THE VARIATION OF ANIMALS AND PLANTS UNDER DOMESTICATION.

By Charles Darwin

IN TWO VOLUMES.-Vol. I.

WITH ILLUSTRATIONS



CONTENTS OF VOLUME I.

FOOTNOTES

INTRODUCTION ... Page 1

CHAPTER I.

DOMESTIC DOGS AND CATS.

ANCIENT VARIETIES OF THE DOGRESEMBLANCE OF DOMESTIC DOGS IN VARIOUS COUNTRIES TO NATIVE CANINE SPECIESANIMALS NOT ACQUAINTED WITH MAN AT FIRST FEARLESSDOGS RESEMBLING WOLVES AND JACKALSHABIT OF BARKING ACQUIRED AND LOSTFERAL DOGSTAN-COLOURED EYE-SPOTSPERIOD OF GESTATIONOFFENSIVE ODOURFERTILITY OF THE RACES WHEN CROSSEDDIFFERENCES IN THE SEVERAL RACES IN PART DUE TO DESCENT FROM DISTINCT SPECIESDIFFERENCES IN THE SKULL AND TEETHDIFFERENCES IN THE BODY, IN CONSTITUTIONFEW IMPORTANT DIFFERENCES HAVE BEEN FIXED BY SELECTIONDIRECT ACTION OF CLIMATEWATER-DOGS WITH PALMATED FEETHISTORY OF THE CHANGES WHICH CERTAIN ENGLISH RACES OF THE DOG HAVE GRADUALLY UNDERGONE THROUGH SELECTIONEXTINCTION OF THE LESS IMPROVED SUB-BREEDS.

CATS, CROSSED WITH SEVERAL SPECIESDIFFERENT BREEDS FOUND ONLY IN SEPARATED COUNTRIESDIRECT EFFECTS OF THE CONDITIONS OF LIFEFERAL CATSINDIVIDUAL VARIABILITY ... Page 15

CHAPTER II.

HORSES AND ASSES.

HORSE.DIFFERENCES IN THE BREEDSINDIVIDUAL VARIABILITY OFDIRECT EFFECTS OF THE CONDITIONS OF LIFECAN WITHSTAND MUCH COLDBREEDS MUCH MODIFIED BY SELECTIONCOLOURS OF THE HORSEDAPPLINGDARK STRIPES ON THE SPINE, LEGS, SHOULDERS, AND FOREHEADDUN-COLOURED HORSES MOST FREQUENTLY STRIPEDSTRIPES PROBABLY DUE TO REVERSION TO THE PRIMITIVE STATE OF THE HORSE.

ASSES.BREEDS OFCOLOUR OFLEG- AND SHOULDER-STRIPESSHOULDER-STRIPES SOMETIMES ABSENT, SOMETIMES FORKED ... Page 49

CHAPTER III.

PIGS—CATTLE—SHEEP—GOATS.

PIGS BELONG TO TWO DISTINCT TYPES, SUS SCROFA AND INDICATORF-SCHWEINJAPAN PIGFERTILITY OF CROSSED PIGSCHANGES IN THE SKULL OF THE HIGHLY CULTIVATED RACESCONVERGENCE OF CHARACTERGESTATIONSOLID-HOOFED SWINECURIOUS APPENDAGES TO THE JAWSDECREASE IN SIZE OF THE TUSKSYOUNG PIGS LONGITUDINALLY STRIPEDFERAL PIGSCROSSED BREEDS.

CATTLE.ZEBU A DISTINCT SPECIESEUROPEAN CATTLE PROBABLY DESCENDED FROM THREE WILD FORMSALL THE RACES NOW FERTILE TOGETHERBRITISH PARK CATTLEON THE COLOUR OF THE ABORIGINAL SPECIESCONSTITUTIONAL DIFFERENCESSOUTH AFRICAN RACESSOUTH AMERICAN RACESNIATA CATTLEORIGIN OF THE VARIOUS RACES OF CATTLE.

SHEEP.REMARKABLE RACES OFVARIATIONS ATTACHED TO THE MALE SEXADAPTATIONS TO VARIOUS CONDITIONSGESTATION OFCHANGES IN THE WOOLSEMI-MONSTROUS BREEDS.

GOATS.REMARKABLE VARIATIONS OF ... Page 65

CHAPTER IV.

DOMESTIC RABBITS.

