Fig. 71.—The Rye District.
"The grains extending farthest north in Europe are barley and oats. Rye is the next, and is the prevailing grain in Sweden and Norway, and all the lands bordering on the Baltic, the North of Germany, and part of Siberia. Wheat follows rye; it is cultivated in the middle and South of France, England, part of Scotland, part of Germany, Hungary, Crimea, and the Caucasus. We next come to a district where wheat still abounds, but no longer exclusively furnishes bread,—rice and maize becoming frequent. To this zone belong Portugal, Spain, part of France, Italy, and Greece, Persia, Northern India, Arabia, Egypt, the Canary Islands, &c. Wheat can be reared wherever the mean temperature of the whole year is not under 37° or 39° F., and the mean summer heat, for a period of at least three or four months, is above 55°. It succeeds best on the limits of the sub-tropical region. In the Scandinavian peninsula, the cultivation of barley extends to 70° N. latitude, rye to 67°, and oats to 65°. The cultivation of rice prevails in Eastern and Southern Asia, and it is a common article of subsistence in various countries bordering on the Mediterranean. Maize succeeds best in the hottest and dampest parts of tropical climates. It may be reared as far as 40° N. and S. latitude on the American continent, on the western side; while in Europe it can grow even to 50° or 52° of latitude. It is now cultivated in all regions in the tropical and temperate zones which are colonised by Europeans. Millet of different kinds is met with in the hottest parts of Africa, in the South of Europe, in Asia Minor, and in the East Indies."[80]
Professor Houston furnishes the following table in illustration of the distribution of wheat and barley. It also shows the mean temperature which they require:—
BARLEY.
| Winter | Summer | Annual | ||
| Mean. | Mean. | Mean. | ||
| N. Lat. 62½° | Faroe, | 39° | 51° | 45° |
| 70 | Lapland, | 22 | 46 | 33 |
| 67·30' | Russia, | 9 | 46 | 32 |
| 57·30' | Siberia, | 0 | 60 | 32 |
WHEAT.
| Winter | Summer | Annual | ||
| Mean. | Mean. | Mean. | ||
| Lat. 58° | Scotland, | 36° | 57° | 64° |
| Norway, | 23 | 59 | 39 | |
| Sweden, | 23 | 59 | 39 | |
| Russia, | 15 | 60 | 37 | |
| 30° | Cairo, | 57 | 88 | 72 |
| Macao, | 64 | 82 | 73 | |
| Rio Janeiro, | 68 | 78 | 74 | |
| Havannah, | 71 | 82 | 77 | |
| Bourbon, | 71 | 80 | 77 |
"Winds, water, and animals, are also instrumental in disseminating plants. Many seeds, with winged and feathery appendages, are easily wafted about; others are carried by rivers and streams, and some can be transported by the ocean currents to a great distance, with their generating powers unimpaired."
Mushrooms or Agarics (Order Fungi).
Every year—principally in autumn—we are startled by hearing or reading of cases of poisoning by mushrooms. Erudite connoisseurs, however, who have profited by Dr Badham's book on "Esculent Fungi," do not suffer themselves to be intimidated by these sad narratives, though, unfortunately, they are frequently too well founded; because they know how to distinguish the good from the ill, the true from the false, the edible from the poisonous mushroom. But this security ought not to embolden the inexperienced amateur in risking his life for the sake of a delicacy. It is true, nevertheless, that we frequently see men's lives exposed for something less.
"Look, what a splendid mushroom I have discovered!" a lady said to me, the other day; a lady who knew something of flowers, but nothing of cryptogams. "Take care!" I replied, taking from her hand the supposed prize; "this is the false mushroom; and would suffice, if served up at your table, to poison yourself and all your guests."
I ought here to observe that my friend had narrowly escaped death two years before, through regaling herself with a dish of mushrooms of very dubious character. Among the symptoms which she experienced, and which she described with medical exactness, she particularly dwelt upon the cold sweats, accompanied by a sentiment of undefinable terror, which is nothing else than the dread of death: it was the special symptom of poisoning with an unwholesome fungus.
