0.005; length of the spines 0.1, basal breadth 0.02.
Habitat.—Antarctic Sea, Station 157, depth 1950 fathoms.
15. Hexacontium asteracanthion, Haeckel.
Haliomma asteracanthion, Haeckel, 1860, Monatsber. d. k. preuss. Akad. d. Wiss. Berlin, p. 816.
Actinomma asteracanthion, Haeckel, 1862, Monogr. d. Radiol., p. 441, Taf. xxiii. figs. 5, 6.
Cortical shell thin walled, spiny; its pores circular, without hexagonal frame (as a rule very regular, but in other specimens more or less irregular), often very variable in size, three to six times as broad as the bars, commonly seven to eight on the radius. Radial proportion of the three spheres = 1 : 2 : 4, or sometimes 1 : 3 : 8. Pores of the two inner shells also circular, but much smaller; those of the middle shell about half as broad, those of the inner shell one-fourth to one-sixth as broad as the pores of the outer shell. Six radial spines strong, three-sided prismatic, about as long as (or somewhat longer than) the radius of the outer shell, as broad as one of its large meshes. Accessory spines very numerous and very thin, bristle-like, usually half as long as the six main spines. (For the variability of this common species compare my Monograph, p. 442.)
Dimensions.—Diameter of the outer shell 0.1 to 0.12, of the middle 0.04 to 0.06, of the inner 0.02 to 0.03; cortical pores 0.005 to 0.01; length of the six spines 0.06 to 0.08.
Habitat.—Cosmopolitan; Mediterranean, Atlantic, Pacific, surface.
Definition.—Pores of the cortical shell irregular, of different size or form; surface smooth, without radial by-spines or papillæ (other than the six main spines).
16. Hexacontium axophænum, n. sp.
Cortical shell thin walled, smooth. Pores irregular polygonal, three to nine times as broad as the bars; seven to ten on the radius. Radial proportion of the three spheres = 1 : 3 : 8. Both medullary shells with smaller, irregular roundish pores. Six spines three-sided pyramidal, longer than the radius of the outer shell, one to two times as broad as one larger pore.
Dimensions.—Diameter of the outer shell 0.12, middle 0.045, inner 0.015; cortical pores 0.006 to 0.018, bars 0.002; length of the spines 0.06 to 0.12, basal breadth 0.02 to 0.03.
Habitat.—Central Pacific, Stations 266 to 272, at various depths.
17. Hexacontium polygonale, n. sp.
Cortical shell thick walled, smooth. Pores irregular roundish, with polygonal frames, two to four times as broad as the bars; eight to twelve on the radius. Radial proportion of the three spheres = 1 : 3 : 10. Six spines pyramidal, nearly as long as the diameter of the outer shell, with six to nine prominent edges (along the corner number of the polygonal meshes).
Dimensions.—Diameter of the outer shell 0.2, middle 0.06, inner 0.02; cortical pores 0.008 to 0.016, bars 0.004; length of the spines 0.16 to 0.18, basal breadth 0.02.
Habitat.—South Pacific, Station 295, depth 1500 fathoms.
18. Hexacontium antarcticum, n. sp.
Cortical shell thin walled, smooth, with irregular roundish pores, eight to ten on the radius, two to four times as broad as the bars. Radial proportion of the three spheres = 1 : 3 : 9. Both medullary shells with smaller roundish irregular pores. Six spines conical, pyramidal at their origin, about as long as the radius, as broad as one larger mesh.
Dimensions.—Diameter of the outer shell 0.18, middle 0.06, inner 0.02; cortical pores 0.006 to 0.012, bars 0.003; length of the spines 0.1, basal breadth 0.12.
Habitat.—Antarctic Ocean, Station 157, depth 1950 fathoms.
Definition.—Pores of the cortical shell irregular, of different size or form; surface covered with numerous conical papillæ or bristle-shaped, radial by-spines (other than the six main spines).
19. Hexacontium papillosum, n. sp. (Pl. 24, fig. 5).
Cortical shell thick walled, papillose. Pores irregular roundish, often somewhat lobed, five to six on the radius, two to four times as broad as the bars. Radial proportion of the three spheres = 1 : 2.5 : 5. Both medullary shells with subregular circular pores (inner much smaller). Six radial spines short and stout, pommel or club shaped, with three prominent wings, about as long as the shell radius, and as broad as the medullary shell.