DOMESTIC RABBITS DESCENDED FROM THE COMMON WILD RABBITANCIENT DOMESTICATIONANCIENT SELECTIONLARGE LOP-EARED RABBITSVARIOUS BREEDSFLUCTUATING CHARACTERSORIGIN OF THE HIMALAYAN BREEDCURIOUS CASE OF INHERITANCEFERAL RABBITS IN JAMAICA AND THE FALKLAND ISLANDSPORTO SANTO FERAL RABBITSOSTEOLOGICAL CHARACTERSSKULLSKULL OF HALF-LOP RABBITSVARIATIONS IN THE SKULL ANALOGOUS TO DIFFERENCES IN DIFFERENT SPECIES OF HARESVERTEBRÆSTERNUMSCAPULAEFFECTS OF USE AND DISUSE ON THE PROPORTIONS OF THE LIMBS AND BODYCAPACITY OF THE SKULL AND REDUCED SIZE OF THE BRAINSUMMARY ON THE MODIFICATIONS OF DOMESTICATED RABBITS ... Page 103

CHAPTER V.

DOMESTIC PIGEONS.

ENUMERATION AND DESCRIPTION OF THE SEVERAL BREEDSINDIVIDUAL VARIABILITYVARIATIONS OF A REMARKABLE NATUREOSTEOLOGICAL CHARACTERS: SKULL, LOWER JAW, NUMBER OF VERTEBRÆCORRELATION OF GROWTH: TONGUE WITH BEAK; EYELIDS AND NOSTRILS WITH WATTLED SKINNUMBER OF WING-FEATHERS, AND LENGTH OF WINGCOLOUR AND DOWNWEBBED AND FEATHERED FEETON THE EFFECTS OF DISUSELENGTH OF FEET IN CORRELATION WITH LENGTH OF BEAKLENGTH OF STERNUM, SCAPULA, AND FURCULALENGTH OF WINGSSUMMARY ON THE POINTS OF DIFFERENCE IN THE SEVERAL BREEDS ... Page 131

CHAPTER VI.

PIGEONS—continued.

ON THE ABORIGINAL PARENT-STOCK OF THE SEVERAL DOMESTIC RACESHABITS OF LIFEWILD RACES OF THE ROCK-PIGEONDOVECOT-PIGEONSPROOFS OF THE DESCENT OF THE SEVERAL RACES FROM COLUMBA LIVIAFERTILITY OF THE RACES WHEN CROSSEDREVERSION TO THE PLUMAGE OF THE WILD ROCK-PIGEONCIRCUMSTANCES FAVOURABLE TO THE FORMATION OF THE RACESANTIQUITY AND HISTORY OF THE PRINCIPAL RACESMANNER OF THEIR FORMATIONSELECTIONUNCONSCIOUS SELECTIONCARE TAKEN BY FANCIERS IN SELECTING THEIR BIRDSSLIGHTLY DIFFERENT STRAINS GRADUALLY CHANGE INTO WELL-MARKED BREEDSEXTINCTION OF INTERMEDIATE FORMSCERTAIN BREEDS REMAIN PERMANENT, WHILST OTHERS CHANGESUMMARY ... Page 180

CHAPTER VII.

FOWLS.

BRIEF DESCRIPTIONS OF THE CHIEF BREEDSARGUMENTS IN FAVOUR OF THEIR DESCENT FROM SEVERAL SPECIESARGUMENTS IN FAVOUR OF ALL THE BREEDS HAVING DESCENDED FROM GALLUS BANKIVA-REVERSION TO THE PARENT-STOCK IN COLOURANALOGOUS VARIATIONSANCIENT HISTORY OF THE FOWLEXTERNAL DIFFERENCES BETWEEN THE SEVERAL BREEDSEGGSCHICKENSSECONDARY SEXUAL CHARACTERSWING- AND TAIL-FEATHERS, VOICE, DISPOSITION, ETC.OSTEOLOGICAL DIFFERENCES IN THE SKULL, VERTEBRÆ, ETC.EFFECTS OF USE AND DISUSE ON CERTAIN PARTSCORRELATION OF GROWTH ... Page 225

CHAPTER VIII.

DUCKS—GOOSE—PEACOCK—TURKEY—GUINEA-FOWL—CANARY-BIRD—GOLD-FISH—HIVE-BEES—SILK-MOTHS.

DUCKS, SEVERAL BREEDS OFPROGRESS OF DOMESTICATIONORIGIN OF, FROM THE COMMON WILD-DUCKDIFFERENCES IN THE DIFFERENT BREEDSOSTEOLOGICAL DIFFERENCESEFFECTS OF USE AND DISUSE ON THE LIMB-BONES.

GOOSE, ANCIENTLY DOMESTICATEDLITTLE VARIATION OFSEBASTOPOL BREED.

PEACOCK, ORIGIN OF BLACK-SHOULDERED BREED.