Let us endeavour to make ourselves better acquainted with this formidable enemy of epicures. You will have no difficulty in finding it in any warm, close season, but especially in spring and autumn. The toadstool thrives indifferently in the shade of all the forest trees, but seems to prefer the oak and birch to the pine and fir. A patch of soft greensward, at the foot of an old oak, and in the neighbourhood of a "brawling stream" or "tranquil pool," will generally be covered with fungi of this description. The numerous synonyms attaching to it show how greatly it has exercised the classifying spirit of our naturalists. Some call it Agaricus muscarius, as if we should say "kill-fly mushroom;" others, changing only the specific designation, designate it Agaricus pseudo-aurantiacus,—which signifies, literally, "false orange," in allusion to the beautiful yellow colour of the true aurantiacus. What is certain is, that our mushroom, which can kill men as well as flies, belongs to the genus Agaric, so numerous in species that it can be formed into a family, that of the Agariceæ. The Agarics, of which the esculent mushroom (Agaricus edulis) represents the type, are easily recognised by their more or less fleshy pileus, or cap, garnished underneath with lamellæ, or gills, which radiate from the centre to the circumference, when the pedicel is in the centre.
Fig. 72.—"At the foot of an old oak."
But some cryptogamists are unwilling to recognise the orange mushrooms (les oronges), whether true or false, as Agarics. They place them in a separate genus, the genus Amanita, though without informing us where they found the name. Meanwhile, they justify the formation of the new genus by the presence of the white swelling, the volva, or wrapper, of the mycelium, or spawn, which entirely covers both the true and the false mushroom on its emergence from the earth. Each, then, is an Amanita. But now remark their specific difference. The true mushroom, as it develops, ruptures its ovoid wrapper, or volva, leaving the remains entirely at the base of the pedicel; while, in the false mushroom, the débris of the volva are formed, not only at the base of the pedicel, as in the real Agaric, but even upon the red surface of the pileus itself: these are the white irregular warts characteristic of the Amanita, but wholly wanting in the Agaricus. Thus, there are two Amanitas: the Amanita muscaria, or "fly agaric," and the Amanita aurantiaca, or, as the English botanists call it, Agaricus Cæsareus, the imperial mushroom.
This fanciful "study" in nomenclature has the advantage of initiating us into the most essential distinctive characters of the two species in question. However, a few additional details are necessary to complete our history.
The Fly Agaric, or Amanita muscaria.
The species seems to have been expressly created to teach our gourmands the necessity of vigilance; that before enjoying a dainty they must first learn to distinguish, under penalty of death, the poisonous fungus from that which safely and pleasantly tickles the palate. The warning is useful, moreover, as showing that even sensualists are not wholly exempt from the law of work.
Fig. 73.—The Amanita Muscaria.
The Amanita, like most falsehoods, is pleasant to the sight. Its round pileus, of a beautiful orange-red colour, spotted with white warts, and lined with white gills, seems to invite your attention. (Fig. 73.) Strike it down with your stick; the lamellæ or gills underneath resemble the white leaves of a book. Its graceful stalk, ornamented on the upper part by a well-designed necklet, is bulbous in its lower portion; its flesh is dazzlingly white; in short, its entire appearance is attractive. Yet it is a traitor! You little know the depth of its wickedness. Mix a few shreds of its white flesh with a little milk; every fly which drinks of the mixture will, in a few seconds, fall dead, their swollen abdomen bearing testimony to the effect of the poison. And in many districts of Germany, particularly in Thuringia, the peasants make use of it to rid themselves of the swarms of flies which, towards the close of summer, infest their habitations. It is thus that man may frequently turn to his advantage those objects of nature which, at the first glance, appear injurious rather than useful.
Various experiments have been essayed to test the intoxicating influence of our Amanita. Bulliard, author of the "Histoire des Champignons de France," made two cats partake of it. Six hours afterwards, these animals, who are so tenacious of life, were dead.
Haller, in his "Histoire des Plantes Vénéneuses de la Suisse," says that "the Agaricus muscarius (Amanita muscaria) cannot be eaten with impunity, six Lithuanians having died of it; and in Kamtschatka it has been known to excite deadly attacks of delirium, accompanied by so deep a despondency, that those who have eaten of it would fain fling themselves into the fire, or fall upon their knives or daggers." This, however, we take to be an exaggeration. The truth is, that in Kamtschatka it is used to produce intoxication; and such is its strength, it imparts an intoxicating property to the urine of those who swallow it. When the fungus itself is not at hand, the would-be drunkard frequently resorts to this nauseous potion.