Dimensions.—Diameter of the outer shell 0.1, middle 0.05, inner 0.02, cortical pores 0.006 to 0.012, bars 0.003; length of the spines 0.05, bars 0.02.
Habitat.—Central Pacific, Stations 266 to 272, at various depths.
20. Hexacontium gladiatum, n. sp. (Pl. 25, fig. 8).
Cortical shell very thick walled, spiny, with irregular network; its meshes roundish, two to five times as broad as the bars, of very different size, ten to fifteen on the half meridian. Bars between them very strong, three-sided prismatic, armed with a great number of small thorns, and with large spines at the nodal-points between every three meshes. Radial proportion of the three spheres = 1 : 3 : 12. Six strong radial main spines three-sided prismatic, with three prominent, somewhat contorted wings, acute, broader than the diameter of the inner shell and as long as the diameter of the outer shell. These six main spines are situated in the same three dimensive axes as the six thin radial beams connecting the two medullary shells. But the six radial beams which connect the middle with the outer shell alternate with the former and lie in three other dimensive axes,—a very rare and remarkable disposition.
Dimensions.—Diameter of the outer shell 0.2, middle 0.05, inner 0.016; cortical pores 0.01 to 0.03, bars 0.006; length of the six spines 0.2, breadth 0.02 to 0.03.
Habitat.—Tropical Atlantic, Station 342, depth 1445 fathoms.
21. Hexacontium setosum, n. sp.
Cortical shell thin walled, with numerous bristle-shaped, simple, radial by-spines of variable length. Pores irregular roundish, five to seven on the radius, two to eight times as broad as the bars. Radial proportion of the three spheres = 1 : 3 : 9. Main spines three-sided pyramidal, as long as the radius.
Dimensions.—Diameter of the outer shell 0.1 to 0.015, middle 0.035 to 0.05, inner 0.013 to 0.016; cortical pores 0.01 to 0.03, bars 0.003 to 0.004; length of the six spines 0.05 to 0.08, basal breadth 0.012 to 0.015.
Habitat.—Cosmopolitan; Mediterranean, Atlantic, Pacific, surface; also fossil in Barbados.
22. Hexacontium furcatum, n. sp.
Cortical shell thin walled, covered with numerous thin, bristle-shaped by-spines, which are forked and nearly as long as the main spines. Pores irregular roundish, eight to ten on the radius, two to eight times as broad as the bars. Radial proportion of the three spheres = 1 : 2.5 : 10. Main spines triangular pyramidal, shorter then the radius.
Dimensions.—Diameter of the outer shell 0.18, middle 0.045, inner 0.018; cortical pores 0.005 to 0.015, bars 0.002; length of the six spines 0.07, basal breadth 0.015.
Habitat.—North Atlantic, Station 354, surface.
23. Hexacontium drymodes, Haeckel.
Actinomma drymodes, Haeckel, 1862, Monogr. d. Radiol., p. 442, Taf. xxiv. fig. 9.
Cortical shell thin walled, covered with numerous thin, bristle-shaped spines, which are double forked and half as long as the main spines. Pores irregular roundish, eight to ten on the radius, two to eight times as broad as the bars. Radial proportion of the three spheres = 1 : 2 : 4. Main spines three-sided prismatic, with prominent, often somewhat contorted edges, at the distal end cuspidated; longer than the radius of the outer shell.
Dimensions.—Diameter of the outer shell 0.15, middle 0.07, inner 0.035; cortical pores 0.01 to 0.03, bars 0.04; length of the six spines 0.11, basal breadth 0.03.
Habitat.—Mediterranean (Messina), surface; Canary Islands (Lanzerote); Haeckel.
24. Hexacontium periplectum, n. sp.
Cortical shell double, enclosing a simple medullary shell. Radial proportion of the three shells = 1 : 4 : 5. Inner cortical shell thick walled, with irregular roundish pores, two to eight times as broad as the bars; five to seven on the radius. Numerous radial spines, arising from it, are connected below their distal ends (at equal distances from the centre) by delicate branched threads, and so form an outer, irregular, thin, cortical shell, with spiny surface. Six main spines three-sided pyramidal, about as long as the radius.