TURKEY, BREEDS OFCROSSED WITH THE UNITED STATES SPECIESEFFECTS OF CLIMATE ON.

GUINEA-FOWL, CANARY-BIRD, GOLD-FISH, HIVE-BEES.

SILK-MOTHS, SPECIES AND BREEDS OFANCIENTLY DOMESTICATEDCARE IN THEIR SELECTIONDIFFERENCES IN THE DIFFERENT RACESIN THE EGG, CATERPILLAR, AND COCOON STATESINHERITANCE OF CHARACTERSIMPERFECT WINGSLOST INSTINCTSCORRELATED CHARACTERS ... Page 276

CHAPTER IX.

CULTIVATED PLANTS: CEREAL AND CULINARY PLANTS.

PRELIMINARY REMARKS ON THE NUMBER AND PARENTAGE OF CULTIVATED PLANTSFIRST STEPS IN CULTIVATIONGEOGRAPHICAL DISTRIBUTION OF CULTIVATED PLANTS.

CEREALIA.DOUBTS ON THE NUMBER OF SPECIES.WHEAT: VARIETIES OFINDIVIDUAL VARIABILITYCHANGED HABITSSELECTIONANCIENT HISTORY OF THE VARIETIES.MAIZE: GREAT VARIATION OFDIRECT ACTION OF CLIMATE ON.

CULINARY PLANTS.CABBAGES: VARIETIES OF, IN FOLIAGE AND STEMS, BUT NOT IN OTHER PARTSPARENTAGE OFOTHER SPECIES OF BRASSICA.PEAS: AMOUNT OF DIFFERENCE IN THE SEVERAL KINDS, CHIEFLY IN THE PODS AND SEEDSOME VARIETIES CONSTANT, SOME HIGHLY VARIABLEDO NOT INTERCROSS.BEANS.POTATOES: NUMEROUS VARIETIES OFDIFFERING LITTLE, EXCEPT IN THE TUBERSCHARACTERS INHERITED ... Page 305

CHAPTER X.

PLANTS continued—FRUITS—ORNAMENTAL TREES—FLOWERS.

FRUITS.GRAPESVARY IN ODD AND TRIFLING PARTICULARS.MULBERRY.THE ORANGE GROUPSINGULAR RESULTS FROM CROSSING.PEACH AND NECTARINEBUD-VARIATIONANALOGOUS VARIATIONRELATION TO THE ALMOND.APRICOT.PLUMSVARIATION IN THEIR STONES.CHERRIESSINGULAR VARIETIES OF.APPLE.PEAR.STRAWBERRYINTERBLENDING OF THE ORIGINAL FORMS.GOOSEBERRYSTEADY INCREASE IN SIZE OF THE FRUITVARIETIES OF.WALNUT.NUT.CUCURBITACEOUS PLANTSWONDERFUL VARIATION OF.

ORNAMENTAL TREESTHEIR VARIATION IN DEGREE AND KINDASH-TREESCOTCH-FIRHAWTHORN.

FLOWERSMULTIPLE ORIGIN OF MANY KINDSVARIATION IN CONSTITUTIONAL PECULIARITIESKIND OF VARIATION.ROSESSEVERAL SPECIES CULTIVATED.PANSY.DAHLIA.HYACINTH, HISTORY AND VARIATION OF ... Page 332

CHAPTER XI.

ON BUD-VARIATION, AND ON CERTAIN ANOMALOUS MODES OF REPRODUCTION AND VARIATION.

BUD-VARIATIONS IN THE PEACH, PLUM, CHERRY, VINE, GOOSEBERRY, CURRANT, AND BANANA, AS SHOWN BY THE MODIFIED FRUITIN FLOWERS: CAMELLIAS, AZALEAS, CHRYSANTHEMUMS, ROSES, ETC.ON THE RUNNING OF THE COLOUR IN CARNATIONSBUD-VARIATIONS IN LEAVESVARIATIONS BY SUCKERS, TUBERS, AND BULBSON THE BREAKING OF TULIPSBUD-VARIATIONS GRADUATE INTO CHANGES CONSEQUENT ON CHANGED CONDITIONS OF LIFECYTISUS ADAMI, ITS ORIGIN AND TRANSFORMATIONON THE UNION OF TWO DIFFERENT EMBRYOS IN ONE SEEDTHE TRIFACIAL ORANGEON REVERSION BY BUDS IN HYBRIDS AND MONGRELSON THE PRODUCTION OF MODIFIED BUDS BY THE GRAFTING OF ONE VARIETY OR SPECIES ON ANOTHERON THE DIRECT OR IMMEDIATE ACTION OF FOREIGN POLLEN ON THE MOTHER-PLANTON THE EFFECTS IN FEMALE ANIMALS OF A FIRST IMPREGNATION ON THE SUBSEQUENT OFFSPRINGCONCLUSION AND SUMMARY ... Page 373

LIST OF ILLUSTRATIONS.