The flesh of the Amanita is not yellow. Yet Vicat speaks of poisonings produced by the Yellow Amanita. "I had much difficulty," says this physician, "in saving two families of Lausanne, poisoned through eating a very small quantity of mushrooms, which the father in the one case, and the mother in the other, had mistaken for Agarici Cæsarei, though they were both esteemed great connoisseurs, and especially in this species; nor had they been once deceived for upwards of thirty years, until they indulged themselves in this delicious but deceitful dish."
These poisonings could have been occasioned only by the Amanita. Some varieties exist, in which the under surface of the pileus is yellow, but the flesh is white. To one of these varieties Vicat's anecdote probably refers.
Thus, the Amanita formosa of Persoon has pedicel, pileus, and warts of the pileus, of a citron yellow. It is but a variety of the Amanita muscaria.
The Amanita umbrina of the same botanist (the Agaricus pantherinus of De Candolle) has an olive-coloured pileus; its surface, like that of Amanita muscaria, is covered with white scales.
The Amanita solitarius is distinguished by the size of its umbilical cap,—sometimes depressed in the centre,—which is furnished with a great number of white or pale brown scales; when fully developed, it measures from thirty to forty centimetres in diameter. The pedicel is bulbous, with a membranous ring, white as snow, clasped around it; at the base it is clothed in pellicles, the remains of its scaly volva. The flesh is firm, thick, and white. You rarely meet with more than two or three individuals in the same locality; hence its name of solitarius. Bulliard speaks of the flesh as good to eat, when cooked on a gridiron, and seasoned with fresh butter, salt, and pepper. It is possible. But as it is so easily confused with the poisonous species, the author of the "Histoire des Champignons" would have done better to prohibit its consumption, whether eatable or not.
We may now turn to the method adopted by Dr Vicat to save the lives of the two families at Lausanne, who, as we have seen, were poisoned by partaking of the Amanita:—
I dissolved, he says, six grains of tartar emetic in a litre of water, and from time to time administered a spoonful to my patients; moreover, I made them swallow floods of warm water, sweetened with a little honey,—that is, a large tea-spoonful of honey to a cupful of water.
I had much difficulty to get one of the sufferers, who was sixty years old, to swallow the first few spoonfuls. He was plunged into a lethargic insensibility differing in no respect from complete apoplexy; his teeth were closely set. Those whom I had ordered to administer the mixture had given it up, after several useless attempts; and in all probability the old man would have sunk, had I not had the patience to hold, for some hours, against his teeth the back of the blade of a small silver knife, so as to profit by the few moments when the teeth were a little less firmly clenched. I used some force to make the blade act as a wedge, and after a while opened up a passage to the handle, which, serving as a lever, forced the jaws sufficiently apart to admit the introduction of a spoonful of the emetic. It was not, however, until fully two hours had passed that the patient, having undoubtedly swallowed the necessary dose, began to vomit, with strenuous efforts and frightful cries. This was at midnight. Four in the morning arrived before, after numerous alternations of vomiting and profound lethargy, he began to speak, and then like a man in delirium. After the first vomit, which was inconsiderable, the convulsions of his whole body were so very violent as to require four men to hold him, while I continued to make use of my knife as at first. Nor did I desist until I was satisfied that his stomach had been sufficiently cleansed. After this, I applied two strong blisters to the back of his legs. As these acted, the purging subsided, and at the end of twenty-four hours it had passed away entirely, the invalid finding himself as well as could be expected after sustaining so severe a shock. The other patients, who were not in so much danger, experienced twitchings and tremblings in the face, which quite disfigured them; the brain seemed a blank; though awake, they felt as in a dream, and their visions were most frightful.
It is evident, from these particulars, that mushroom-poisoning specially affects the encephalic nervous system, and that the best remedies are emetics and antispasmodics. In our present ignorance of what are the poisonous principles in the Amanita, we can adopt no other method than a chemical neutralisation.
Agaricus Cæsareus, or Imperial Mushroom.
Fig. 74.—The Imperial Mushroom.
