Ray-flowers fertile.  156

156. Pappus of two or more rows of bristles, the outer of which are shorter.  157

Pappus of subequal bristles sometimes intermingled with a few shorter ones.  159

157. Pappus-bristles in 3 or more rows. Heads without ray-flowers.—Species
10. Tropical and South Africa and Egypt.  Iphiona Cass.

Pappus-bristles in 2 rows.  158

158. Pappus of 5 inner and 10 outer bristles. Heads without ray-flowers.
Shrubs.—Species 1. South Africa.  Anisothrix O. Hoffm.

Pappus of 10 inner and 10 outer bristles. Heads with ray-flowers. Undershrubs.—Species
1. South Africa.  Minurothamnus DC.

159. Heads few-flowered, without ray-flowers, arranged in panicles or corymbs.
Involucre of few bracts.—Species 2. Egypt. (Under Iphiona Cass.)  Varthemia DC.

Heads many-flowered. Involucre of many bracts.  160

160. Involucral bracts leathery, the outer sticky at the tip. Heads with ray-flowers, solitary. Pappus-bristles in one row. Glandular-hairy shrubs.—Species
1. South Africa.  Homochaete Benth.

Involucral bracts herbaceous or the inner scarious.—Species 30. Some of them yield vermin-poison or are used in medicine. (Including Bojeria
DC., Pentatrichia Klatt, Schizogyne Cass., and Vicoa Cass.)  Inula L.

161. Pappus-scales united below.  162

Pappus-scales free.  163

162. Pappus-bristles about 5. Female marginal flowers wanting. Heads in corymbs. Shrubs.—Species 2. Canary Islands.  Allagopappus Cass.

Pappus-bristles 7 or more. Female marginal flowers present. Heads solitary at the ends of the branches. Herbs.—Species 30. Some of them yield vermin-poison or are used in medicine. (Including Francoeuria
Cass.)  Pulicaria Gaertn.

163. Pappus-scales rather broad, fringed. Fruits 10-ribbed. Heads without ray-flowers. Herbs. (See 69.)  Pegolettia Cass.

Pappus-scales very narrow.  164

164. Fruits constricted into a short neck, 10-ribbed. Heads without ray-flowers.
Shrubs. Leaves pinnately divided.—Species 3. North
Africa. (Under Grantia Boiss.)  Perralderia Coss.

Fruits not constricted above. Heads with ray-flowers.  165

165. Fruits 4-5-ribbed. Pappus-bristles 5-10. Shrubs.—Species 1. Canary
Islands.  Viraea Webb

Fruits many-ribbed. Pappus-bristles numerous. Undershrubs.—Species
1. North-west Africa.  Jasonia Cass.

166. (149.) Female or neuter marginal flowers with a strap-shaped corolla.  167

Female or neuter marginal flowers with a thread-shaped corolla or wanting.
[Tribe INULEAE, subtribes GNAPHALINAE and RELHANINAE.]  176

167. Leaves grooved or rolled inwards on the upper side, small. Heath-like plants. [Tribe INULEAE, subtribe RELHANINAE.]  168

Leaves flat or rolled back from the margins. Not heath-like plants.  172

168. Heads one-flowered, some hermaphrodite, the others female, or 2-flowered with a hermaphrodite and a female or neuter flower. Pappus of feathery bristles united at the base. Shrubs.—Species 8. South Africa.  Disparago Gaertn.

Heads many-flowered.  169

169. Pappus wanting. Shrubs.—Species 2. South Africa.  Anaglypha DC.

Pappus present.  170

170. Pappus of numerous scales sometimes united below.—Species 13. South
Africa.  Nestlera Spreng.

Pappus of bristles.  171

171. Pappus-bristles feathery. Shrubs.—Species 5. South Africa.  Amphiglossa DC.

Pappus-bristles simple. Herbs. Flowers red.—Species 1. South
Africa.  Bryomorphe Harv.