1. Dun Devonshire Pony, with shoulder, spinal, and leg stripes ... PAGE 56
2. Head of Japan or Masked Pig ... 69
3. Head of Wild Boar, and of "Golden Days," a pig of the Yorkshire large breed ... 72
4. Old Irish Pig, with jaw-appendages ... 75
5. Half-lop Rabbit ... 108
6. Skull of Wild Rabbit ... 117
7. Skull of large Lop-eared Rabbit ... 117
8. Part of Zygomatic Arch, showing the projecting end of the malar-bone, and the auditory meatus, of Rabbits ... 118
9. Posterior end of Skull, showing the inter-parietal bone, of Rabbits ... 118
10. Occipital Foramen of Rabbits ... 118
11. Skull of Half-lop Rabbit ... 119
12. Atlas Vertebræ of Rabbits ... 121
13. Third Cervical Vertebræ of Rabbits ... 121
14. Dorsal Vertebræ, from sixth to tenth inclusive, of Rabbits ... 122
15. Terminal Bone of Sternum of Rabbits ... 123
16. Acromion of Scapula of Rabbits ... 123
17. The Rock-Pigeon, or Columbia Livia ... 135
18. English Pouter ... 137
19. English Carrier ... 140
20. English Barb ... 145
21. English Fantail ... 147
22. African Owl ... 149
23. Short-faced English Tumbler ... 152
24. Skulls of Pigeons, viewed laterally ... 163
25. Lower Jaws of Pigeons, seen from above ... 164
26. Skull of Runt, seen from above ... 165
27. Lateral view of Jaws of Pigeons ... 165
28. Scapulæ of Pigeons ... 167
29. Furculæ of Pigeons ... 167
30. Spanish Fowl ... 226
31. Hamburgh Fowl ... 228
32. Polish Fowl ... 229
33. Occipital Foramen of the Skulls of Fowls ... 261
34. Skulls of Fowls, viewed from above, a little obliquely ... 262
35. Longitudinal sections of Skulls of Fowls, viewed laterally ... 263
36. Skull of Horned Fowl, viewed from above, a little obliquely ... 265
37. Sixth Cervical Vertebræ of Fowls, viewed laterally ... 267
38. Extremity of the Furcula of Fowls, viewed laterally ... 268
39. Skulls of Ducks, viewed laterally, reduced to two-thirds of the natural size ... 282
40. Cervical Vertebræ of Ducks, of natural size ... 283
41. Pods of the Common Pea ... 328
42. Peach and Almond Stones, of natural size, viewed edgeways ... 337
43. Plum Stones, of natural size, viewed laterally ... 345






THE VARIATION OF ANIMALS AND PLANTS UNDER DOMESTICATION.

By Charles Darwin

IN TWO VOLUMES.-Vol. II.

WITH ILLUSTRATIONS.

INDEX

CONTENTS OF VOLUME II.

CHAPTER XII.

INHERITANCE.

WONDERFUL NATURE OF INHERITANCEPEDIGREES OF OUR DOMESTICATED ANIMALSINHERITANCE NOT DUE TO CHANCETRIFLING CHARACTERS INHERITEDDISEASES INHERITEDPECULIARITIES IN THE EYE INHERITEDDISEASES IN THE HORSELONGEVITY AND VIGOURASYMMETRICAL DEVIATIONS OF STRUCTUREPOLYDACTYLISM AND REGROWTH OF SUPERNUMERARY DIGITS AFTER AMPUTATIONCASES OF SEVERAL CHILDREN SIMILARLY AFFECTED FROM NON-AFFECTED PARENTSWEAK AND FLUCTUATING INHERITANCE: IN WEEPING TREES, IN DWARFNESS, COLOUR OF FRUIT AND FLOWERS, COLOUR OF HORSESNON-INHERITANCE IN CERTAIN CASESINHERITANCE OF STRUCTURE AND HABITS OVERBORNE BY HOSTILE CONDITIONS OF LIFE, BY INCESSANTLY RECURRING VARIABILITY, AND BY REVERSIONCONCLUSION ... Page 1

CHAPTER XIII.

INHERITANCE continued—REVERSION OR ATAVISM.