In this mushroom, for which, as we have seen, the Amanita is too frequently mistaken, the inside as well as the outside is yellow; the upper surface of the pileus, which is equally free from scales and warts, is, however, of a reddish yellow, like that of an orange (whence the popular French name, la vrai oronge); all the other parts are of a beautiful citron hue. This agaric exhales an agreeable odour, combined apparently of the scent of the vanilla and the truffle. It decomposes rapidly, and when in a state of advanced putridity, the fragrance I speak of is succeeded by—well, by a fearful stench! When young, and still completely covered with its wrapper or volva (this, in the Amanita, is imperfect), it is very like a hen's egg which has been partly buried in the ground so as to expose only the larger end. It seems partial to solitude; more than four or five are seldom found in the same locality. Moreover, in autumn it affects the same habitats as the Amanita,—which is unfortunate.
It would seem that our imperial mushroom was specially appreciated by the ancients, and it is said that Nero pronounced it a dish fit for the gods. In this circumstance originated the scientific name which has now become popular, and which was first applied to it by the cryptogamist Fries, Agaricus Cæsareus.
The Boletus of Pliny appears to have been our Agaricus, and not one of our Boleti, which are easily recognised by the numerous tubercular projections covering the under part of the pileus. In proof of this I would point out that the Roman naturalist, after speaking of the Boleti as genuine delicacies, immediately inveighs against them as dangerously poisonous. He relates that it was with one of these, or rather with one of the false mushrooms so easily mistaken for the true, that Agrippina poisoned the Emperor Claudius, to secure the imperial crown for her son Nero.[81]
The virtues of the mushrooms have been sung by Juvenal and Martial. The latter accurately distinguishes the true from the false, when reproaching Cæcilianus with his gluttony. "Ah, you are used to devour your Boleti alone, in the face of your invited guests; eat then, the Boletus which Claudius ate!"
The best known and most valuable species of Agarici may be briefly enumerated:—
Agaricus campestris, or Common Mushroom, found in nearly all temperate regions: pileus convex, and white, with a tinge of brown; thick set on the under side with dark brown gills; stem firm and fleshy, and surrounded by a white membranous ring.
Agaricus Cæsareus, or Imperial Mushroom, the Kaiserling of the Germans, already described.
Agaricus deliciosus, or Orange-milked Agaric, found in coverts of fir and juniper; pileus viscid, orange, and upwards of four inches broad; gills and juice of a fine orange colour.
Agaricus procerus, or Parasol Mushroom, found in the shade of trees, on meadows with a sandy soil; stem from eight to twelve inches high, with a thick spongy ring; pileus bell-shaped, and covered with brown scales.
Agaricus Virgineus, or White Field Agaric, found in rich moist
pastures; pileus whitish, and convex; gills of a light chocolate shade; stem nearly two inches broad.
Agaricus eburneus, or Ivory Mushroom, found in beech woods; grayish-yellow pileus; broad gills; stem long and scaly.
Agaricus Georgii, St George's Agaric, or Whitecaps, found in moist pastures, and in the shelter of old barns, farmhouses, and churches; flesh yellow; gills yellowish-white; pileus twelve to eighteen inches broad; the least valuable of British species of agaric, but useful in ketchup-making.
Agaricus oreades, Fairy-ring Mushroom, or Scotch bonnets, found in meadows, where it grows in circles known as "fairy rings;" pileus seldom exceeds an inch in diameter; stem solid, tough, and fibrous, with a boss or umbo in the centre, of a light brown colour; the flesh white, and of a pleasant odour.
Agaricus odorus, Anise Mushroom, or Sweet-scented Agaric; pileus slightly convex, about three inches broad, and with pale gills; the scent like that of anise; stem strong, fleshy, but not very tall.
Agaricus formosus, or Smoky Mushroom; so called from the colour of the upper surface of the pileus; the stalk and gills of a pale yellow; grows in fir woods.
Agaricus primulus, or Mousseron, grows in woods and pastures, where the soil is sandy; pileus convex, yellow, about three or four inches in diameter; gills change from white to flesh colour.
How many Vegetable Species exist over the whole surface of the Globe?
