The genus Spirema begins the series of the Spiremida, or of those Lithelida in which the medullary shell presents a simple latticed sphere or ellipsoid, never composed of a double, trizonal or Larnacilla-shaped shell. In the present state of our knowledge we cannot say whether this simple medullary shell be a primary formation, or effected by secondary means, by reduction of a double Larnacilla-shaped medullary shell, which is constantly found in the Larcospirida. The species of this genus (as of all Lithelida) are difficult to distinguish, are transformistic, and incline very much to variations and abnormalities. The spiral may be simple or double.

Subgenus 1. Spiremarium, Haeckel.

Definition.—Spiral convolutions of the cortical shell simple.

1. Spirema lentellipsis, n. sp.

Cortical shell lentelliptical, with smooth surface; proportion of its three dimensive axes = 4 : 5 : 6. In the median plane are visible four perfect turnings of the simple spiral, the breadth of which gradually increases towards the third convolution, finally decreasing; the broadest (third) turning three times as broad as the simple spherical medullary shell.

Dimensions.—Length of the lentelliptical cortical shell 0.18, breadth 0.15, height 0.12; diameter of the spherical medullary shell 0.02.

Habitat.—North Atlantic, Station 353, surface.

2. Spirema melonia, n. sp. (Pl. 49, fig. 1).

Cortical shell nearly spherical, with smooth surface; proportion of its three dimensive axes = 1.4 : 1.5 : 1.6. In the median plane are visible three perfect turnings of the simple spiral, all of the same breadth as the simple spherical medullary shell; the breadth of each convolution somewhat greater at the poles of the principal than at the poles of the sagittal axis.

Dimensions.—Length of the shell 0.16, breadth 0.15, height 0.14; medullary shell 0.018.

Habitat.—Pacific, central area, Station 271, surface.

3. Spirema flustrella, Haeckel.

Flustrella haliomma, Ehrenberg (1861), Abhandl. d. k. Akad. d. Wiss. Berlin, 1872, p. 293, Taf. ii. fig. 6.

Cortical shell egg-shaped, with thorny surface; proportion of its three axes = 4 : 5 : 6. In the median plane are visible three perfect turnings of the simple spiral, the first and second of about the same breadth as the simple spherical medullary shell, the third suddenly increasing, and finally three to four times as broad. Network of the surface irregular, with roundish pores.

Dimensions.—Length of the shell 0.2, breadth 0.17, height 0.14; medullary shell 0.013.

Habitat.—North Atlantic, Greenland, 1000 fathoms, Ehrenberg; Færöe Channel, John Murray.

Subgenus 2. Spiremidium, Haeckel.

Definition.—Spiral convolutions of the cortical shell double.

4. Spirema diplospira, n. sp.

Cortical shell lentelliptical, with smooth surface; proportion of its three dimensive axes = 6 : 7 : 8. In the median plane are visible three perfect turnings of a double spiral, the breadth of which gradually increases; the broadest (third) convolution three times as broad as the simple lentelliptical medullary shell.

Dimensions.—Length of the shell 0.24, breadth 0.21, height 0.18; medullary shell 0.02.

Habitat.—Tropical Pacific, Philippines, Station 200, depth 250 fathoms.

5. Spirema subglobosum, n. sp.

Cortical shell nearly spherical, with thorny surface; proportion of its three axes = 2 : 2.1 : 2.2. In the median plane are visible two perfect turnings of a double spiral, the breadth of which in the second convolution is four times as great as that of the first convolution and the simple spherical medullary shell.

Dimensions.—Length of the shell 0.22, breadth 0.21, height 0.2; medullary shell 0.02.

Habitat.—Tropical Atlantic, off Sierra Leone, Station 348, depth (2450) fathoms.

Genus 305. Lithelius,[345] Haeckel, 1862, Monogr. d. Radiol., p. 519.

Definition.Lithelida with simple, spherical or subspherical, medullary shell, and lentelliptical or subspherical, spirally constructed cortical shell; surface covered with numerous, simple or branched, radial spines.