DIFFERENT FORMS OF REVERSIONIN PURE OR UNCROSSED BREEDS, AS IN PIGEONS, FOWLS, HORNLESS CATTLE AND SHEEP, IN CULTIVATED PLANTSREVERSION IN FERAL ANIMALS AND PLANTSREVERSION IN CROSSED VARIETIES AND SPECIESREVERSION THROUGH BUD-PROPAGATION, AND BY SEGMENTS IN THE SAME FLOWER OR FRUITIN DIFFERENT PARTS OF THE BODY IN THE SAME ANIMALTHE ACT OF CROSSING A DIRECT CAUSE OF REVERSION, VARIOUS CASES OF, WITH INSTINCTSOTHER PROXIMATE CAUSES OF REVERSIONLATENT CHARACTERSSECONDARY SEXUAL CHARACTERSUNEQUAL DEVELOPMENT OF THE TWO SIDES OF THE BODYAPPEARANCE WITH ADVANCING AGE OF CHARACTERS DERIVED FROM A CROSSTHE GERM WITH ALL ITS LATENT CHARACTERS A WONDERFUL OBJECTMONSTROSITIESPELORIC FLOWERS DUE IN SOME CASES TO REVERSION ... Page 28

CHAPTER XIV.

INHERITANCE continued—FIXEDNESS OF CHARACTER—PREPOTENCY—SEXUAL LIMITATION—CORRESPONDENCE OF AGE.

FIXEDNESS OF CHARACTER APPARENTLY NOT DUE TO ANTIQUITY OF INHERITANCEPREPOTENCY OF TRANSMISSION IN INDIVIDUALS OF THE SAME FAMILY, IN CROSSED BREEDS AND SPECIES; OFTEN STRONGER IN ONE SEX THAN THE OTHER; SOMETIMES DUE TO THE SAME CHARACTER BEING PRESENT AND VISIBLE IN ONE BREED AND LATENT IN THE OTHERINHERITANCE AS LIMITED BY SEXNEWLY-ACQUIRED CHARACTERS IN OUR DOMESTICATED ANIMALS OFTEN TRANSMITTED BY ONE SEX ALONE, SOMETIMES LOST BY ONE SEX ALONEINHERITANCE AT CORRESPONDING PERIODS OF LIFETHE IMPORTANCE OF THE PRINCIPLE WITH RESPECT TO EMBRYOLOGY; AS EXHIBITED IN DOMESTICATED ANIMALS; AS EXHIBITED IN THE APPEARANCE AND DISAPPEARANCE OF INHERITED DISEASES; SOMETIMES SUPERVENING EARLIER IN THE CHILD THAN IN THE PARENTSUMMARY OF THE THREE PRECEDING CHAPTERS ... Page 62

CHAPTER XV.

ON CROSSING.

FREE INTERCROSSING OBLITERATES THE DIFFERENCES BETWEEN ALLIED BREEDSWHEN THE NUMBERS OF TWO COMMINGLING BREEDS ARE UNEQUAL, ONE ABSORBS THE OTHERTHE RATE OF ABSORPTION DETERMINED BY PREPOTENCY OF TRANSMISSION, BY THE CONDITIONS OF LIFE, AND BY NATURAL SELECTIONALL ORGANIC BEINGS OCCASIONALLY INTERCROSS; APPARENT EXCEPTIONSON CERTAIN CHARACTERS INCAPABLE OF FUSION; CHIEFLY OR EXCLUSIVELY THOSE WHICH HAVE SUDDENLY APPEARED IN THE INDIVIDUALON THE MODIFICATION OF OLD RACES, AND THE FORMATION OF NEW RACES, BY CROSSINGSOME CROSSED RACES HAVE BRED TRUE FROM THEIR FIRST PRODUCTIONON THE CROSSING OF DISTINCT SPECIES IN RELATION TO THE FORMATION OF DOMESTIC RACES ... Page 85

CHAPTER XVI.

CAUSES WHICH INTERFERE WITH THE FREE CROSSING OF VARIETIES—INFLUENCE OF DOMESTICATION ON FERTILITY.

DIFFICULTIES IN JUDGING OF THE FERTILITY OF VARIETIES WHEN CROSSEDVARIOUS CAUSES WHICH KEEP VARIETIES DISTINCT, AS THE PERIOD OF BREEDING AND SEXUAL PREFERENCEVARIETIES OF WHEAT SAID TO BE STERILE WHEN CROSSEDVARIETIES OF MAIZE, VERBASCUM, HOLLYHOCK, GOURDS, MELONS, AND TOBACCO, RENDERED IN SOME DEGREE MUTUALLY STERILEDOMESTICATION ELIMINATES THE TENDENCY TO STERILITY NATURAL TO SPECIES WHEN CROSSEDON THE INCREASED FERTILITY OF UNCROSSED ANIMALS AND PLANTS FROM DOMESTICATION AND CULTIVATION ... Page 100

CHAPTER XVII.