I will not do my readers the injustice to suppose that they are unacquainted with the writings of our greatest English poetess, Elizabeth Barrett Browning. They will not fail to have been attracted by the prodigal genius, the superabundant power, the exquisite imagery, the profound spirit of tenderness, the high, pure thoughts, which render almost every page such delightful reading. Successful as she was, however, in giving expression to the most subtle emotions and the intensest feeling, I think she was even happier in her descriptions of scenery. These are invariably aglow with life and colour, and have all the fidelity of Creswick with the imaginative insight of Turner. Turning over her "Aurora Leigh," the other day, I lighted on the following beautiful picture:—
Fig. 75.—"And cattle grazing in the watered vales."
As I read this fine passage, the thought occurred to me, how many thousands there are who, in such a scene as it so vividly depicts, would see no beauty whatever, whose heart would not respond to it, whose sympathies would not be aroused by all its variety of outline and all its rich magnificence of colour! Yet not in so wide a landscape alone, but in the smallest nook,—in the little clump of elms by the side of the stream, in yonder grassy knoll rising straight up from the old churchyard, in the quiet angle of the green pasture-meadows,—there is a whole world of wonder and beauty for him who has eyes to see and a heart to feel! Look at the flowery bank which runs along the side of an English lane. Is it not crowded with objects of the rarest and purest interest? Count the many varieties of grasses which clothe it so abundantly, count the many species of flowers and herbs which adorn it with a grace beyond all human skill, and acknowledge that in itself it might supply the inquirer with matter for years of study and meditation.
Pursuing this train of thought, I was led to think of the number of genera and species into which the plant world is divided,—a remarkable proof, not only of the power and wisdom, but of the goodness of the Creator, of His desire to furnish man with inexhaustible sources of pleasure and entertainment; and finally, to put to myself the question, How many vegetable species exist over the whole surface of the globe? If this corner of a leafy English lane is so rich in variety, what must be the case with "the wide, wide world?"
I was now brought to see that a question so difficult could, like so many others, be usefully approached only by its inferior limit; in other words, that in the actual condition of botanical science, we can but affirm the number which certainly exceeds the sum of the vegetable species scattered over the surface of our earth. To determine this total with mathematical accuracy, we should need to have explored the terrestrial crust, liquid and solid, land and water, from the bed of ocean to the line of perpetual snow, and from the equator to the poles. And as yet we are very far from having obtained so complete a possession of the planet which has been assigned as a dwelling-place to our poor humanity,—alas, more presumptuous than powerful!
The number of plants mentioned by Theophrastus, Dioscorides, and Pliny, whom we take to be the representatives of ancient botany, does not exceed five hundred species. How very few, compared with the presumable total! The Middle Ages added scarcely anything to the botanical researches of antiquity. It is only since the discovery of America that we have seen the domain of Flora extending itself in unexpected proportions. But we must come down to the epoch of Linnæus (the middle of the eighteenth century) before we can obtain an accurate list of species, scientifically classified. Murray's edition of the "Specilegium" of Linnæus contains two thousand and forty-two species, including the Cryptogams. Wildmore, in another edition of the same great work, raised the total to twenty thousand. And this was the point at which our botanists had arrived when the nineteenth century opened.
But it was not long before they perceived that all these estimates, large as they seemed, fell immeasurably short of the reality. In attempting to distribute the different species among the then known regions of the globe, Alexander von Humboldt arrived at a total of forty-four thousand species, Phanerogams and Cryptogams included. De Candolle extended the estimate to upwards of fifty-six thousand.
Let us divide, in fancy, the earth into two parts,—one which has been visited by travellers, and one which still remains to be explored. Can you determine which would present the larger area? The latter.
Thus we possess but a very imperfect knowledge of the luxuriant, the glowing vegetation of the tropical and sub-tropical regions of the New World, in spite of the labours of Bates, Agassiz, Wallace, and others. To the north of the equator, we know very little of the flora of Yucatan, Guatemala, Nicaragua, the isthmus of Panama, the Chaco of Antioquius, the province of Los Pastos. We are not much better acquainted with the vegetation of the countries south of the equator. What do we know of the manifold species flourishing in Paraguay, in the province of the Missions, in the immense wooded region between the Ucayali, the Rio de la Madeira, and the Tocantin, three affluents of the mighty river Amazon? We know scarcely anything.