The genus Lithelius, founded by me in 1862, and represented by two Mediterranean species, was at that time the only known form of this family, which now contains six genera and twenty-seven species. It differs from the foregoing Spirema in the possession of numerous radial spines on the surface. These may be either simple or branched. The spiral may be simple or double, and according to this latter modification we distinguish two different subgenera.

Subgenus 1. Lithospira, Haeckel.

Definition.—Spiral convolutions of the cortical shell simple.

1. Lithelius spiralis, Haeckel.

Lithelius spiralis, Haeckel, 1862, Monogr. d. Radiol., p. 519, Taf. xxvii. figs. 6, 7.

Cortical shell lentelliptical, one and a third times as long as broad, covered with very numerous (one hundred to one hundred and fifty or more) simple, bristle-shaped radial spines, about as long as the shell. Spiral turnings simple, all nearly of the same breadth and scarcely broader than the simple spherical medullary shell.

Dimensions.—Length of the cortical shell (with six spiral convolutions) 0.15, breadth 0.13; diameter of the medullary shell 0.012.

Habitat.—Mediterranean, Messina, Haeckel, surface; Atlantic, Stations 348 to 353, surface.

2. Lithelius primordialis, R. Hertwig.

Lithelius primordialis, R. Hertwig, 1879, Organism. d. Radiol., p. 54, Taf. vi. figs. 4, 4a.

Cortical shell subspherical, covered with numerous simple, bristle-shaped radial spines, longer than the shell. Spiral turnings simple, with gradually increasing breadth, so that the beginning of the third spiral is twice as broad as the first and as the simple spherical medullary shell.

Dimensions.—Diameter of the cortical shell (with two spiral convolutions) 0.12; diameter of the medullary shell 0.02.

Habitat.—Mediterranean, Messina, R. Hertwig, surface.

3. Lithelius alveolina, Haeckel, 1862.

Lithelius alveolina, Haeckel, 1862, Monogr. d. Radiol., p. 520, Taf. xxvii. figs. 8, 9.

Cortical shell spherical, covered with simple, very numerous (two hundred to three hundred or more), short, bristle-shaped radial spines, scarcely half as long as the radius of the shell. Spiral turnings simple, with gradually increasing breadth, so that the beginning of the third spiral is three times as broad as the first and as the simple spherical medullary shell.

Dimensions.—Diameter of the cortical shell (with four spiral convolutions) 0.2; diameter of the medullary shell 0.01.

Habitat.—Cosmopolitan; Mediterranean, Atlantic, Pacific, on many Stations, surface.

4. Lithelius capreolus, n. sp.

Cortical shell lentelliptical, one and a fourth times as long as broad, covered with numerous (eighty to one hundred and twenty or more) short, branched radial spines, about half as long as the radius of the shell; each spine once or twice dichotomous, with curved divergent branches. Spiral turnings simple, with gradually increasing breadth, so that the beginning of the third spiral is twice as broad as the first, and as the simple spherical medullary shell.

Dimensions.—Length of the cortical shell (with four spiral convolutions) 0.18, breadth 0.14; diameter of the medullary shell 0.015.

Habitat.—Pacific, central area, Station 274, surface.

Subgenus 2. Drymospira, Haeckel, 1881, Prodromus, p. 464.

Definition.—Spiral convolutions of the cortical shell double.

5. Lithelius solaris, n. sp. (Pl. 49, fig. 2).

Cortical shell spherical, covered with simple, very numerous (two hundred to three hundred) bristle-shaped radial spines, longer than the diameter of the shell. Spiral turnings double, both of the same breadth, gradually increasing with the growth of the shell and several times surpassing the diameter of the simple spherical medullary shell. (Pl. 49, fig. 2, exhibits only the first convolutions in the centre of the shell.)

Dimensions.—Diameter of the cortical shell (with four spiral convolutions) 0.18; diameter of the medullary shell 0.02.

Habitat.—Pacific, central area, Stations 266 to 272, surface and in various depths.