ON THE GOOD EFFECTS OF CROSSING, AND ON THE EVIL EFFECTS OF CLOSE INTERBREEDING.

DEFINITION OF CLOSE INTERBREEDINGAUGMENTATION OF MORBID TENDENCIESGENERAL EVIDENCE ON THE GOOD EFFECTS DERIVED FROM CROSSING, AND ON THE EVIL EFFECTS FROM CLOSE INTERBREEDINGCATTLE, CLOSELY INTERBRED; HALF-WILD CATTLE LONG KEPT IN THE SAME PARKSSHEEPFALLOW-DEERDOGSRABBITSPIGSMAN, ORIGIN OF HIS ABHORRENCE OF INCESTUOUS MARRIAGESFOWLSPIGEONSHIVE-BEESPLANTS, GENERAL CONSIDERATIONS ON THE BENEFITS DERIVED FROM CROSSINGMELONS, FRUIT-TREES, PEAS, CABBAGES, WHEAT, AND FOREST-TREESON THE INCREASED SIZE OF HYBRID PLANTS, NOT EXCLUSIVELY DUE TO THEIR STERILITYON CERTAIN PLANTS WHICH EITHER NORMALLY OR ABNORMALLY ARE SELF-IMPOTENT, BUT ARE FERTILE, BOTH ON THE MALE AND FEMALE SIDE, WHEN CROSSED WITH DISTINCT INDIVIDUALS EITHER OF THE SAME OR ANOTHER SPECIESCONCLUSION ... Page 114

CHAPTER XVIII.

ON THE ADVANTAGES AND DISADVANTAGES OF CHANGED CONDITIONS OF LIFE: STERILITY FROM VARIOUS CAUSES.

ON THE GOOD DERIVED FROM SLIGHT CHANGES IN THE CONDITIONS OF LIFESTERILITY FROM CHANGED CONDITIONS, IN ANIMALS, IN THEIR NATIVE COUNTRY AND IN MENAGERIESMAMMALS, BIRDS, AND INSECTSLOSS OF SECONDARY SEXUAL CHARACTERS AND OF INSTINCTSCAUSES OF STERILITYSTERILITY OF DOMESTICATED ANIMALS FROM CHANGED CONDITIONSSEXUAL INCOMPATIBILITY OF INDIVIDUAL ANIMALSSTERILITY OF PLANTS FROM CHANGED CONDITIONS OF LIFECONTABESCENCE OF THE ANTHERSMONSTROSITIES AS A CAUSE OF STERILITYDOUBLE FLOWERSSEEDLESS FRUITSTERILITY FROM THE EXCESSIVE DEVELOPMENT OF THE ORGANS OF VEGETATIONFROM LONG-CONTINUED PROPAGATION BY BUDSINCIPIENT STERILITY THE PRIMARY CAUSE OF DOUBLE FLOWERS AND SEEDLESS FRUIT ... Page 145

CHAPTER XIX.

SUMMARY OF THE FOUR LAST CHAPTERS, WITH REMARKS ON HYBRIDISM.

ON THE EFFECTS OF CROSSINGTHE INFLUENCE OF DOMESTICATION ON FERTILITYCLOSE INTERBREEDINGGOOD AND EVIL RESULTS FROM CHANGED CONDITIONS OF LIFEVARIETIES WHEN CROSSED NOT INVARIABLY FERTILEON THE DIFFERENCE IN FERTILITY BETWEEN CROSSED SPECIES AND VARIETIESCONCLUSIONS WITH RESPECT TO HYBRIDISMLIGHT THROWN ON HYBRIDISM BY THE ILLEGITIMATE PROGENY OF DIMORPHIC AND TRIMORPHIC PLANTSSTERILITY OF CROSSED SPECIES DUE TO DIFFERENCES CONFINED TO THE REPRODUCTIVE SYSTEMNOT ACCUMULATED THROUGH NATURAL SELECTIONREASONS WHY DOMESTIC VARIETIES ARE NOT MUTUALLY STERILETOO MUCH STRESS HAS BEEN LAID ON THE DIFFERENCE IN FERTILITY BETWEEN CROSSED SPECIES AND CROSSED VARIETIESCONCLUSION ... Page 173

CHAPTER XX.

SELECTION BY MAN.