Dimensions.—Diameter of the outer shell 0.22, middle 0.2, inner 0.045; cortical pores (of both outer shells) 0.01 to 0.03, bars 0.001 to 0.005; length of the spines 0.1, basal breadth 0.02.
Habitat.—North Pacific, Station 241, surface.
Definition.—Cubosphærida with three concentric lattice-spheres and six branched spines of equal size.
The genus Hexadendron differs from Hexacontium in the ramification of the six dimensive spines, and from the similar Hexancistra in the duplication of the medullary shell. As in the latter instance, each spine can bear either three simple lateral branches or three rows of pinnate lateral branches.
1. Hexadendron quadricuspis, n. sp.
All three shells spherical, with radial proportion = 1 : 2 : 6. Pores of both medullary shells regular circular, twice as broad as the bars. Pores of the cortical shell irregular roundish, four to six times as broad as the bars; surface a little thorny. Six radial spines prismatic, with three prominent wing-like edges, which are prolonged below the distal end in three curved lateral branches. (Differs from Hexalonche quadricuspis, Pl. 22, fig. 11, mainly in the double medullary shell.)
Dimensions.—Diameter of the outer shell 0.15, middle 0.05, inner 0.025; length of the spines 0.12, breadth 0.01.
Habitat.—Central Pacific, Station 272, depth 2600 fathoms.
2. Hexadendron bipinnatum, n. sp. (Pl. 23, fig. 1).
All three shells regular octahedral, with very delicate network of irregular polygonal meshes, and very thin, thread-like bars between them; their radial proportion = 1 : 2.5 : 7.5. Surface of the cortical shell covered with numerous bristle-shaped by-spines, as long as the radius. Six main spines very large, three-sided prismatic, with three rows of pinnate, lateral branches, on the three wing-like, spirally twisted edges (similar to those of Hexancistra mirabilis, p. 189, Pl. 23, fig. 3).
Dimensions.—Diameter of the outer shell 0.12, middle 0.04, inner 0.016.
Habitat.—Central Pacific, Station 272, surface.
Definition.—Cubosphærida with three concentric lattice-spheres and six simple spines of different size; one opposite pair larger than the two others.
The genus Hexacontarium exhibits to its ancestral form Hexacontium the same relation that Hexaloncharium bears to Hexalonche; the former is developed from the latter by duplication of the medullary shell. As two opposite spines of one pair are larger than the four others, they correspond to the three axes of a quadratic crystal.
1. Hexacontarium dentatum, n. sp.
Cortical shell with regular circular, hexagonally framed pores, twice as broad as the bars, with smooth surface. Radial proportion of the three spheres = 1 : 2 : 5. Six radial spines three-sided prismatic, half as broad as the inner medullary shell, with three dentated edges. Two opposite major spines longer than the shell diameter; four other minor spines only half as long as the radius. (Similar to Hexacontium clavigerum, Pl. 23, fig. 5, but distinct by the prolongation of the spines of one axis.)
Dimensions.—Diameter of the outer shell 0.15, middle 0.06, inner 0.03; length of the major spines 0.2, minor 0.04.
Habitat.—Central Pacific, Station 266, depth 2750 fathoms.
2. Hexacontarium clavatum, n. sp.
Cortical shell covered with short conical by-spines and irregular roundish pores, three to four times as broad as the bars. Radial proportion of the three spheres = 1 : 3 : 8. Six radial spines three-sided prismatic, at the distal end club-shaped; two major spines three times as long as the four others, which are equal to the shell radius.
Dimensions.—Diameter of the outer shell 0.16, middle 0.06, inner 0.02.
Habitat.—Central Pacific, Station 272, depth 2600 fathoms.
Definition.—Cubosphærida with four concentric spherical lattice-shells.
Definition.—Cubosphærida with four concentric lattice-spheres and six simple spines of equal size.
The genus Hexacromyum possesses four concentric, spherical, or octahedral lattice-shells; two inner medullary shells within the central capsule, two outer cortical outside it. The four spheres are connected by six radial beams, which are prolonged outside into simple spines of equal size, opposite in pairs in the three dimensive axes. This genus can be derived from Hexacontium by duplication of the cortical shell.