6. Lithelius arborescens, n. sp.

Cortical shell lentelliptical, one and a half times as long as broad, covered with numerous (fifty to eighty or more) branched radial spines, about as long as the greatest diameter of the shell; each spine with two to four lateral branches, which are again branched or dichotomous. Spiral turnings double, both of little different breadth, which increases considerably with the growth of the shell, so that the third turn is four times as broad as the simple spherical medullary shell.

Dimensions.—Length of the cortical shell (with three spiral convolutions) 0.18, breadth 0.12; diameter of the medullary shell 0.01.

Habitat.—North Atlantic, Færöe Channel, surface, John Murray.

Subfamily 2. Larcospirida, Haeckel.

Definition.Lithelida with double, trizonal, or Larnacilla-shaped medullary shell.

Genus 306. Larcospira,[346] n. gen.

Definition.Lithelida with double, trizonal, or Larnacilla-shaped medullary shell; cortical shell subspherical or lentelliptical, constructed of a simple or double spiral of the transverse girdle (or primary cortical girdle); the spiral lamella revolving around the principal axis.

The genus Larcospira begins the interesting series of the Larcospirida, or of those Lithelida in which the medullary shell is formed by a trizonal or Larnacilla-shaped lattice-shell, and the cortical shell by spiral turnings of one of the three girdles, which compose the cortical shell of the Pylonida. In Larcospira, as the oldest and most simple form of Larcospirida, the spiral is formed by the transverse girdle, or the first girdle of the Diplozonaria, the only cortical girdle of Amphipyle. If in this genus one of both wings of the transverse girdle grow stronger than the other and overgrow the latter, turning around the principal axis, we reach the characteristic form of Larcospirema, the first subgenus of Larcospira; but if afterwards the second wing follow the example of the first, and overgrow it from the other side, we reach the typical form of the second subgenus, Larcospironium. In this latter subgenus the spiral becomes double, whilst in the former it remains simple.

Subgenus 1. Larcospirema, Haeckel.

Definition.—Spiral convolutions or turns of the cortical shell simple; only one single wing of the transverse girdle turning around the principal axis.

1. Larcospira lentelliptica, n. sp.

Cortical shell with smooth surface, four times as long as the lentelliptical medullary shell. Perimeter of the lateral plane elliptical, one and a half times as long as broad, without constrictions. One lateral wing of the transverse girdle is more strongly developed and turns around the other in two to three simple spiral turns.

Dimensions.—Length of the cortical shell 0.2, breadth 0.16; length of the medullary shell 0.05, breadth 0.04.

Habitat.—Pacific, central area, Station 265, depth 2900 fathoms.

2. Larcospira quadrangula, n. sp. (Pl. 49, fig. 3).

Cortical shell with thorny surface, three times as long as the lentelliptical medullary shell. Perimeter of the lateral plane nearly quadrangular, with four rounded corners (on the poles of two crossed diagonal axes), one and a third times as long as broad, with one sagittal constriction at the poles of the principal axis. One lateral wing of the transverse girdle turns around the other in one and a half to two simple spiral turns.

Dimensions.—Length of the cortical shell 0.18, breadth 0.14; length of the medullary shell 0.06, breadth 0.04.

Habitat.—Pacific, central area, Station 274, depth 2750 fathoms.

3. Larcospira sexangula, n. sp.

Cortical shell with spiny surface, four times as long as the hexagonal medullary shell. Perimeter of the lateral plane nearly hexagonal, one and a half times as long as broad, with six rounded corners (two on the poles of the principal axis, four on the poles of two crossed diagonal axes), with three slight ring-like constrictions. One lateral wing of the transverse girdle turns around the other in two to two and a half simple spiral turns.

Dimensions.—Length of the cortical shell 0.24, breadth 0.16; length of the medullary shell 0.06, breadth 0.04.

Habitat.—Pacific, central area, Station 271, depth 2425 fathoms.

Subgenus 2. Larcospironium, Haeckel.

Definition.—Spiral convolutions or turns of the cortical shell double; both wings of the transverse girdle turning around the principal axis.

4. Larcospira oliva, n. sp.

Cortical shell with smooth surface, six times as long as the lentelliptical medullary shell. Perimeter of the lateral plane elliptical, one and a fourth times as long as broad, without constrictions. Both lateral wings of the transverse girdle turn round one another and form one and a half to two double spiral turns.