172. Pappus wanting. Inner fruits compressed. Heads solitary. Herbs or undershrubs. (See 124.)  Dimorphotheca Moench

Pappus present. [Tribe INULEAE, subtribe ATHRIXINAE.]  173

173. Pappus of the outer fruits consisting of scales, of the inner of bristles or of scales and bristles. Fruits glabrous. Herbs or undershrubs.
(See 101.)  Leyssera L.

Pappus consisting of bristles.  174

174. Involucral bracts narrow, acuminate. Herbs or undershrubs.—Species
15. Southern and tropical Africa.  Athrixia Ker

Involucral bracts blunt, scarious at the apex. Pappus-bristles in two rows. Shrubs.  175

175. Fruits hairy. Pappus-bristles thick and stiff. Involucre hemispherical.—Species
3. South Africa.  Heterolepis Cass.

Fruits glabrous, but with a hairy swelling at the base. Pappus-bristles thin. Involucre narrow-campanulate.—Species 3. Central and
South Africa.  Antithrixia DC.

176. (166.) Hermaphrodite flowers fewer than the female.  177

Hermaphrodite flowers as many as or more than the female, or all flowers hermaphrodite.  184

177. Fruits without a pappus. Heads small, in dense cymes. Tall herbs.—Species
1. East Africa.  Chiliocephalum Benth.

Fruits, at least the inner, crowned by a pappus.  178

178. Inner fruits with a pappus of feathery bristles, outer without a pappus.
Heads small, in glomerules. Low herbs. (See 132.)  Amphidoxa DC.

Inner and outer fruits crowned by a pappus.  179

179. Pappus of the inner fruits of bristles and united scales, that of the outer only of scales united below. Heads solitary. Herbs.—Species 1.
East Africa.  Artemisiopsis S. Moore

Pappus of all fruits formed of bristles.  180

180. Pappus-bristles feathery. Heads in glomerules. Herbs.—Species
2. South and North Africa.  Lasiopogon Cass.

Pappus-bristles not feathery.  181

181. Fruits with a long beak. Herbs. Leaves radical or opposite. Heads solitary.—Species 1. Island of Tristan da Cunha.  Chevreulia Cass.

Fruits without a beak.  182

182. Heads 4-8-flowered, cylindrical, in dense cymes arranged in panicles.
Undershrubs. (See 110.)  Achyrocline Less.

Heads many-flowered, hemispherical ovoid or campanulate.  183

183. Anthers very shortly tailed. Shrubs or undershrubs. Heads solitary or in long-stalked glomerules.—Species 15. North and Central Africa.  Phagnalon Cass.

Anthers distinctly tailed. Herbs.—Species 50. “Cudweed.”  Gnaphalium L.

184. (176.) Heads 1-flowered.  185

Heads 2- or more-flowered.  188

185. Flowers partly hermaphrodite, partly female. Pappus of feathery bristles. Heads in glomerules. Shrubs. (See 168.)  Disparago Gaertn.

Flowers all hermaphrodite.  186

186. Pappus wanting. Heads in glomerules. Shrubs.—Species 2. South
Africa.  Perotriche Cass.

Pappus formed of bristles.  187

187. Pappus-bristles feathery. Inner involucral bracts scarious. Heath-like shrubs.—Species 35. Southern and tropical Africa.  Stoebe L.

Pappus-bristles feathery only at the apex or not feathery. Involucral bracts scarious, coloured. Heads in panicled cymes. Herbs.—Species
12. Madagascar, Mauritius, and South Africa to Damaraland.
Some are used medicinally.  Stenocline DC.

188. Pappus wanting. Heads 2-3-flowered, in leafy corymbs.  189

Pappus formed of bristles.  190

189. Leaves small. Undershrubs.—Species 1. Madagascar.  Syncephalum DC.

Leaves rather large. Shrubs. Involucre woolly at the base.—Species 1.
Madagascar.  Astephanocarpa Bak.