1. Hexacromyum elegans, n. sp. (Pl. 24, fig. 9).
Shell composed of four concentric shells, with radial proportion = 1 : 2.5 : 7.5 : 10. First (innermost) shell with very small circular pores, second shell with larger circular pores. Third shell (inner cortical shell) with large, subregular, circular, hexagonally framed pores (eight to nine on the radius), twice as broad as the bars; from the elevated nodal-points of the hexagonal frames (between every three pores) arise thin bristle-shaped radial beams, which are united at the distal end by vaulted branches forming the delicate fourth shell. Surface smooth. Radial spines three-sided pyramidal, as long as the radius, as broad at the base as the innermost shell.
Dimensions.—Diameter of the four shells—(A) 0.02, (B) 0.05, (C) 0.15, (D) 0.2; length of the six radial spines 0.1, basal breadth 0.02.
Habitat.—Central Pacific, Station 271, surface.
2. Hexacromyum quadrigatum, n. sp.
Shell composed of four concentric shells, with radial proportion = 1 : 3 : 8 : 10. Structure of all four shells the same, with regular, circular pores, twice to three times as broad as the bars; size of the pores gradually increasing from the innermost to the outermost shell; surface smooth. Radial spines three-sided pyramidal, as long as the radius, half as broad at the base as the innermost shell.
Dimensions.—Diameter of the four shells—(A) 0.025, (B) 0.08, (C) 0.20, (D) 0.25; length of the spines 0.12, basal breadth 0.012.
Habitat.—North Pacific, Station 253, depth 3125 fathoms.
3. Hexacromyum arachnoides, n. sp.
Hexacromidium arachnoides, Haeckel, 1881, Prodromus, p. 453.
Shell composed of four concentric shells, with radial proportion = 1 : 2 : 3 : 4. Innermost shell with regular, hexagonal meshes; the three other shells with irregular, polygonal meshes; bars between the large meshes in all four shells very thin, cobweb-like. Surface covered with thin bristle-shaped by-spines, as long as the radius. Six radial main spines three-sided prismatic, longer than the diameter of the whole shell.
Dimensions.—Diameter of the four shells—(A) 0.08, (B) 0.16, (C) 0.24, (D) 0.32; length of the spines 0.4, breadth 0.01.
Habitat.—South Pacific, Station 300, depth 1375 fathoms.
4. Hexacromyum octahedrum, n. sp. (Pl. 23, fig. 2).
Shell composed of four concentric shells which are not spherical (as in the three preceding species), but regular octahedral. Radial proportion = 1 : 2.5 : 6 : 9. Network in all four shells delicate, with irregular polygonal meshes and thin bars; the thickness of the bars and size of the meshes increasing from the innermost to the outermost shell. Six radial spines three-sided prismatic, increasing slowly in breadth towards the distal end, much longer than the shell diameter.
Dimensions.—Diameter of the four shells—(A) 0.02, (B) 0.05, (C) 0.12, (D) 0.18; length of the radial spines 0.2 to 0.3 and more, breadth 0.01.
Habitat.—Central Pacific, Station 263, depth 2650 fathoms.
Definition.—Cubosphærida with five or more concentric lattice-shells.
Definition.—Cubosphærida with five to six or more concentric lattice-shells and six simple spines of equal size.
The genus Cubosphæra is developed from the preceding Hexacromyum by further addition of the concentric lattice-shells, their number amounting to five, six, or more. The innermost two of these are medullary shells, the others being cortical shells. All are connected by six radial beams, prolonged outside into six simple spines of equal size; these lie opposite in pairs in three dimensive axes, corresponding to the three axes of a cube.
1. Cubosphæra cubaxonia, n. sp. (Pl. 24, fig. 8).
Hexacromyon cubaxonium, Haeckel, 1881, Prodrom. et Atlas.