Dimensions.—Length of the cortical shell 0.25, breadth 0.2; length of the medullary shell 0.04, breadth 0.035.

Habitat.—Antarctic Ocean, Station 157, depth 1950 fathoms.

Genus 307. Pylospira,[347] n. gen.

Definition.Lithelida with double, trizonal or Larnacilla-shaped medullary shell; cortical shell subspherical or lentelliptical, constructed of a single or double spiral of the lateral girdle (or second cortical girdle); the spiral lamella revolving round the sagittal axis.

The genus Pylospira follows after Larcospira as the second genus of Larcospirida; but in this latter the spiral of the cortical shell is formed by the transverse girdle (or the first lattice-girdle of the Diplozonaria), whilst in Pylospira it is produced by the lateral girdle, or the second lattice-girdle of that group. Therefore Pylospira may be derived phylogenetically from Tetrapyle in the same manner as Larcospira from Amphipyle. Whilst in this latter the first cause of the spiral turning, the unequal growth of both girdle-wings, proceeds from the transverse girdle, in Pylospira it proceeds from the lateral girdle. One of its wings overgrows the other, turning around the sagittal axis. If the second wing do not become developed, the spiral remains simple and represents the subgenus Pylospirema; but if afterwards the second wing follow the example of the first and overgrow it from the other side, we reach the typical form of the second subgenus, Pylospironium, with a double spiral.

Subgenus 1. Pylospirema, Haeckel.

Definition.—Spiral convolutions or turns of the cortical shell simple, only one single wing of the lateral girdle turning around the sagittal axis.

1. Pylospira tetrapyle, n. sp.

Cortical shell with thorny surface, five times as long as the lentelliptical medullary shell. Perimeter of the lateral plane elliptical, one and a half times as long as broad. Four internal gates (as in Tetrapyle, between the complete lateral wings of the transverse girdle) kidney-shaped. One principal wing of the lateral girdle turns around the other in one and a half to two simple spiral turns.

Dimensions.—Length of the cortical shell 0.21, breadth 0.15; length of the medullary shell 0.04, breadth 0.03.

Habitat.—Pacific, central area, Station 268, depth 2900 fathoms.

2. Pylospira octopyle, n. sp. (Pl. 49, fig. 4).

Cortical shell with thorny surface, three times as long as the lentelliptical medullary shell. Perimeter of the lateral plane elliptical or nearly hexagonal, with four rounded corners (on the poles of two crossed diagonal axes), one and a third times as long as broad. Eight internal egg-shaped gates (as in Octopyle), between the complete lateral wings of the transverse girdle, and two axial beams in the principal axis. One single wing of the lateral girdle turns around the other in two to three simple spiral turns.

Dimensions.—Length of the cortical shell 0.18, breadth 0.14; length of the medullary shell 0.06, breadth 0.035.

Habitat.—North Pacific, Station 253, depth 3125 fathoms.

Subgenus 2. Pylospironium, Haeckel.

Definition.—Spiral convolutions or turns of the cortical shell double; both wings of the lateral girdle turning around the sagittal axis.

3. Pylospira cymbium, n. sp.

Cortical shell with smooth surface, seven times as long as the lentelliptical medullary shell. Perimeter of the lateral plane elliptical, one and a third times as long as broad. Four internal kidney-shaped gates between the complete lateral wings of the transverse girdle (as in Tetrapyle). Both principal wings of the lateral girdle turn round one another in two to two and a half double spiral turns.

Dimensions.—Length of the cortical shell 0.22, breadth 0.17; length of the medullary shell 0.03, breadth 0.025.

Habitat.—Indian Ocean, Madagascar, Rabbe, surface.

Genus 308. Tholospira,[348] n. gen.

Definition.Lithelida with double, trizonal or Larnacilla-shaped medullary shell; cortical shell subspherical or lentelliptical constructed of a simple or double spiral of the sagittal girdle (or third cortical girdle); the spiral lamella revolving around the transverse axis.