190. Pappus-bristles feathery from the base.  191

Pappus-bristles feathery at the tip only or not feathery.  192

191. Heads 2-10-flowered. Heath-like shrubs.—Species 3. South Africa.  Pterothrix DC.

Heads many-flowered. Involucral bracts scarious, coloured. Not heath-like plants. (See 84.)  Helipterum DC.

192. Pappus-bristles in 1 row.  193

Pappus-bristles in 2 or more rows.  197

193. Pappus-bristles with bladdery inflated cells at the tip. Heads small, in glomerules. Prostrate herbs.—Species 1. South Africa.  Eriosphaera Less.

Pappus-bristles without bladdery inflated cells.  194

194. Habit heath-like. Shrubs. Leaves small, grooved or rolled inwards on the upper face. Flowers all hermaphrodite.  195

Habit not heath-like.  196

195. Pappus-bristles feathery at the tip, united at the base and surrounded by
a ring- or cup-shaped rim. Heads few-flowered.—Species 7. South
Africa. Some are used medicinally.  Elytropappus Cass.

Pappus-bristles simple or thickened at the tip. Inner involucral bracts coloured above.—Species 25. South Africa.  Metalasia R. Br.

196. Fruits large, with 8-10 prominent ribs, glabrous or short-haired. Heads
2-6-flowered, in panicled cymes. Herbs. (See 187.)  Stenocline DC.

Fruits small, not prominently 8-10-ribbed. (See 84.)  Helichrysum Gaertn.

197. Pappus-bristles in 2 rows. Heath-like shrubs.  198

Pappus-bristles in 3 or more rows. Not heath-like herbs or undershrubs.  199

198. Heads few-flowered. Involucre oblong, of oblong bracts. Receptacle glabrous. Fruits ribbed. Leaves oblong.—Species 1. Madagascar.  Cullumiopsis Drake

Heads many-flowered. Involucre top-shaped, of linear bracts. Receptacle bristly at the margin. Fruits angular, hairy. Leaves linear.
(See 83.)  Lachnospermum Willd.

199. Fruits beaked, hairy. Heads few-flowered. Inner involucral bracts yellow.—Species 1. South Africa.  Pachyrhynchus DC.

Fruits not beaked, glabrous. Heads in cymes.—Species 8. South
Africa.  Leontonyx Cass.

200. (67.) Hermaphrodite disc-flowers sterile, their style without a stigma and usually entire or shortly toothed.  201

Hermaphrodite flowers, at least some of them, fertile.  238

201. Anthers arrow-shaped, with acuminate halves. Corolla of the marginal flowers strap-shaped. Fruits glabrous, usually large. Pappus wanting, rarely cup-shaped. Receptacle glabrous, rarely bristly. [Tribe CALENDULEAE.]  202

Anthers entire or shortly auricled at the base, rarely (Adelostigma) distinctly arrow-shaped, but then corolla of the marginal flowers thread-shaped, fruits hairy, and pappus bristly.  207

202. Outer fruits of several kinds. Heads solitary, yellow-flowered. Herbs or undershrubs.  203

Outer fruits all alike.  204

203. Fruits curved. Heads middle-sized. (See 122.)  Calendula L.

Fruits straight. Heads small. (See 122.)  Oligocarpus Less.

204. Fruits with 3 wings and a cup-shaped apical appendage. Involucral bracts in one row. (See 123.)  Tripteris Less.

Fruits without distinct wings or other appendages.  205

205. Fruits very hard, smooth or indistinctly ribbed. Involucral bracts in two
or more rows. Ray-flowers yellow. (See 124.)  Osteospermum L.

Fruits not very hard, 3-angled, usually tubercled. Herbs or undershrubs.
Heads solitary.  206

206. Involucral bracts in one row or in two indistinct rows. (See 124.)  Dimorphotheca Moench

Involucral bracts in several rows. Ray-flowers blue or white. Leaves pinnately divided.—Species 6. South Africa. Some are used medicinally.  Garuleum Cass.