SELECTION A DIFFICULT ARTMETHODICAL, UNCONSCIOUS, AND NATURAL SELECTIONRESULTS OF METHODICAL SELECTIONCARE TAKEN IN SELECTIONSELECTION WITH PLANTSSELECTION CARRIED ON BY THE ANCIENTS, AND BY SEMI-CIVILISED PEOPLEUNIMPORTANT CHARACTERS OFTEN ATTENDED TOUNCONSCIOUS SELECTIONAS CIRCUMSTANCES SLOWLY CHANGE, SO HAVE OUR DOMESTICATED ANIMALS CHANGED THROUGH THE ACTION OF UNCONSCIOUS SELECTIONINFLUENCE OF DIFFERENT BREEDERS ON THE SAME SUB-VARIETYPLANTS AS AFFECTED BY UNCONSCIOUS SELECTIONEFFECTS OF SELECTION AS SHOWN BY THE GREAT AMOUNT OF DIFFERENCE IN THE PARTS MOST VALUED BY MAN ... Page 192

CHAPTER XXI.

SELECTION—continued.

NATURAL SELECTION AS AFFECTING DOMESTIC PRODUCTIONSCHARACTERS WHICH APPEAR OF TRIFLING VALUE OFTEN OF REAL IMPORTANCECIRCUMSTANCES FAVOURABLE TO SELECTION BY MANFACILITY IN PREVENTING CROSSES, AND THE NATURE OF THE CONDITIONSCLOSE ATTENTION AND PERSEVERANCE INDISPENSABLETHE PRODUCTION OF A LARGE NUMBER OF INDIVIDUALS ESPECIALLY FAVOURABLEWHEN NO SELECTION IS APPLIED, DISTINCT RACES ARE NOT FORMEDHIGHLY-BRED ANIMALS LIABLE TO DEGENERATIONTENDENCY IN MAN TO CARRY THE SELECTION OF EACH CHARACTER TO AN EXTREME POINT, LEADING TO DIVERGENCE OF CHARACTER, RARELY TO CONVERGENCECHARACTERS CONTINUING TO VARY IN THE SAME DIRECTION IN WHICH THEY HAVE ALREADY VARIEDDIVERGENCE OF CHARACTER, WITH THE EXTINCTION OF INTERMEDIATE VARIETIES, LEADS TO DISTINCTNESS IN OUR DOMESTIC RACESLIMIT TO THE POWER OF SELECTIONLAPSE OF TIME IMPORTANTMANNER IN WHICH DOMESTIC RACES HAVE ORIGINATEDSUMMARY ... Page 224

CHAPTER XXII.

CAUSES OF VARIABILITY.

VARIABILITY DOES NOT NECESSARILY ACCOMPANY REPRODUCTIONCAUSES ASSIGNED BY VARIOUS AUTHORSINDIVIDUAL DIFFERENCESVARIABILITY OF EVERY KIND DUE TO CHANGED CONDITIONS OF LIFEON THE NATURE OF SUCH CHANGESCLIMATE, FOOD, EXCESS OF NUTRIMENTSLIGHT CHANGES SUFFICIENTEFFECTS OF GRAFTING ON THE VARIABILITY OF SEEDLING-TREESDOMESTIC PRODUCTIONS BECOME HABITUATED TO CHANGED CONDITIONSON THE ACCUMULATIVE ACTION OF CHANGED CONDITIONSCLOSE INTERBREEDING AND THE IMAGINATION OF THE MOTHER SUPPOSED TO CAUSE VARIABILITYCROSSING AS A CAUSE OF THE APPEARANCE OF NEW CHARACTERSVARIABILITY FROM THE COMMINGLING OF CHARACTERS AND FROM REVERSIONON THE MANNER AND PERIOD OF ACTION OF THE CAUSES WHICH EITHER DIRECTLY, OR INDIRECTLY THROUGH THE REPRODUCTIVE SYSTEM, INDUCE VARIABILITY ... Page 250

CHAPTER XXIII.

DIRECT AND DEFINITE ACTION OF THE EXTERNAL CONDITIONS OF LIFE.