The genus Tholospira represents the third genus of Larcospirida. Whilst the spiral growth of the cortical shell is produced in Larcospira by the first (transverse) girdle of the Diplozonaria, in Pylospira by the second (lateral) girdle, in Tholospira it is effected by the third or sagittal girdle, which we found complete in Pylonium. If in this genus one wing of the sagittal girdle overgrow the other remaining one and turn around the transverse axis, we get Tholospirema, the first subgenus of our genus, with simple spiral; but if afterwards the second wing follow the example of the first, and overgrow it from the other side, we get Tholospironium, with double spiral.

Subgenus 1. Tholospirema, Haeckel.

Definition.—Spiral convolutions or turns of the cortical shell simple, only one single wing of the sagittal girdle turning around the transverse axis.

1. Tholospira nautiloides, n. sp.

Cortical shell with smooth surface, five times as long as the lentelliptical medullary shell. Perimeter of the lateral plane elliptical, one and a half times as long as broad. Four internal kidney-shaped gates (between the lateral wings of the transverse girdle, as in Tetrapyle). One wing of the sagittal girdle turns around the other in two to two and a half simple spiral turns.

Dimensions.—Length of the cortical shell 0.24, breadth 0.18; length of the medullary shell 0.05, breadth 0.04.

Habitat.—Indian Ocean, between Ceylon and Socotra, Haeckel, surface.

2. Tholospira spinosa, n. sp.

Cortical shell covered with numerous (sixty to eighty or more) simple, bristle-like radial spines, about the length of the shell. Perimeter of the lateral plane nearly quadrangular, with four rounded corners (on the poles of two crossed diagonal axes), one and a half times as long as broad. Eight internal egg-shaped gates (between the lateral wings of the transverse girdle and two axial beams in the principal axes, as in Octopyle). One wing of the sagittal girdle turns around the other in two to three simple spiral turns.

Dimensions.—Length of the cortical shell 0.27, breadth 0.18; length of the medullary shell 0.04, breadth 0.03.

Habitat.—South Pacific, Station 288, surface.

3. Tholospira dendrophora, n. sp. (Pl. 49, fig. 6).

Cortical shell covered with numerous (forty to fifty or more) branched radial spines, about half as long as the shell; each spine with two to six dichotomous branches. Perimeter of the lateral plane elliptical, one and a third times as long as broad. Eight internal egg-shaped gates, as in the foregoing species. One single wing of the sagittal girdle turns around the other in three to four spiral turns.

Dimensions.—Length of the cortical shell 0.22, breadth 0.17; length of the medullary shell 0.05, breadth 0.04.

Habitat.—Pacific, central area, Station 272, depth 2600 fathoms.

Subgenus 2. Tholospironium, Haeckel.

Definition.—Spiral convolutions of the cortical shell double; both wings of the sagittal girdle turning around the transverse axis.

4. Tholospira hystrix, n. sp.

Cortical shell covered with numerous (sixty to eighty or more) simple conical spines, about half as long as the shell. Perimeter of the lateral plane hexagonal, one and a half times as long as broad. Four internal kidney-shaped gates, as in Tetrapyle. Both wings of the sagittal girdle turn round one another in two to two and a half double spiral turns.

Dimensions.—Length of the cortical shell 0.21, breadth 0.14; length of the medullary shell 0.04, breadth 0.03.

Habitat.—North Atlantic, Station 354, surface.

5. Tholospira cervicornis, n. sp. (Pl. 49, fig. 5).

Cortical shell covered with numerous (forty to fifty or more) branched radial spines; each spine antler-shaped, about as long as the medullary Larnacilla-shell, with eight to twelve dichotomous branches (similar to Cromyodrymus abietinus, Pl. 30, fig. 6). Perimeter of the lateral plane elliptical, one and a third times as long as broad. Four internal kidney-shaped gates, as in Tetrapyle. Both wings of the sagittal girdle turn round one another in one and a half to two double spiral turns.

Dimensions.—Length of the cortical shell 0.2, breadth 0.15; length of the medullary shell 0.03, breadth 0.025.