207. (201.) Receptacle covered with chaffy scales.  208

Receptacle glabrous or hairy, without scales between the flowers.  213

208. Heads with the inner flowers male (apparently hermaphrodite), the outer female and provided with a more or less strap-shaped, rarely a thread-shaped corolla.  209

Heads unisexual, some with all the flowers male (apparently hermaphrodite), the others with all the flowers female and provided with a tubular corolla or without a corolla.  211

209. Involucral bracts in one row, united below when young. Outer fruits hairy, without a pappus, the inner with a pappus of numerous one-ranked bearded bristles. Heads solitary. Shrubs.—Species 1.
South Africa. (Under Eriocephalus L.)  Lasiocoma Bolus

Involucral bracts in two or more rows. Pappus of 2-3 bristles or wanting.  210

210. Involucral bracts all alike. Corolla of the female flowers 2-toothed.
Pappus of the outer fruits of 2-3 bristles. Heads in panicles.—Species
1. Naturalized in tropical and South Africa. Used medicinally.  Parthenium L.

Involucral bracts of two kinds, the inner united below. Corolla of the female flowers entire or 3-toothed. Pappus none. Heads in racemes or umbels, or solitary.—Species 20. South Africa to Damaraland.
Some are used medicinally.  Eriocephalus L.

211. Heads dioecious, many-flowered. Involucral bracts in 3 rows. Corolla
4-toothed, yellow. Anthers cohering, auricled at the base. Style cleft.
Fruits hairy. Pappus of 2-3 bristles. Trees. Leaves opposite.
Heads in leafy panicles.—Species 2. Island of St. Helena.  Petrobium R. Br.

Heads monoecious, the male many-flowered with a 5-toothed corolla, the female 1-2-flowered with an involucre of partly united bracts, and without a corolla. Anthers free or slightly cohering, entire at the base.
Style of the male flowers undivided. Pappus none. Herbs or undershrubs.
[Tribe HELIANTHEAE, subtribe AMBROSINAE.]  212

212. Involucral bracts of the male heads free. Filaments united. Female heads
2-flowered. Heads solitary or in glomerules in the axils of the leaves.—Species
4. North and Central Africa and Mascarene Islands; naturalized in South Africa. They yield dyes and medicaments and are noxious
to pasturing cattle.  Xanthium L.

Involucral bracts of the male heads united below. Filaments free or nearly so. Female heads 1-flowered. Male heads in spikes or racemes.—Species
2. Northern and tropical Africa; naturalized in South Africa.
Used medicinally.  Ambrosia L.

213. (207.) Pappus of the outer fruits ring-, crown-, or ear-shaped, sometimes produced into two small points, or wanting.  214

Pappus of the outer fruits consisting of bristles.  223

214. Pappus of the inner fruits of bristles, of the outer of 1-2 small points or wanting. Involucral bracts in few rows. Receptacle pitted. Corolla of the female flowers strap-shaped. Style cleft, with lanceolate appendages.
Fruits flattened. Shrubs or undershrubs. Leaves entire.—Species
8. South Africa. (Heteractis DC.)  Gymnostephium Less.

Pappus of all fruits alike or wanting.  215

215. Outer flowers with a strap-shaped corolla.  216

Outer flowers with a tubular corolla or without a corolla.  220

216. Corolla of the outer flowers very shortly strap-shaped, of the inner 4-toothed, yellow in all flowers. Involucral bracts in two rows.—Species
40. Some of them are used as ornamental or medicinal plants. (Including
Brocchia Vis. and Cenia Juss.)  Cotula L.

Corolla of the outer flowers distinctly strap-shaped. Involucral bracts rarely in two rows, and then corolla of the inner flowers 5-toothed.  217

217. Involucral bracts in one row and united at the base. Corolla-limb of the hermaphrodite (male) central flowers bell-shaped, 5-cleft. Style-branches of the same ending in a blunt appendage. Fruits flattened, without ribs. Branching herbs. Leaves pinnately divided. Heads solitary.—Species 4. South Africa.  Steirodiscus Less.