Shell composed of five concentric spheres, with the radial proportion = 1 : 3 : 8 : 10 : 13. The two medullary shells with small regular, circular pores of the same breadth as the bars. Inner cortical shell (third shell) with large regular, circular pores, four times as broad as the bars, hexagonally framed. From each hexagon-corner arises a thin bristle-shaped radial by-spine, which at a fixed distance from the centre is united with the middle cortical shell (fourth shell), which has very small circular pores. The beginning of the fifth shell (outermost) is indicated by six small reticula, produced by the six main spines at equal distances from the centre. All five shells are united by six prismatic radial beams, ending outside in pyramidal furrowed spines.
Dimensions.—Diameter of the five shells—(A) 0.2, (B) 0.15, (C) 0.12, (D) 0.045, (E) 0.015.
Habitat.—Central Pacific, Station 272, depth 2600 fathoms.
2. Cubosphæra concentrica, n. sp.
Shell composed of six concentric spheres, with the radial proportion = 1 : 2 : 6 : 7.5 : 9 : 11. The two medullary shells with regular, circular pores, twice as broad as the bars. The four cortical shells of the same structure, with irregular, roundish pores, three to four times as broad as the bars. The size of these pores and the breadth of their bars gradually increase towards the smooth surface. All six shells are connected by six thin three-sided prismatic radial beams, which are prolonged outside in six stronger spines, angular pyramidal, with smooth edges, as long as the shell radius.
Dimensions.—Diameter of the six shells—(A) 0.02 (B) 0.04, (C) 0.12, (D) 0.15, (E) 0.18, (F) 0.22.
Habitat.—Central Pacific, Station 268, depth 2900 fathoms.
Definition.—Cubosphærida with five to six or more concentric lattice-shells and six branched spines of equal size.
The genus Hexacaryum is distinguished from the foregoing Cubosphæra by ramification of the six radial spines, and therefore exhibits the same relation to it that Hexancistra bears to Hexastylus, &c.
1. Hexacaryum arborescens, n. sp. (Pl. 23, figs. 4, 4a).
Shell composed of five, six, or more concentric shells, which are united by six very large radial spines. The two medullary shells spherical, inner with regular, circular, outer with irregular polygonal pores. All cortical shells (third and following) not spherical, but regular octahedral, with irregular polygonal meshes and thin bars. Radial spines prismatic, with three wing-like, spirally twisted edges, which at equal distances send out thin forked lateral branches (six on each verticil); by further ramification and communication of these branches the triangular net-plates arise, filling out the sides of the octahedral cortical shells. Diameter of all shells little different. The outer free parts of the six spines are arborescent, twice to three times as long as the enclosed inner parts, and bear six to eight verticils of free lateral branches, decreasing in size towards the distal end (similar to Arachnosphæra).
Dimensions.—Diameter of the six shells—(A) 0.015, (B) 0.04, (C) 0.1, (D) 0.16, (E) 0.22, (F) &c.; average distance of the concentric octahedra = 0.06; length of the radial spines 0.3 to 0.4 and more, breadth 0.02.
Habitat.—Central Pacific, Station 274, surface.
Definition.—Cubosphærida with spongy spherical or octahedral shell (with or without enclosed concentrical lattice-shells).
Definition.—Cubosphærida with solid spongy spherical or octahedral shell, without latticed medullary shell in the centre, and with six simple radial spines of equal size.
The genus Cubosphæra may be regarded as a form of Styptosphæra, which develops six radial spines, opposite in pairs in the three dimensive axes. The solid spongy framework of the shell assumes the outer form either of a sphere, or of a regular octahedron.
1. Cubaxonium spongiosum, n. sp.
Spongy shell spherical, composed of a very dense spongy framework of nearly uniform structure; the meshes three to four times as broad as the bars; surface almost smooth. Six spines cylindrical, twice to three times as long as the diameter of the spongy sphere, about as broad as one half mesh.
Dimensions.—Diameter of the sphere 0.2, meshes 0.006 to 0.008, bars 0.002; length of the spines 0.4 to 0.5, breadth 0.004.
Habitat.—South Pacific, Station 288, surface.
2. Cubaxonium octahedrum, n. sp.
Spongy shell octahedral, composed of a loose spongy framework of nearly uniform structure; the meshes ten to twelve times as broad as the bars; surface thorny. Six spines three-sided pyramidal, longer than the diameter of the octahedron, arising from its six corners, as broad at the base as one mesh.
