GEOLOGICAL TIME
Figures to the left denote millions of years that have elapsed up to recent time
| CENOZOIC | ||
| Age of Man | ||
| RECENT | Man and his Culture | |
| 1 | PLEISTOCENE | Last of Mammoths & Mastodons |
| Age of Mammals | ||
| 7 | PLIOCENE | Horses modernized |
| 20 | MIOCENE | Grasses and Grazing Animals |
| Three-toed Horses, Rhinos, Camels | ||
| 35 | OLIGOCENE | Specialization of Primitive Ancestors |
| 60 | EOCENE | Decline of archaic types |
| Mammals flourishing | ||
| MESOZOIC | ||
| Age of Reptiles | ||
| 125 | CRETACEOUS | Last of Great Reptiles |
| Specialization of Dinosaurs | ||
| 160 | JURASSIC | Bony Fishes thriving |
| Flowering plants advance | ||
| Cycads | ||
| Birds and Flying Reptiles | ||
| 200 | TRIASSIC | Few small mammals of lower orders |
| Dinosaurs become prominent | ||
| PALEOZOIC | ||
| Age of Amphibians | ||
| 225 | PERMIAN | Reptiles advancing |
| Amphibians dominant insects | ||
| 300 | CARBONIFEROUS | Dense forests of spore-bearing plants |
| Age of Fishes | ||
| 350 | DEVONIAN | Shark-like Fishes |
| Land floras established | ||
| 375 | SILURIAN | First land animals (scorpions) |
| Armored Fishes prominent | ||
| Age of Invertebrates | ||
| 425 | ORDOVICIAN | Corals and Bryozoa |
| Progress among Mollusks | ||
| 500 | CAMBRIAN | Brachiopods gaining |
| Trilobites dominant | ||
| Advance of shelled animals | ||
| PROTEROZOIC | ||
| EARLIEST LIFE | ||
| 1000 | UPPER PRE-CAMBRIAN | Small marine invertebrates |
| Lowest Forms of Plant and Animal Life | ||
| Few Fossils | ||
| ARCHEOZOIC | ||
| 2000 | LOWER PRE-CAMBRIAN | Some chemical evidence of life |
| No fossils | ||
Such aids have been devised and revised from time to time. No figures have been offered as final or absolutely “right” since the beginning of scientific investigations. Time divisions have been proposed that are not yet in common use while others have been abandoned or modified. Sources of information are so numerous that appropriate credit cannot be given fairly for anything that is up-to-date. The combined chart and outline here provided is based on time calculations of recent date but with figures slightly rounded off for the sole purpose of making them easier to remember. In view of the still existent probability of error it is felt that the slight alteration of figures may justify itself. It need not be regarded as misleading if the present purpose be considered—the stimulation of a natural history interest which is not vitally concerned with the little difference between a thousand million years and nine hundred ninety-nine million years.
The whole of geological time has been divided and subdivided according to varying practices. The development of life is perhaps the one outstanding feature of the time divisions, but for the most part the changes in floras and faunas have been gradual rather than abrupt, and this makes it very difficult to draw sharp lines or to visualize beginnings and endings of the various stages of development. Occasionally there is good excuse for drawing a line, where the record is broken and resumed again after a long lapse of time. The principal cause of such breaks is the elevation of great land masses, which brings on an interval of erosion and surface destruction for the areas uplifted.
These movements of parts of the earth’s crust have been exceptionally pronounced at certain times, often culminating in the production of mountain systems, and because of the extreme changes they introduce are known as revolutions. The major divisions of prehistoric time have been established, at least in part, by such revolutions; crustal, climatic, or other disturbances, on a smaller scale and recurring with greater frequency, may be regarded as establishing boundaries for the minor divisions. Hence we have five great Eras of geological history, and these are divided again into Periods. The time chart shows an arrangement commonly used in America. In the first column the names of the Eras are stated in technical form. Closely coinciding with these terms are the popular names of the Ages which appear in the second column. These names, describing the dominant life of each age, are very convenient. The more scientific terms used for the eras, while serving essentially the same purpose, are a little more systematic and generalized in that they refer to ancient life (Paleozoic), middle life (Mesozoic), and recent life (Cenozoic), without being specific as to any class of animals or plants for any one division of time.