Our ignorance increases if from America we pass to Africa. Nearly the whole interior of this continent, from 15° N. latitude to 20° S. latitude, is, botanically speaking, a blank to us. The same is the case with the greater portion of Central Asia. The floras of the south and south-east of Arabia are still sealed letters,—treasuries to which we have not found the key. As much may be said of the floras of the countries situated between the Thian-Schan, the Kuenlung, and the Himalaya, as well as of the floras of western China, and most of the trans-Gangetic countries. We know still less of the vegetation of the interior of Madagascar, Borneo, New Guinea, and the greater part of Australia. To conclude: we are probably not acquainted with more than one-fifth of the vegetable species which cover the surface of our globe.
There are regions, moreover, which we imagine will always lie outside of our sphere of investigation; such, for instance, are the Polar regions, properly so called. Undoubtedly, it is open to us to conjecture that the Poles—those two extremities of our axis of planetary rotation—are not the home of any form of life. But this is only a conjecture; we are even without an analogy for it; since we have found, as shown in an earlier chapter of the present volume, living beings, plants, and animals, among the snows of our loftiest mountains. Moreover, might not the auroras, whose maximum of intensity occurs exactly at the poles, render life possible in regions where we at present suppose it to be impossible? Conjecture for conjecture,—acknowledge that we here touch in both cases upon an element completely beyond our human power.
These, then, are the reasons why, at present, we can only venture upon defining the lower limit, the restricted number, above which we are unable to fix the total of vegetable species living on the surface of our planet.
The method to be adopted has been indicated by Alexander von Humboldt in his "Ansichten der Natur" ("Pictures of Nature"). His method consists in the comparison of the vegetable families whose numerical relations are known, with the number of species contained in our herbariums, or cultivated in our botanical gardens.
But here, at the outset, a difficulty confronts us. Does any relation exist between the classification of plants by natural families and by their geographical distribution?
To group plants according to their analogies of structure, we study them from an abstract point of view, and without any regard to the medium in which they flourish. The question grows complicated if we also take into consideration their characteristic conditions and their distribution over the terrestrial surface. Families are then split asunder, and the importance of our scientific classifications disappears.
The gathering together of a small number of species, represented by innumerable individuals, confined within the same area, may suffice to communicate to a landscape its characteristic physiognomy: as, for example, is the case with the Asiatic steppes, the landes of Brittany, the moors of Scotland, the palm-groves and the clumps of Cactaceæ of tropical America. By the side of species which impress us by their mass—that is, by the frequent reproduction of the same individuals at an infinitesimal distance from one another—are placed those much more numerous species which are everywhere very thinly sown.
But do the plants themselves follow, from the equator to the poles, the same law of decrease as obtains from the base to the summit of the loftiest equatorial mountains?
Under identical isothermal lines, is the ratio of families known and identified to the probable aggregate of Phanerogams the same, in the temperate zone, on either side of the equator?
What are the vegetable families which preponderate at the two extremes, represented by the torrid and the frigid zones?
Under the same geographical latitude, or between the same isothermal lines, are the Synantheræ, the Gramineæ, the Leguminosæ, the Labiatæ, the Cruciferæ, the Umbelliferæ, more numerous in the Old than in the New World?
What families, either through their mass of individuals or their number of species, take precedence of the other Phanerogams?
How many species of one and the same family belong to any particular country?
What groups or families are characteristic of each zone?
Is the present classification of genera and species in all respects what could be desired?
These are questions that require to be considered, and to some of them we shall presently attempt replies.
Herbariums, though their classification is too frequently imperfect, may furnish us with data of great utility. The great herbarium of Benjamin Delessert was estimated, after his death, to contain 86,000 species,—a total not widely differing from that which Lindley, in 1835, estimated as the probable aggregate of the vegetable species of the world.
Great in importance are botanic gardens. Loudon, in his Hortus Britannicus (ed. 1832), places at 22,660 the number of Phanerogams cultivated in the gardens of the Bristol amateur botanists. With this number we must not confound the living species exhibited, in other counties, in gardens designed for the instruction of students, nor the grand total reared for a similar purpose at Kew. Kunth's enumeration, in 1861, of the plants at the Botanic Gardens at Berlin, one of the richest in Europe, amounted to upwards of 14,000 species, including 375 heaths. Among the Phanerogams were 1600 Synantheræ, 1150 Leguminosæ, 428 Labiatæ, 370 Umbelliferæ, 460 Orchidaceæ, 60 Palmaceæ, 600 Gramineæ and Cyperaceæ, &c. By comparing these data with the number of species described in the works of De Candolle, Walpers, Bentham, Lindley, Kunth, and others, we find that in the Berlin gardens are cultivated only one-seventh of the known species of the Synantheræ, one-eighth of the Leguminosæ, one-ninth of the Gramineæ, and about one-fiftieth of the smaller families, such as the Labiatæ and Umbelliferæ.