Involucral bracts in two or more rows.  218

218. Involucral bracts in two rows. Corolla of the marginal flowers red, of the central yellow, the latter 5-toothed. Style-branches of the hermaphrodite
(male) flowers pointed. Fruits beaked, without ribs. Herbs.
Leaves undivided, radical. Heads solitary.—Species 1. Island of
Tristan da Cunha.  Lagenophora Cass.

Involucral bracts in 3 or more rows. Corolla of the marginal flowers white, yellow, or blue, more rarely red, but then style-branches of the hermaphrodite (male) flowers truncate and fruits ribbed.  219

219. Style of the hermaphrodite (male) flowers with pointed branches or undivided. Corolla of the marginal flowers blue, rarely white. Fruits wrinkled or smooth. Branching herbs or undershrubs. Leaves pinnately divided. (See 206.)  Garuleum Cass.

Style of the hermaphrodite (male) flowers with truncate branches. Corolla of the marginal flowers white, yellow, or red. Fruits 5-10-ribbed.—Species
50. North Africa, Abyssinia, Madagascar, and South Africa.
Some species yield condiments, medicaments, or insect-poison, or serve
as ornamental plants. (Including Argyranthemum Webb, Ismelia Cass.,
Leucanthemum DC., Monoptera Schultz, Myconia Neck., Pinardia Cass.,
Plagius L’Hér., Preauxia Schultz, Prolongoa Boiss., Pyrethrum Gaertn., and Tanacetum L.)  Chrysanthemum L.

220. Corolla of the hermaphrodite (male) flowers 2-4-toothed, of the female
2-3-toothed, entire, or wanting. Involucral bracts in two rows.
Herbs. Flowers yellow.  221

Corolla of the hermaphrodite (male) flowers 5-toothed, of the female 2-4-toothed.  222

221. Heads sessile between the leaves. Female flowers in several rows, without
a corolla. Outer fruits winged, with a persistent style. Leaves divided.—Species
1. Naturalized in the Island of Madeira.  Soliva Ruiz & Pav.

Heads stalked, at the ends of the branches. (See 216.)  Cotula L.

222. Heads in corymbs. Involucral bracts in two rows. Fruits with marginal ribs or wings. Herbs or undershrubs.—Species 4. South Africa.
Used medicinally.  Hippia L.

Heads not in corymbs. Involucral bracts in several rows. Fruits without strong ribs.—Species 20. Some of them (especially A. Absinthium
L., wormwood) are used as ornamental or medicinal plants, as pot-herbs, or for preparing liquors and vinegar.  Artemisia L.

223. (213.) Pappus of 1 or 2 rows of bristles.  224

Pappus, at least that of the outer fruits, of 3 or more rows of bristles.  233

224. Pappus-bristles feathery.  225

Pappus-bristles not feathery.  226

225. Corolla of all flowers yellow. Small shrubs.—Species 1. South Africa.  Homochroma DC.

Corolla of the marginal flowers white or red, of the central ones yellow.
Herbs or undershrubs.—Species 13. South Africa. Used as ornamental plants.  Mairia Nees

226. Involucral bracts in one row, sometimes surrounded by some much shorter ones. Herbs or undershrubs. Heads in corymbs or panicles.  227

Involucral bracts in two rows and all nearly of the same size, or in 3 or more rows.  228

227. Involucral bracts united at the base. Style of the hermaphrodite (male) flowers undivided. Inner fruits without a pappus.—Species 3. South
Africa.  Gymnodiscus Less.

Involucral bracts free. Style cleft.—Species 45. Southern and tropical
Africa.  Cineraria L.

228. Corolla of the female marginal flowers thread-shaped. Herbs. Heads solitary on the ends of the branches. Style of the central flowers entire or shortly toothed.  229

Corolla of the female marginal flowers, at least of the outer ones, strap-shaped, rarely (Psiadia) thread-shaped, but then shrubs, heads in
corymbs, and style of the central flowers 2-cleft.  230

229. Involucral bracts in two rows. Corolla of the hermaphrodite (male) flowers 4-toothed. Anthers entire at the base. Fruits compressed, glabrous. (See 216.)  Cotula L.