The period names, in the central column, have been derived from miscellaneous sources, some of them from geographical districts, some from descriptive references to prominent features of the rocks, others indicating a degree of approach to recent time. In paleontology (fossil study) it has long been a practice to cut the periods into lower, middle, and upper divisions, and in a few cases it has been found desirable to make two periods out of an old one. What was once known as the Lower Carboniferous is now commonly recognized as the Mississippian period while the upper portion has become the Pennsylvanian. The Lower Cretaceous is now the Comanchean of some authors.
Both old and new practices are responsible for a little confusion at the present time. A former division into Primary, Secondary, Tertiary, and Quaternary eras has been partly abandoned, but the term “Quaternary” still applies to the Age of Man, while “Tertiary time” remains in good usage for the balance of the Cenozoic era. Among the newer introductions may be mentioned the use of a Paleocene period which precedes the Eocene. Geologists are not entirely in agreement as to the necessity for this addition and many would not give it equivalent rank with other periods. In the interest of simplicity these modern refinements have been omitted from the chart.
The figures appearing in the third column, between the Ages and Periods, indicate the millions of years that have elapsed up to present time. They denote the age of the rocks at the beginning of each period. The age of a plant or animal which lived in Eocene time would be, according to this scale, somewhere between 35 million and 60 million years. In practice it is usually possible to determine whether a fossil was embedded in the rocks during an early or late portion of the period, and thus its age may be established within a shorter range, but it is impossible to be exact, even in terms of millions of years, with regard to anything as far back in prehistory as the Eocene period.
The period in which we are living today is known as Recent. It began at the close of the Ice Age or Pleistocene period about ten thousand years ago and represents so little of earth history since the beginning of life that a chart many times the length of this page would be required to show the rest of the periods in proportion. The Cambrian period is an early chapter in which the story of prehistoric life suddenly becomes clear and richly varied. It is, however, much farther from the beginning of the record than it is from the present, and the Pre-Cambrian eras would require a great deal more space in order to show their relative lengths. The Archeozoic and Proterozoic eras have to some extent been divided into periods, but the great antiquity of the rocks has obscured much of their history, and divisions established for one locality have been of little service elsewhere. Consequently, the period names are in less general use and the common practice is to refer to all this great stretch of time as Pre-Cambrian.
In the last column, at the right of the chart, some of the historical features are indicated. This column should be read from bottom to top in order to get the proper development of the story, and at best this sketchy outline of events can be no more than suggestive of the progress and decline through which the earth’s inhabitants have passed.
Rocks of every period except probably the Silurian are known to have been deposited somewhere in the Colorado area, although in most cases the record for each period is far from complete. Formations are too numerous and too varied locally to be shown on a chart of this type.
In the study of fossils there are two important field aids usually available. For any locality there should be a geological map and a section showing the sequence and character of the strata. On a small-scale map many of the local details have to be omitted, but the position of the larger exposures is indicated and, with this information at hand, the fossil-bearing strata may be located with the help of a geological section. The latter is frequently obtained from technical reports published by State and National Geological Surveys. Frequently, however, it is possible to obtain only a general plan for a given locality, and a great deal of literature may have to be scanned in order to get that. Excellent geological maps of Colorado have been published by the Colorado Geological Survey and the United States Geological Survey.
It often happens that a formation is not where we expect to find it, this being due to several possible factors. The sediments may not have been deposited there, or they may have been removed by erosion. Where the structure has been disturbed by folding and faulting, a multitude of complications is introduced. The expected sequence is sometimes inverted and repeated through a series of folds. Formations also may be moved miles out of place by faulting. Both thickness and character of sediments may vary considerably within a formation. In some regions the geology is very simple, in others extremely difficult to understand.