Habitat.—Pacific, central area, Station 271, depth 2425 fathoms.

Genus 309. Spironium,[349] n. gen.

Definition.Lithelida with double, trizonal or Larnacilla-shaped medullary shell; cortical shell subspherical or lentelliptical, constructed of two crossed spirals, which arise from both lateral wings of one girdle (commonly the lateral girdle) and revolve in an opposite diagonal direction around the principal axis.

The genus Spironium differs in a very remarkable manner from all foregoing Lithelida, and is distinguished by a quite peculiar mode of growth. It is most nearly allied to Larcopyle, and may, like this, be derived from Amphipyle (or rather from Larnacilla, beginning to transform into Amphipyle). But whilst in Larcopyle one of the two wings of the lateral girdle overgrows the other in the direction of the transverse axis (turning around the principal axis), here in Spironium both lateral wings begin at the same time to grow out from the lateral sides of the Larnacilla-shaped medullary shell; the most remarkable thing is, however, that the direction of growth in the wings is diverse from the beginning: the left wing grows downwards and turns around the lower (aboral) pole of the principal axis, the right wing grows upwards and turns around the upper (oral) pole of the same axis. Thus both wings of the lateral girdle are crossed in diagonal axes, and with increasing growth one overgrows the other in the direction of these diagonals, so as to resemble the figure 8 in shape. The open gates remaining between the turnings of the girdle become afterwards closed on the surface by irregular lattice-work, and so the whole cortical shell assumes finally a spherical, ellipsoidal, or lentelliptical form. Its surface sometimes becomes covered with simple or branched radial spines. In the interior the eight characteristic egg-shaped gates of Octopyle are commonly (or constantly?) visible, two strong radial beams in the principal axis arising from the poles of the lentelliptical medullary shell.

Subgenus 1. Spironetta, Haeckel.

Definition.—Surface of the shell smooth or rough, but not with radial spines.

1. Spironium octonium, n. sp. (Pl. 49, fig. 7).

Cortical shell lentelliptical, its breadth (or transverse axis) surpassing considerably the length (or the principal axis). Surface thorny and somewhat hump-backed. The spiral wings of the transverse girdle have about the same breadth as the eight internal gates between them and the axial beams.

Dimensions.—Length of the cortical shell (or principal axis) 0.15, breadth (or transverse axis) 0.2; length of the medullary shell 0.05, breadth 0.04.

Habitat.—South Atlantic, west of Tristan da Cunha, Station 332, depth 2200 fathoms.

2. Spironium diagonale, n. sp.

Cortical shell nearly spherical, four times as great as the subspherical medullary shell. Surface rough. The spiral wings of the transverse girdle about half as broad as the eight internal gates between them and the axial beams.

Dimensions.—Diameter of the cortical shell 0.16, of the medullary shell 0.04.

Habitat.—Indian Ocean, Zanzibar, Pullen, depth 2200 fathoms.

Subgenus 2. Spironilla, Haeckel.

Definition.—Surface of the shell covered with simple or branched radial spines.

3. Spironium spinosum, n. sp.

Cortical shell subspherical, five times as great as the subspherical medullary shell. Surface covered with numerous (sixty to eighty or more) simple, bristle-like radial spines, longer than the shell. The spiral wings of the transverse girdle of about the same breadth as the eight internal gates between them and the axial beams.

Dimensions.—Diameter of the cortical shell 0.2, of the medullary shell 0.04.

Habitat.—Pacific, central area, Station 274, surface.

4. Spironium arbustum, n. sp.

Cortical shell lentelliptical, its breadth surpassing its length considerably. Surface covered with numerous (forty to sixty or more) thin radial spines, about as long as the greatest diameter of the shell; each spine with two to six lateral branches, which are either simple or again branched (similar to Cromyodrymus abietinus, Pl. 30, fig. 6). The spiral wings of the transverse girdle only half as broad as the eight internal gates between them and the strong beams of the principal axis.

Dimensions.—Length of the cortical shell 0.12, breadth 0.15; length of the hexagonal medullary shell 0.05, breadth 0.04.