Dimensions.—Diameter of the shell 0.15, meshes 0.01 to 0.012, bars 0.001; length of the spines 0.2, basal breadth 0.01.
Habitat.—South Pacific, Station 300, surface.
Definition.—Cubosphærida with spongy spherical shell and one simple latticed medullary shell in its centre, having six simple spines of equal size.
The genus Hexadoras exhibits the same structure of the spongy shell as Spongoplegma, but differs from it in the production of six dimensive spines. In the centre lies one simple, latticed, medullary shell, which is either spherical or octahedral. In the latter case the six spines arise from the six corners of the octahedron.
1. Hexadoras axophæna, n. sp.
Medullary shell spherical, with regular hexagonal meshes, twice as broad as the bars. Cortical shell entirely enclosing it, with loose irregular framework and thorny surface. Diameter of the outer shell three times as large as that of the inner. Six radial spines arising from the medullary shell, four to five times as long as the radius of the cortical shell, three-sided prismatic, with straight dentated edges.
Dimensions.—Diameter of the outer shell 0.15, inner 0.05; length of the spines 0.3 to 0.4.
Habitat.—Central Pacific, Station 265, depth 2900 fathoms.
2. Hexadoras lychnosphæra, n. sp.
Medullary shell spherical, with regular, circular, hexagonally framed pores, three times as broad as the bars. Cortical shell enveloping it, with loose irregular framework and spiny surface. Diameter of the outer shell eight times as large as that of the inner. Six radial spines arising from the inner shell, three-sided prismatic, with dentated, spirally contorted edges, seven times as long as the diameter of the medullary shell, with three lateral branches at the distal end (similar to Lychnosphæra, Pl. 11, fig. 1).
Dimensions.—Diameter of the outer shell 0.5, inner 0.06; total length of the spines 0.4, breadth 0.01.
Habitat.—Central Pacific, Station 270, depth 2925 fathoms.
3. Hexadoras octahedrum, n. sp.
Medullary shell regular octahedral, with irregular polygonal meshes, five times as broad as the bars. Cortical shell enveloping it, with dense spongy framework, also octahedral with rough surface. Diameter of the outer shell five times as large as that of the inner. Six radial spines very long, arising from the six corners of the inner and piercing the spongy mass of the outer shell, considerably exceeding it at the free distal end, three-sided prismatic, with elegantly denticulate edges.
Dimensions.—Diameter of the outer shell 0.2, inner 0.04; total length of the spines 0.3 or more, breadth 0.02.
Habitat.—Central Pacific, Station 266, depth 2750 fathoms.
Definition.—Cubosphærida with spongy spherical shell and two concentric latticed medullary shells in its centre, having six simple spines of equal size.
The genus Hexadoridium differs from Hexadoras in the duplication of the medullary shell, and exhibits therefore the same relation to it that Spongodictyon bears to Spongoplegma. In the only known species the six spines are opposite, arranged quite regularly in pairs in the three dimensive axes, and consequently represent the three axes of a regular crystal or cube.
1. Hexadoridium streptacanthum, n. sp. (Pl. 25, figs. 1, 1a).
Both medullary shells spherical, with small regular, circular pores, twice as broad as the bars; outer twice as broad as the inner. Spongy cortical shell enclosing it with dense framework, five times as broad as the outer medullary shell, regular octahedral. Six radial spines, arising from the latter, are thinned at the inner end, three to five times as long as the diameter of the cortical shell, and nearly as broad as the inner medullary shell, with three dentated and spirally contorted edges. (Very similar to the common Spongosphæra streptacantha, with irregular and variable number and dispositions of spines; possibly its ancestral form?).
Dimensions.—Diameter of the cortical shell 0.2, of the outer medullary shell 0.04, inner 0.02; length of the spines 1 mm. and more, breadth 0.02.
Habitat.—Central Pacific, Station 271, depth 2425 fathoms.
Astrosphærida, Haeckel, 1881, Prodromus, p. 449.
Definition.—Sphæroidea with numerous (eight to twelve or more, commonly between twenty and sixty) radial spines on the surface of the spherical shell; living solitary (not associated in colonies).