FORMATIONS of the DENVER-FOOTHILLS REGION
A GENERALIZED SECTION SHOWING SOME OF THE SURFACE FEATURES
RED BEDS HOGBACKS TABLE MOUNTAIN DENVER DISTRICT
REGION OF MOUNTAIN-MAKING UPLIFT
Formations bordering the mountains have been bent into upright positions.
| PERIODS | ||||
|---|---|---|---|---|
| RECENT | ||||
| PLEISTOCENE | ||||
| PLIOCENE | ||||
| MIOCENE | ||||
| OLIGOCENE | ||||
| EOCENE | ||||
| CRETACEOUS | THICKNESS | |||
| SOFT SANDSTONES GRITS & CLAYS | DENVER & ARAPAHOE | 2000 ft. | ||
| SANDSTONES, SHALES & LIGNITE | LARAMIE | 1000 ft. | ||
| YELLOWISH SANDS & SHALES | FOX HILLS | 1000 ft. | ||
| SOFT DARK GRAY OR RUSTY SHALE | PIERRE | 5000 ft. | ||
| LIMESTONES & SHALES | NIOBRARA | 500 ft. | ||
| DARK SHALES & LIME | BENTON | 400 ft. | ||
| GRAY OR BUFF SANDSTONES & CLAYS | DAKOTA | 300 ft. | ||
| SHALES, SANDSTONE & LIME | MORRISON | 200 ft. | ||
| JURASSIC | ||||
| TRIASSIC | ||||
| PERMIAN | ||||
| DEEP-RED SANDY SHALES, LIME, GYPSUM | LYKINS | 700 ft. | ||
| CARBONIFEROUS | ||||
| MASSIVE PINK OR WHITE SANDSTONE | LYONS | 200 ft. | ||
| RED OR BROWN SANDSTONE & CONGLOMERATE | FOUNTAIN | 1500 ft. | ||
| DEVONIAN | ||||
| SILURIAN | ||||
| ORDOVICIAN | ||||
| CAMBRIAN | ||||
| PRE-CAMBRIAN | ||||
| METAMORPHIC & INTRUDED ROCKS SCHIST, GNEISS, QUARTZITE | IDAHO SPRINGS (PART) | |||
| BASEMENT ROCKS of IGNEOUS ORIGIN |
A generalized section for the western part of the Denver Basin is introduced here for the use of local students. The formations normally present in this region are shown in their usual position. They are briefly described on the chart, and their thickness is indicated by figures which may be regarded as near the maximum for the district. The section will apply to most of the foothills area between Morrison and Boulder though surface features and thickness of beds will vary considerably from place to place.
Certain of the formations are known to be fossil bearing, others barren or nearly so. When fossils are present they are usually restricted to certain localities, and these may be widely scattered. The following remarks apply to the possibilities for finding fossils in the formations named.
Leaf impressions of palms, ferns, and numerous species of well-known trees and shrubs are common in many localities. Petrified wood is fairly abundant, and a few scattered bones of reptiles and mammals have been found. The two formations are treated as a unit because the Arapahoe is neither conspicuous nor sharply defined. Denver beds are well exposed on the slopes of Table Mountain at Golden; fossils, however, have been obtained from several localities nearer the city of Denver, notably from the hills just west of Overland Park.
Plant material is locally abundant, principally the leaves of familiar deciduous trees, palms, and ferns. Many of the clay pits being worked near Golden are in this formation. Oysters and a few other mollusks may be found in some places.
Better exposures of this formation are located to the north of Denver. Marine mollusks are most frequently found.
In addition to the characteristic dark shales, this formation includes some limy material and sandstone beds, both of which are fossiliferous in places. Two types of marine mollusks are characteristic: Inoceramus, generic name for several species of clam-like bivalves readily identified by concentric elevations which produce a rippled effect on the shell surfaces; and Baculites, cephalopods with straight, chambered shells which often break at the suture lines, where the fossil is weakened by the chamber walls. Small oyster shells are fairly common also. The formation is to be found some distance to the east of the prominent hogback where it weathers into smooth surfaces in the form of broad valleys and flats, with rounded contours on the few elevations that may be present. It forms a soft, flaky soil when dry, is a sticky “gumbo” when wet. The clay is generally of a rather dark grayish color when freshly exposed but it takes on a rusty appearance after weathering. At various levels there are numerous iron-cemented concretions, many of which contain fossil shells.