Habitat.—Pacific, central area, Station 271, depth 2425 fathoms.

Family XXX. Streblonida, n. fam. (Pl. 49, figs. 8, 9).

Definition.Larcoidea with asymmetrical, spiral, polythalamous shell, composed of a variable number of roundish chambers, which form together an ascending spiral; both halves of the shell unequal. Primordial chamber either simple or Larnacilla-shaped.

The family Streblonida comprises those Larcoidea in which a number of chambers is arranged in an ascending spiral, round a simple or trizonal primordial chamber, like winding stairs. They show the same spiral structure as in the foregoing family, but whilst in the Lithelida the spiral line lies in one plane (as in Nautilus), in the Streblonida it ascends like a screw (as in Helix). Therefore the former have the same relation, regarding the spiral structure, to the nautiloid Polythalamia as the latter to the turbinoid Foraminifera. Indeed the single forms of Streblemida repeat in their special structure the characteristic genera of Turbinoida, such as Globigerina, Rosalina, Pulvinulina, Hastigerina, &c. As in these calcareous turbinoid Foraminifera, so also in the analogous siliceous Streblonida the distinction of species is very difficult and open to many objections.

The number of species in this family is very small; all are rare and for the most part very opaque and difficult to understand, so that the following distinction of a dozen species can have only a provisional value. To get a complete idea of their peculiar structure, the shell must be turned and observed from different sides, and thus their full study requires yet much time and work. There are to be found evident transitional forms between them and the Lithelida on the one hand and the Soreumida on the other. Besides this, most species of Streblonida seem to have more inclination to individual varieties and abnormalities than the majority of the other Radiolaria.

The general form of the whole shell is in the Streblonida sometimes more egg-shaped or even subspherical, at other times more top-like or conical, sometimes nearly discoidal. The height of the shell (or the vertical axis of the ascending spiral) is occasionally larger, at other times smaller than the breadth (or the greatest horizontal diameter, perpendicular to the height). Some very flat forms seem to approach the Lithelida. With regard to the internal screw-formation, the shell of all Streblonida is asymmetrical.

The number of the aggregated incomplete chambers is commonly between ten and twenty, but ascends sometimes to thirty, forty, or more. Sometimes the size of all the chambers is nearly the same, sometimes they increase gradually, occasionally also very rapidly. The primordial chamber (or the first and oldest) seems to be commonly the smallest, and inversely, the last and youngest chamber, the largest. But sometimes (in Streblopyle) also the contrary may be the case. The form of the single chambers is very variable, from the spherical or hemispherical through all transitions leading to irregular roundish or longish forms. The network is commonly irregular, with small roundish pores of different sizes, but sometimes also regular, circular. The surface of the shell is commonly smooth or rough, rarely covered with radial spines. In most species the reticulation and particularly the separation of the chambers is more or less incomplete.

As in the Lithelida, so also in the Streblonida we can distinguish two subfamilies. In the Streblacanthida (Streblonia, Streblacantha) the primordial chamber is a simple, spherical, subspherical, or lentelliptical latticed shell. In the Streblopylida (Streblopyle) the primordial chamber is trizonal or Larnacilla-shaped, as in the greater number of Larcoidea, composed of three elliptical latticed girdles of unequal size, perpendicular one to another, and surrounding a simple central chamber. As in the Lithelida, so also here we cannot certainly say whether the former have originated from the latter by reduction of the Larnacilla-shell, or whether both groups be of different origin. The latter is perhaps more probable. This family as well as the foregoing requires a much more careful study than I could give to it.

Synopsis of the Genera of Streblonida.

I. Subfamily Streblacanthida.

Primordial chamber simple, spherical or lentelliptical.

brace Shell without radial spines, 310. Streblonia.
Shell with radial spines, 311. Streblacantha.

II. Subfamily Streblopylida.

Primordial chamber trizonal or Larnacilla-shaped

Shell without radial spines, 312. Streblopyle.
I. Subfamily Streblacanthida. Primordial chamber simple, spherical or lentelliptical.
Shell without radial spines,
310. Streblonia.
Shell with radial spines,
311. Streblacantha.
II. Subfamily Streblopylida. Primordial chamber trizonal or Larnacilla-shaped
Shell without radial spines,
312. Streblopyle.
Genus 310. Streblonia,[350] n. gen.