The family Astrosphærida, the largest and most varied of all Sphæroidea, is distinguished from the other members of this group by the possession of numerous radial spines, which are either regularly or irregularly disposed on the surface of the spherical shell. The extreme variability and richness of form in this family is mainly due to the different size, shape, and disposition of these radial spines.
The simplest Astrosphærida are the Coscinommida, with a single spherical or polyhedral lattice-shell. To this ancestral group all other subfamilies can be opposed as "Astrosphærida composita," since their skeleton is composed of two or more concentric lattice-shells: two in the Haliommida, three in the Actinommida, four in the Cromyommida, five or more in the Caryommida. In these four subfamilies the concentric shells are all simple (not spongy) fenestrated spheres or endospherical polyhedra. In the sixth subfamily, the Spongiommida, the shell is wholly or partially composed of spongy irregular wicker-work, with or without a medullary shell in the centre.
The Number of the Radial Spines in the Astrosphærida is extremely variable, and ranges from eight to forty or more; in many cases more than one hundred. Often each nodal-point of the network develops on the shell surface one spine. Still more frequently the number of the spines is less than that of the nodal-points. In all concentric Astrosphærida, having two or more concentrical shells, we can distinguish "primary spines," as outer prolongations of the inner radial beams connecting the shells, and "secondary spines," developed only on the outer surface of the shell. Naturally the former are of much greater importance than the latter. But we can also often distinguish among the latter larger "main spines" and smaller "by-spines," the latter commonly much more numerous than the former.
The Disposition of the Radial Spines, either regular or irregular, is a subject of great morphological interest, and remains to be exhausted by further observations. The following cases of regular disposition have been observed by me—(A) eight spines, opposite in pairs in four axes corresponding to the four diagonal axes of a cube; (B) nine spines, regularly disposed at equal distances (?) (not opposed in pairs); (C) ten spines, disposed at equal distances (?); (D) twelve spines, regularly disposed, corresponding to the twelve corners of the regular icosahedron; (E) fourteen spines, quite regularly disposed (six corresponding to the three axes of a regular octahedron, eight to the central points of its eight faces); (F) sixteen spines, regularly disposed (?); (G) twenty spines (very common!), either disposed in the same manner (after the law of Johannes Müller) as in the Acantharia (?), or corresponding to the twenty corners of the regular or pentagonal dodecahedron, or disposed in the same manner as in many Larcoidea (Tholonida, &c., to be described afterwards); (H) twenty-four spines, regularly disposed (?); (I) thirty-two spines, quite regularly disposed (twenty corresponding to the twenty corners of the regular dodecahedron, twelve to the central points of its twelve faces); (K) forty spines, nearly regularly (or quite symmetrically?) disposed. If the number of the spines amounts to more than forty, it is as a rule impossible to determine their regular disposition in a satisfactory manner.
|
I. Subfamily Coscinommida. (One single latticed shell.) |
brace | Spines all simple, not branched and not tubulous. | brace | All spines of the same kind, | 088. Acanthosphæra. |
| Larger main spines and smaller by-spines, | 089. Heliosphæra. | ||||
| Spines hollow conical tubes with porous walls, | 090. Conosphæra. | ||||
| Between simple spines the pores prolonged in hollow tubes, | 091. Coscinomma. | ||||
| Spines branched or forked. | brace | Spines with lateral branches, | 092. Cladococcus. | ||
| Spines forked or dichotomous, | 093. Elaphococcus. | ||||
|
II. Subfamily Haliommida. (Two concentric latticed shells.) |
brace | A. Elatommida, one medullary and one cortical shell. | brace | All spines equal, simple, | 094. Haliomma. |