Marine Beds of the Benton Formation, Northeastern Colorado
A stream channel has cut deeply into the formation, uncovering and partly destroying a plesiosaur skeleton which was found at the level where the men are standing.
Plesiosaur Bones in Place
Benton formation. Surface rubble has been cleared away, and several vertebrae are partially uncovered in the area at the right of the hammer.
The formation contains fossils rather similar to those of the Pierre. Shark’s teeth have been found in some of the lower beds. Limestone is a prominent feature, often forming a well defined ridge near the foot of the eastern slope of the main hogback. The limestones commonly have a chalky character.
The formation is not especially productive in this region. Marine shells are numerous in some localities, and bones of marine reptiles have been found at various places. As usually seen, it is almost entirely composed of impure clay shales, very dark, brownish-gray to almost black, and commonly interbedded with thin patches of white bentonite, yellow ochre, gypsum, and limestone.
This formation produces the high hogback which is usually present some distance east of the Red Rocks. There are generally two or three layers of massive, light-colored sandstone separated by clays which are used extensively in the making of bricks and pottery. Leaf impressions and some fish scales are found in the clays and occasionally in the sandstone. The hogback is a good marker from which to locate other formations, because of its prominence in the foothills landscape.
Good dinosaur material has been taken from the Canon City and Morrison districts. The formation is to be found on the lower west slope of the Dakota hogback. It consists of continental deposits of the stream and lake types. There is considerable sandstone in this formation and a little limestone is to be found here and there, but the most characteristic feature is in the shales. When freshly exposed, the shales are delicately tinted with gray, green, and maroon, a bronze-green being rather prominent. This formation is highly variable in character, with much of the clay often buried under the valley floor. In addition to the bones of reptiles, there are plant fossils, usually of poor quality, and fresh-water gastropods more or less abundant in some localities.
Outcrops are not prominent, owing to the small amount of weather-resisting materials. The sandy clays are commonly of a deep red color mottled with spots of light gray. A white limestone is sometimes present near the middle of these deposits, and gypsum beds are included locally. The formation is often indicated only by red soil in the depressions between ridges. Few fossils have been reported.
This formation is usually prominent as the eastern wall of the uplifted Red Rocks series. In some localities it forms a ridge of pink or white sandstone distinctly separated from the older sediments to the west. Very few fossils are found.
Exposures usually are brown to red in color, though sometimes a dirty white. The prominent rocks are rather coarse sandstone, commonly with a gritty texture due to the angular character of the sand or gravel from which they were made. These are the westernmost of the Red Beds and the oldest of the uplifted sedimentary rocks bordering the foothills in most of our area. Fossils have been found in the formation, but it is practically barren for the territory here considered.
* * * * * * * *
This geological section also illustrates a method of dating crustal movements and the birth of mountain ranges, for the folding of the strata along the flanks of the Rocky Mountains has a great deal of significance in this connection. The sedimentary layers were originally deposited over much of the present mountain area in a horizontal position, and only those formations in existence at the time could be distorted by the upheavals which produced the new elevations. Of the series generally involved in the movement the Laramie beds are the youngest. Since these beds had not been formed until near the close of the Cretaceous period it is to be assumed that the mountains must be of more recent date, younger than the topmost of the deformed beds and at least as old as the lowermost of the undisturbed formations overlying them.
Some disturbance is evident also in the Arapahoe and Denver beds which overlie the Laramie, but this is believed to have occurred sometime after the occasion of the first great uplift. Volcanic materials in these beds lead to the belief that the sediments were deposited during a period of volcanic activity brought on by the crustal folding which terminated the Mesozoic era. Hence the conclusion arises that the age of the Denver and Arapahoe beds must coincide closely with some of the earlier stages in the history of the mountain system. This interval is often referred to as Post-Laramie time.