Definition.Streblonida with simple, spherical, subspherical, or lentelliptical primordial chamber, beginning the screw-like series of spirally ascending chambers. Surface smooth or thorny, without radial spines.

The genus Streblonia contains those Streblonida in which a variable number of roundish, subspherical, or longish chambers form a screw-like aggregate, beginning with a quite simple primordial chamber. The special order of the complex spiral offers interesting resemblances to different genera of the calcareous Foraminifera, from which I have taken the corresponding name of the species. The whole form of the shell is sometimes more egg-shaped or subspherical, at other times more top-like or conical, occasionally very flat. Its surface is smooth or rough, but not covered with radial spines.

1. Streblonia globigerina, n. sp.

Shell subspherical, thick walled, clustered, with eight to ten nearly spherical chambers, of rapidly increasing size, the tenth chamber about twelve times as broad as the first. Breadth of the shell nearly equal to the height. Pores subregular, circular, hexagonally formed, of about the same breadth as the bars; about sixteen on the breadth of the tenth chamber. (Resembles very much the common Globigerina.)

Dimensions.—Breadth of the shell 0.18, height 0.16.

Habitat.—North Pacific, Station 253, depth 3125 fathoms.

2. Streblonia uvigerina, n. sp.

Shell nearly egg-shaped, clustered, with eight to eleven subspherical chambers of gradually increasing size, the tenth chamber about six times as broad as the first. Breadth of the shell about half its height. Pores of the shell subregular, circular, about twice as broad as the bars; about twenty on the breadth of the tenth chamber. (Resembles some species of Uvigerina.)

Dimensions.—Breadth of the shell 0.11, height 0.23.

Habitat.—South Atlantic, Station 332, depth 2200 fathoms.

3. Streblonia polymorphina, n. sp.

Shell egg-shaped, thin walled, with twelve to fourteen roundish chambers of rapidly increasing size, the tenth chamber about seven times as broad as the first. Breadth of the shell about two-thirds of the height. Pores irregular roundish, twice as broad as the bars; about twenty on the tenth chamber. (Resembles certain forms of Polymorphina.)

Dimensions.—Breadth of shell 0.22, height 0.14.

Habitat.—Pacific, central area, Station 266, depth 2750 fathoms.

4. Streblonia bulimina, n. sp.

Shell nearly egg-shaped, thick walled, clustered, with fourteen to eighteen egg-shaped chambers of rapidly increasing size, the tenth chamber about eight times as broad as the first. Breadth of the shell about two-thirds of the height. Pores irregular, roundish, half as broad as the bars; about twelve on the breadth of the tenth chamber. (Resembles closely Bulimina.)

Dimensions.—Breadth of the shell 0.17, height 0.24.

Habitat.—Pacific, central area, Station 268, depth 2900 fathoms.

5. Streblonia rosalina, n. sp.

Shell top-shaped or flatly conical, with twelve to sixteen chambers of gradually increasing size, the tenth chamber about four times as broad as the first. Breadth of the shell twice as large as the height. Pores subregular, circular, very small, of the same breadth as the bars; about twelve on the breadth of the tenth chamber. (Resembles some forms of Rosalina.)

Dimensions.—Breadth of the shell 0.27, height 0.13.

Habitat.—Pacific, central area, Station 271, depth 2425 fathoms.

6. Streblonia planorbulina, n. sp.

Shell very flatly conical, nearly lenticular or discoidal, with twenty to twenty-five chambers of nearly equal size, the tenth chamber a little broader than the first. Breadth of the shell exceeds five to six times the height. Pores subregular, circular, very small, half as broad as the bars; about eight on the breadth of the tenth chamber. (Resembles closely Planorbulina.)

Dimensions.—Breadth of the shell 0.3 to 0.4, height 0.06 to 0.07.

Habitat.—Indian Ocean, Zanzibar, Pullen, depth 2200 fathoms.