| All spines simple, of two different kinds, | 095. Heliosoma. | ||||
| Spines branched, | 096. Elatomma. | ||||
| B. Diplosphærida, both shells cortical. | brace | No by-spines, | 097. Leptosphæra. | ||
| Inner by-spines, | 098. Diplosphæra. | ||||
| Outer by-spines, | 099. Drymosphæra. | ||||
| Outer and inner by-spines, | 100. Astrosphæra. | ||||
|
III. Subfamily Actinommida. (Three concentric latticed shells.) |
brace | Radial spines not branched. | brace | All spines equal, | 101. Actinomma. |
| Larger main spines and smaller by-spines, | 102. Echinomma. | ||||
| Radial spines branched or forked, | 103. Pityomma. | ||||
|
IV. Subfamily Cromyommida. (Three concentric latticed shells.) |
brace | Radial spines not branched. | brace | All spines equal, | 104. Cromyomma. |
| Larger main spines and smaller by-spines, | 105. Cromyechinus. | ||||
| Radial spines branched or forked, | 106. Cromyodrymus. | ||||
|
V. Subfamily Caryommida. (Numerous, five to ten or more, concentric latticed shells.) |
brace | Two medullary shells and three or more cortical shells (lattice work ordinary), | 107. Caryomma. | ||
| No medullary shells (lattice work arachnoidal); three-sided prismatic spines, with verticils of three forked branches. (Arachnosphærida.) | brace | Triangular meshes, simple bars, without diagonal bars, | 108. Arachnopila. | ||
| Triangular meshes, simple bars; diagonal bars between the shells, | 109. Arachnopegma. | ||||
| Polygonal meshes, bars branched, | 110. Arachnosphæra. | ||||
| I. Subfamily Coscinommida. (One single latticed shell.) | ||||||
| Spines all simple, not branched and not tubulous. | ||||||
| All spines of the same kind, | ||||||
| 88. Acanthosphæra. | ||||||
| Larger main spines and smaller by-spines, | ||||||
| 89. Heliosphæra. | ||||||
| Spines hollow conical tubes with porous walls, | ||||||
| 90. Conosphæra. | ||||||
| Between simple spines the pores prolonged in hollow tubes, | ||||||
| 91. Coscinomma. | ||||||
| Spines branched or forked. | ||||||
| Spines with lateral branches, | ||||||
| 92. Cladococcus. | ||||||
| Spines forked or dichotomous, | ||||||
| 93. Elaphococcus. | ||||||
| II. Subfamily Haliommida. (Two concentric latticed shells.) | ||||||
| A. Elatommida, one medullary and one cortical shell. | ||||||
| All spines equal, simple, | ||||||
| 94. Haliomma. | ||||||
| All spines simple, of two different kinds, | ||||||
| 95. Heliosoma. | ||||||
| Spines branched, | ||||||
| 96. Elatomma. | ||||||
| B. Diplosphærida, both shells cortical. | ||||||
| No by-spines, | ||||||
| 97. Leptosphæra. | ||||||
| Inner by-spines, | ||||||
| 98. Diplosphæra. | ||||||
| Outer by-spines, | ||||||
| 99. Drymosphæra. | ||||||
| Outer and inner by-spines, | ||||||
| 100. Astrosphæra. | ||||||
| III. Subfamily Actinommida. (Three concentric latticed shells.) | ||||||
| Radial spines not branched. | ||||||
| All spines equal, | ||||||
| 101. Actinomma. | ||||||
| Larger main spines and smaller by-spines, | ||||||
| 102. Echinomma. | ||||||
| Radial spines branched or forked, | ||||||
| 103. Pityomma. | ||||||
| IV. Subfamily Cromyommida. (Three concentric latticed shells.) | ||||||
| Radial spines not branched. | ||||||
| All spines equal, | ||||||
| 104. Cromyomma. | ||||||
| Larger main spines and smaller by-spines, | ||||||
| 105. Cromyechinus. | ||||||
| Radial spines branched or forked, | ||||||
| 106. Cromyodrymus. | ||||||
| V. Subfamily Caryommida. (Numerous, five to ten or more, concentric latticed shells.) | ||||||
| Two medullary shells and three or more cortical shells (lattice work ordinary), | ||||||
| 107. Caryomma. | ||||||
| No medullary shells (lattice work arachnoidal); three-sided prismatic spines, with verticils of three forked branches. (Arachnosphærida.) | ||||||
| Triangular meshes, simple bars, without diagonal bars, | ||||||
| 108. Arachnopila. | ||||||
| Triangular meshes, simple bars; diagonal bars between the shells, | ||||||
| 109. Arachnopegma. | ||||||
| Polygonal meshes, bars branched, | ||||||
| 110. Arachnosphæra. | ||||||