The rocks of Pre-Cambrian time have been buried deeply under the accumulation of younger sediments, and the resulting pressure in many places has been tremendous. In addition to the effects of pressure there also is recorded in these ancient formations the repeated movements of the materials since they were first deposited. Vertical and side adjustments of parts, with relation to other parts, have distorted the original arrangement of the rock particles to such an extent that ordinary fossils would eventually become unrecognizable. These crushing, grinding, and kneading forces working through millions of years alone would account for the absence of fossils from the older deposits. Frequently the rocks have become so changed in form that their original character can only be conjectured, and because of this change they are known as metamorphic rocks.
A few beds of Archeozoic age remain in nearly their original condition, but they are either without fossils or they have produced very questionable and unsatisfactory specimens. The existence of life during these early stages of earth history is indicated largely by chemical rather than fossil evidence. Much of the ancient limestone has been converted into marble, but it is not unreasonable to believe that plants and animals were instrumental in the production of this type of rock as they are today. Certain varieties of iron ore deposits are now being built up by the aid of plants, and similar ores in the ancient rocks may have had a like origin. The presence of great quantities of carbon, in the form of graphite, may be regarded also as a sign of life, for this substance is accumulated on a large scale by living plants, and may be retained in a solid form after the partial decay of the plant tissues.
So far as the direct evidence goes, there is no sign of any creature of large size or of such complicated structure as the common plants and animals of today. The chemistry of the mineral deposits is not entirely convincing as to the presence of life, but it is regarded as highly probable that microscopic, single-celled plants and animals, comparable to modern algae and protozoa, were in existence during Archean time. Throughout later eras there is unmistakable evidence of gradual development from simpler to more elaborate life-forms and the Archeozoic is commonly regarded as a time of preparation during which simple organisms of some kind were becoming adapted to early conditions which could not support life on a higher plane. The importance of the work done by such lowly creatures in the preparation of suitable environments for more advanced modes of living is overlooked almost entirely.
During the next era, the Proterozoic, the record of life becomes somewhat clearer. Fossils are hardly to be regarded as abundant but there were several well-defined types of animals which left shells and other parts composed of mineral matter. Among these may be mentioned the Radiolaria, Foraminifera, Bryozoa, and Sponges. Radiolaria produced delicate, often lace-like shells of many patterns adorned with the radiating filaments or spines which have suggested the name for this group. Foraminifera produced minute shells, sometimes many chambered, and often bearing a confusing resemblance to the work of snails. Common chalk is composed almost entirely of such shells and fragments of them.
Sponges and Bryozoa are animals of slightly higher organization. They are many-celled instead of one-celled and the cells have special work to perform, which is a most important step in the direction of the specialization which characterizes the structural and life pattern of later arrivals. The Bryozoa lived in moss-like colonies which have been important rock-makers; the fossil forms bear some resemblance to corals. Sponges are too well known to require description although the familiar article of commerce is merely the framework of once-living animals. They represent the earliest organization of true animal bodies even though in appearance they may have a resemblance to plants.
Actual plants of this era were of the algae class, aquatic in habit as were their animal neighbors, the first to leave a record in the form of fossils. This record, obscure and distorted, has long been a source of perplexity to investigators. Without well-defined floras and faunas to guide them, and with rocks frequently in chaotic relationships, early geologists were content to regard it all as a “Pre-Cambrian complex.” Recent studies have contributed a great deal of information not available some years ago. It is quite possible that more advanced types of life were in abundance before the close of the second era, but material on which to base sound opinion is still scarce.
Rocks of Pre-Cambrian age are plentiful in the foothills region west of Denver. The schists, gneisses, and quartzites exposed for some miles immediately beyond the red-beds are part of this great complex. The Idaho Springs formation is known to be one of the oldest in this district, although its exact age has not been determined. Other formations are recognized among the metamorphic rocks of the region but none has contributed to our knowledge of early life.