SLIGHT MODIFICATIONS IN PLANTS FROM THE DEFINITE ACTION OF CHANGED CONDITIONS, IN SIZE, COLOUR, CHEMICAL PROPERTIES, AND IN THE STATE OF THE TISSUESLOCAL DISEASESCONSPICUOUS MODIFICATIONS FROM CHANGED CLIMATE OR FOOD, ETC.PLUMAGE OF BIRDS AFFECTED BY PECULIAR NUTRIMENT, AND BY THE INOCULATION OF POISONLAND-SHELLSMODIFICATIONS OF ORGANIC BEINGS IN A STATE OF NATURE THROUGH THE DEFINITE ACTION OF EXTERNAL CONDITIONSCOMPARISON OF AMERICAN AND EUROPEAN TREESGALLSEFFECTS OF PARASITIC FUNGICONSIDERATIONS OPPOSED TO THE BELIEF IN THE POTENT INFLUENCE OF CHANGED EXTERNAL CONDITIONSPARALLEL SERIES OF VARIETIESAMOUNT OF VARIATION DOES NOT CORRESPOND WITH THE DEGREE OF CHANGE IN THE CONDITIONSBUD-VARIATIONMONSTROSITIES PRODUCED BY UNNATURAL TREATMENTSUMMARY ... Page 271

CHAPTER XXIV.

LAWS OF VARIATION—USE AND DISUSE, ETC.

NISUS FORMATIVUS, OR THE CO-ORDINATING POWER OF THE ORGANISATIONON THE EFFECTS OF THE INCREASED USE AND DISUSE OF ORGANSCHANGED HABITS OF LIFEACCLIMATISATION WITH ANIMALS AND PLANTSVARIOUS METHODS BY WHICH THIS CAN BE EFFECTEDARRESTS OF DEVELOPMENTRUDIMENTARY ORGANS ... Page 293

CHAPTER XXV.

LAWS OF VARIATION, continued—CORRELATED VARIABILITY.

EXPLANATION OF TERMCORRELATION AS CONNECTED WITH DEVELOPMENTMODIFICATIONS CORRELATED WITH THE INCREASED OR DECREASED SIZE OF PARTSCORRELATED VARIATION OF HOMOLOGOUS PARTSFEATHERED FEET IN BIRDS ASSUMING THE STRUCTURE OF THE WINGSCORRELATION BETWEEN THE HEAD AND THE EXTREMITIESBETWEEN THE SKIN AND DERMAL APPENDAGESBETWEEN THE ORGANS OF SIGHT AND HEARINGCORRELATED MODIFICATIONS IN THE ORGANS OF PLANTSCORRELATED MONSTROSITIESCORRELATION BETWEEN THE SKULL AND EARSSKULL AND CREST OF FEATHERSSKULL AND HORNSCORRELATION OF GROWTH COMPLICATED BY THE ACCUMULATED EFFECTS OF NATURAL SELECTIONCOLOUR AS CORRELATED WITH CONSTITUTIONAL PECULIARITIES ... Page 319

CHAPTER XXVI.

LAWS OF VARIATION, continued—SUMMARY.

ON THE AFFINITY AND COHESION OF HOMOLOGOUS PARTSON THE VARIABILITY OF MULTIPLE AND HOMOLOGOUS PARTSCOMPENSATION OF GROWTHMECHANICAL PRESSURERELATIVE POSITION OF FLOWERS WITH RESPECT TO THE AXIS OF THE PLANT, AND OF SEEDS IN THE CAPSULE, AS INDUCING VARIATIONANALOGOUS OR PARALLEL VARIETIESSUMMARY OF THE THREE LAST CHAPTERS ... Page 339

CHAPTER XXVII.

PROVISIONAL HYPOTHESIS OF PANGENESIS.

PRELIMINARY REMARKSFIRST PART:THE FACTS TO BE CONNECTED UNDER A SINGLE POINT OF VIEW, NAMELY, THE VARIOUS KINDS OF REPRODUCTIONTHE DIRECT ACTION OF THE MALE ELEMENT ON THE FEMALEDEVELOPMENTTHE FUNCTIONAL INDEPENDENCE OF THE ELEMENTS OR UNITS OF THE BODYVARIABILITYINHERITANCEREVERSION.

SECOND PART:STATEMENT OF THE HYPOTHESISHOW FAR THE NECESSARY ASSUMPTIONS ARE IMPROBABLEEXPLANATION BY AID OF THE HYPOTHESIS OF THE SEVERAL CLASSES OF FACTS SPECIFIED IN THE FIRST PARTCONCLUSION ... Page 357

CHAPTER XXVIII.

CONCLUDING REMARKS.

DOMESTICATIONNATURE AND CAUSES OF VARIABILITYSELECTIONDIVERGENCE AND DISTINCTNESS OF CHARACTEREXTINCTION OF RACESCIRCUMSTANCES FAVOURABLE TO SELECTION BY MANANTIQUITY OF CERTAIN RACESTHE QUESTION WHETHER EACH PARTICULAR VARIATION HAS BEEN SPECIALLY PREORDAINED ... Page 405

Index ... Page 433