Now, if we admit that, on the one hand, the number of phanerogamous species cultivated in all the great gardens of Europe is about 30,000, and, on the other, that the cultivated Phanerogams form about one-eighth part of the species described in books and preserved in herbariums, we obtain a total of 24,000 species.
But the Cryptogams, or Agams, such as heaths, mosses, lichens, mushrooms, fungi, mould, and the like, of which our knowledge, as yet, is very imperfect, are probably much more numerous in species than the Phanerogams; for these vegetables, mostly microscopical, develop themselves wherever life can manifest itself—on the barren and denuded rocks, as well as in the air and in the depths of the ocean. If we suppose that they exceed only by 2000 the estimated number of Phanerogams, we shall obtain a total of just half-a-million!
Such, in our opinion, is the number which approximatively represents the lower limit of the aggregate of vegetable species (phanerogamous and cryptogamous) inhabiting our planet. The innumerable individuals of this half-million of species are born, and live, and reproduce their kind, and die, like the twelve hundred millions of individuals of our solitary human species. The former, it is true, remain fixed to the soil which has witnessed their birth, while the latter wander, more or less freely over the terrestrial surface. Do not animals enjoy the same privilege of locomotion? Undoubtedly. But men boast of the reason and the conscience with which they are endowed. Agreed. But with the exception of a small number—the infinite minority of progress—to what advantage have men employed the reason and the conscience of which they boast?
But this is a digression. We proceed to place before the reader a few final data in illustration of the subject we have been considering—the number of existing vegetable species.
The following is an estimate of the known species of plants on the globe at different dates:—
| Phanerogams. | Cryptogams. | Total. | |||
|---|---|---|---|---|---|
| According to | Linnæus, | 1753, | 5,323 | 615 | 5,938 |
| Pusoon, | 1807, | 19,949 | 6,000 | 25,949 | |
| Stendel, | 1824, | 39,684 | 10,765 | 50,649 | |
| Stendel, | 1841, | 78,000 | 13,000 | 91,000 | |
| Stendel, | 1844, | 80,000 | 15,000 | 95,000 |
The advance made of late years in the knowledge of existing species will be apparent from a consideration of Lindley's estimate in 1846:—
| Genera. | Species. | |
|---|---|---|
| Thallogens, | 939 | 8,394 |
| Acrogens, | 310 | 4,086 |
| Rhizogens, | 21 | 53 |
| Endogens, | 1,420 | 13,684 |
| Dictyogens, | 17 | 268 |
| Gymnogens, | 37 | 210 |
| Exogens, | 6,191 | 16,225 |
| ——— | ——— | |
| Total, | 8,935 | 92,920 |
According to Hinds, the following families are almost entirely restricted to particular divisions of the globe:—
To Europe—Globulariaceæ, Ceratophyllaceæ.
To Asia—Dipterocarpaceæ, Aquilariaceæ, Camelliaceæ, Moringaceæ, Stilaginaceæ.
To Africa—Bruniaceæ, Brexiaceæ, Belvisiaceæ, Penæaceæ.
To North America—Sarraceniaceæ.
To South America—Rhizobolaceæ, Gillesiaceæ, Calyceraceæ, Vochysiaceæ, Simarubaceæ, Monimiaceæ, Humiriaceæ, Papayaceæ, Gesneraceæ, Lacistemaceæ.
To Australasia—Goodeniaceæ, Epacridaceæ, Stackhousiaceæ, Brunoniaceæ, Tremandraceæ.
[A group of plants occurring only in one of the six great divisions of the world is called monomic, (from μονος one, and νομὸς, a region).
A group common to two divisions is dinomic; to three, trinomic; to four, quatrinomic; to all the divisions, polynomic.]