There can be no mistake as to the prolific development of life in Cambrian seas, for fossils of this age are to be found in many parts of the world, where ancient sea bottoms now form part of the land surface. Invertebrate animals appear to have made much progress, but plants were either scarce or too small and delicate to be productive of fossils. It is probable, however, that seaweeds and other algae were flourishing along with the invertebrates, because animal life is directly or indirectly dependent on the existence of plants. The latter sustain themselves by taking carbon and nitrogen from air, water, and soil, but animals must obtain their requirements by eating plants or eating each other. They cannot obtain what they need from the inorganic world without this help from the vegetable kingdom.
One group of animals stands out prominently above all its contemporaries. Known as the trilobites they were by far the most distinguished and most characteristic of Cambrian invertebrates. Trilobites inhabited the warmer seas of this period and several later ones, but were extinct by the end of the Paleozoic era. Hundreds of species have been described, most of them under four inches in length. Well-known distant relatives now living are the shrimps, and other crustaceans. The name Trilobite has reference to the three lobes which are apparent in the form of the upper surface, the central lobe forming a broad ridge extending along the back. Beneath the outer lobes on each side there was, during life, a row of short, jointed legs used for swimming and walking, but these delicate appendages are seldom preserved in the fossils.
Second in importance among the animals of the period were the brachiopods or lamp-shells, not true mollusks although they were provided with similar shells composed of calcium phosphate or calcium carbonate. Shells are of two parts (bivalved) as in the case of clams, but the valves are above and beneath the body instead of on the right and left sides, which is the arrangement among mollusks. Although abundant as individuals, there were only a few species during the earlier part of the period; the number of species increased, however, and the race became very persistent. About seven thousand species have been described, and the race is not yet extinct although the number of living species is relatively small.
Cambrian life evidently included representatives of all the great divisions of invertebrates; sponges, jelly-fishes, worms, and primitive corals have been reported. At the end of the period there was an elaborate molluscan fauna. The closing of the period in North America was apparently a gentle elevation of continental areas and a consequent withdrawal of the sea.
Invertebrate Fossils
Only a few prominent types have been selected from thousands of invertebrates known to zoologists. The forms illustrated are of frequent occurrence as fossils.
Extensive land areas must have subsided again early in the Ordovician period for marine sediments were laid down over a large part of the North American interior, and three epochs or subdivisions of the period have been based on as many invasions of the sea. In these ancient deposits the record of life continues to show new forms. Nothing of a very spectacular sort is recorded other than a great increase in the number of species among types that were established in earlier periods.
Trilobites were at their best, brachiopods continued to flourish, and the mollusks made new progress, especially the cephalopods, a group which includes our cuttle fishes and squids. Some of these predatory creatures attained large size and were no doubt masters of the sea. Typical forms were provided with tapering chambered shells that occasionally reached a length of twelve or more feet. Most of the shells were straight and trumpet-like or but slightly curved. Some were closely coiled and in this respect more like the well-known nautilus now in existence.
The bryozoans became very common in the later part of the period and corals made slight advances. Somewhat of a novelty at this time were the crinoids, commonly known as “stone lilies” although not plants at all. They have been described as starfishes with back turned downward and a thick stem attached beneath. Where they lived in great abundance the limestone deposits may consist almost entirely of their stems. Crinoids continued to produce a variety of forms throughout several of the succeeding periods.
The brachiopods were commonest of all animals representing this period, however, and their wide distribution, together with their known preference for warm waters, is taken to be an indication of mild temperatures prevailing over a large portion of the earth. Land plants are indicated by spore-bearing forms related to the ferns and mosses. Impressions of such plants have been found in Europe but, since most of the known rocks of this age were formed in seas, the marine algae are more abundant as fossils.
In the Colorado area, rocks of Ordovician time are exposed only in mountainous areas where they have been lifted high above their original levels. They are not especially rich in fossils although they have produced some fish remains which are of interest in that they suggest an age of vertebrates which is just ahead.
Since land floras and faunas had not yet become conspicuous the fossil record for this period is limited to those areas which were invaded by the sea. Apparently there was no such invasion of the present Colorado region, for rocks of this age are not in evidence. If they exist at all they are restricted to localized districts which are deeply buried under sediments of later periods. There may have been no Silurian deposition in this area, or such rocks may have been produced only to be destroyed by elevation and consequent subjection to weathering and erosion during a long interval of time, in which they were completely removed. In the region of the Colorado Rockies there is no evidence of returning seas until late Devonian time.
In other parts of the world, however, there was extensive deposition of rock-making sediments in seas which were inhabited by algae and invertebrates of the types previously described. Among the common animals of the time there were still numerous species of brachiopods, trilobites, corals, crinoids, and bryozoans. In addition to the primitive cone-shaped, cup corals there were several advanced types but the habit of building large reefs was not yet established.
“Sea scorpions,” really large crustaceans, flourished during Silurian time, and late in the period there appeared a race of true scorpions which lived on dry land or between high and low tides along the seashore. These were smaller and much like modern descendants but probably they did not wander far from the ocean shores where an abundance of food was available. These little scorpions, the largest measuring only two and a half inches in length, are the oldest air-breathing land animals of the fossil record.
It was not until the period was well advanced that fishes became numerous, and much of our knowledge of the beginning of an “Age of Fishes” has been obtained from European fossils. Although fishes are classed with the vertebrate or backboned animals there are large groups which do not have bony skeletons but are provided instead with a simple framework of cartilage. Among the earlier and more primitive types were the ostracoderms or bony-skinned fishes with no internal bones and only a small amount of bony substance in the armor-like plates and scales which covered the forward portion of the body.
The ostracoderms comprise a small group of fishes about which very little is known. They appear to have been inhabitants of fresh-water streams as well as lagoons bordering the seas, and may have been related to the small sharks of the time. They lived during the Ordovician, Silurian, and Devonian periods, and left no descendants now recognized among living creatures. A much larger type of armored fishes is known as the arthrodires, a name which refers to a pointed neck and an arrangement of the armor plates to permit a movement of the head. These were the most ferocious fishes of the Silurian and Devonian seas, some of them reaching a length of twenty feet though most were much smaller. Their jaws were provided with formidable shearing and crushing plates instead of teeth.
The Devonian is one of the most outstanding of all periods from the viewpoint of life development. Dominance of the fishes is its greatest achievement, the invertebrates remaining about as they were and the higher vertebrates barely in evidence, but life on a large scale was no longer confined to the seas. Fresh-water fishes became prominent and land plants well established. The first forests appeared, with fern-like plants predominating although woody trees of several types and considerable size were included. It is quite possible that extensive land areas had been well supplied with vegetation during earlier times, but the delicate tissues of plants are far less likely to be preserved than the limy parts of animals. The fossil record, therefore, cannot be expected to reveal more than a suggestion of the progress made at this level of living. The story of plant life becomes much clearer in the next period when conditions were more favorable for the production and preservation of plant fossils.
Land animals of the time are almost unknown. A few snails and scorpions have been found, and some footprints made by early amphibians. Insects probably were in existence although the evidence is not quite clear on this point. The increasing number of fresh-water fishes, however, may be regarded as a sure indication that inland conditions were becoming more favorable for plant and animal inhabitants of all kinds.
The extent of development among the fishes cannot be accurately indicated by naming a few types, for it is mainly in the number of species and genera within the larger groups that progress is seen. In general it may be stated that the fishes of the period had not yet acquired the bony skeleton and typical form of familiar modern species. Skeletons were of cartilage, partly hardened in some instances by lime. Armor plates were customary with certain races but were not present among all fishes. Neither were these armored forms exceptionally large, as compared with living sharks. Although occasional giants appeared, the majority were small. Many were sluggish creatures with poorly-developed jaws, living as scavengers on sea and stream bottoms. Tail fins were usually unbalanced as in the sharks, or pointed and rounded rather than evenly forked.
Modernized Types of Fishes from Eocene Shales of Southwestern Wyoming
The great tribe of true bony fishes, such as the cod and perch, which includes more than ninety percent of the fishes living today, was not yet in existence. About one-third of the many kinds of fishes then living were related to the sharks, a group which is relatively insignificant in recent years. Nearly one-fourth of the total belonged to a tribe of enamel-scaled fishes, now represented only by a few sturgeon and gar-pike.
Lung fishes have never been a large group but it is noteworthy that they have had existence since Middle Devonian time. Living members of the race, inhabitants of Africa and South America, make a practice of burrowing into the mud of stream channels during dry seasons and are provided with lungs which enable them to breathe air in the manner of higher vertebrates. They survive the complete drying-up of the streams and live for months without water. Other forms, with less development of lungs, frequent stagnant pools and come to the surface occasionally for a breath of air. All are provided with gills also, which enables them to obtain their oxygen as other fishes do. They are believed to be a connecting link between the fishes and the early amphibians. More accurately, perhaps, they should be regarded as holding an intermediate position without being directly ancestral to any higher type of vertebrate animal.
Still dominant among the invertebrates were the brachiopods, on the whole averaging a little larger in size, and otherwise indicating congenial times for that type of organism. They reached the peak of their development during this period. Trilobites were declining although a few new and strangely ornamented varieties made a brief appearance. Crinoids apparently found living conditions less favorable during Devonian time, but in a later era they again became prominent. Corals were favored only at times and in certain localities. Along with the crinoids they appear to have suffered from the presence of an unusual amount of mud in the waters of their customary habitats. Both had a preference for clear water as indicated by the absence of fossils from limestones containing more than a very small percentage of muddy sediments. Crustaceans, similar to the sea-scorpions and better known as eurypterids, became prominent among fresh-water animals. Some were unusually large for creatures of this class, lengths of several feet being recorded from fragments. Gastropod mollusks came into prominence in localities where living conditions were favorable. Bivalves continued to thrive but the cephalopods had a rather meager development considering the heights they were to achieve in subsequent periods.
In western North America the large expanse of territory known as the Great Plains was evidently well above sea level during this entire period, for no beds of this age are found in eastern Colorado. West of the Front Range, however, there was some deposition of marine sediments during late Devonian time. Formations of this age are exposed near Salida and Glenwood Springs, on the White River Plateau, and in the San Juan region.
The Carboniferous period gets its name from the vast deposits of coal which were developed during that time in many parts of the northern hemisphere. Depressed land surfaces bordering the continents, and extending well into the interior of present boundaries, supported dense growths of vegetation and provided the swampy conditions most favorable to coal production. Varieties of plants which are now of small size and lowly position in the botanical world acquired the proportions of large trees.
Best-known fossils of the period are carbonized portions of the larger trees, and impressions left in the muds and sands of ancient bogs. Forest trees of several kinds reached the height of a hundred feet, with a trunk diameter of two to six feet. This size often is exceeded in modern forests, but by trees of an entirely different type. Considering the amount of development among the plants of earlier periods, Carboniferous forests provide an outstanding spectacle of advancing life.
Quite common among the larger trees were two varieties of club-mosses, also known as scale trees. They were cone-bearing evergreens with only slight resemblance to modern conifers. Instead of seeds they produced spores, a method of reproduction which is practiced among ferns. The trunks were marked from bottom to top with uniform patterns of cushions and scars indicating the points at which leaves were attached during the earlier stages of growth. In the Lepidodendrons the rows of scale-like cushions wind spirally upward while among the Sigillaria there is a vertical arrangement of leaf-scars which resemble the imprints of a seal, these impressions being in straight and parallel rows on a surface which may be either ribbed or smooth. The leaves of scale trees were stiff and slender, and arranged in grass-like tufts at the top.
Calamites, related to our horsetail rushes, were somewhat smaller than the scale trees. Their trunks consisted of a thin, woody cylinder with a pithy interior, and were marked at intervals by nodes which gave them the “jointed” appearance of a bamboo stem. Leaves were arranged in circles around the nodes of main stem or branches. Spore-bearing cones appeared at the tips of the stems.