Involucral bracts in 3 or more rows. Corolla of the hermaphrodite
(male) flowers 5-toothed. Anthers arrow-shaped. Fruits nearly terete, hairy.—Species 2. Central Africa.  Adelostigma Steetz

230. Corolla of the female marginal flowers yellow, thread- or shortly strap-shaped.
Fruits scarcely compressed, 3-6-ribbed. Shrubs. Heads in corymbs.—Species 40. Tropics. Some are used as pot-herbs.  Psiadia Jacq.

Corolla of the female marginal flowers white, red or blue, strap-shaped.
Fruits compressed.  231

231. Stem woody, at least at the base. Marginal flowers in 1-2 rows. Pappus-bristles usually in one row.—Species 65. South and Central Africa.
Some are used as ornamental plants. (Including Asterosperma Less.,
Detris Adans., and Diplopappus DC., under Aster L.)  Felicia Cass.

Stem herbaceous.  232

232. Involucral bracts in 2 rows, membranous. Marginal flowers usually in several rows, with a linear corolla. Style-branches with a mostly short triangular appendage. Pappus-bristles usually in one row.—Species
13, one of them only naturalized. Several species are used medicinally.  Erigeron L.

Involucral bracts in 3 or more rows. Marginal flowers in 1-2 rows, with a usually oblong corolla. Style-branches with a lanceolate appendage.
Pappus-bristles in two rows.—Species 10. South and
North Africa. Some are used as ornamental or medicinal plants.
(Including Linosyris Cass.)  Aster L.

233. (223.) Involucral bracts in several rows. (See 232.)  Aster L.

Involucral bracts in one row, but sometimes surrounded by some much smaller ones.  234

234. Heads of two kinds, some with most of the flowers hermaphrodite (male), the others with most of them female. Corolla red or white; limb shortly strap-shaped in the female flowers. Herbs. Leaves radical, broad.
Heads in racemes or corymbs.—Species 1. North Africa. Used as an ornamental plant.  Petasites Gaertn.

Heads all alike.  235

235. Female marginal flowers in several rows. Corolla yellow; limb long strap-shaped in the female flowers. Herbs. Leaves radical, broad.
Heads solitary.—Species 1. North Africa. Used medicinally. “Coltsfoot.”  Tussilago L.

Female marginal flowers in one row.  236

236. Inner fruits without a pappus, outer with a pappus of interwoven hairs.
Style 2-cleft. Herbs. Leaves pinnately divided. Heads solitary.—Species
3. South Africa.  Ruckeria DC.

Inner and outer fruits with a pappus of bristles.  237

237. Style of the central flowers 2-cleft. Involucral bracts finally free.
Undershrubs.—Species 7. South and North Africa. Some are used as ornamental plants. (Othonnopsis Jaub. et Spach)  Hertia Less.

Style of the central flowers undivided. Involucral bracts more or less united.—Species 110. South Africa and southern Central Africa.
Some are used as ornamental plants. (Including Doria Less.)  Othonna L.

238. (200.) Style-branches of the hermaphrodite flowers rather long, semi-cylindrical, covered with stigmatic papillae on the inner surface, with hairs on the outer; hairs also clothing the upper part of the style below the point of division. Anthers more or less arrow-shaped.  239

Style-branches of the hermaphrodite flowers with marginal rows of stigmatic papillae, hairy only in their upper part, more rarely down to the point of division; no hairs on the upper part of the style below that point.  262

239. Heads with inner hermaphrodite and outer female flowers.  240

Heads with all the flowers hermaphrodite, rarely (Vernonia) subdioecious by incomplete development of one sex.  242

240. Female marginal flowers with a thread-shaped corolla. Style-branches blunt. Pappus of bristles. Herbs.—Species 17. Tropical and
South Africa and Egypt.  Laggera Schultz.

Female marginal flowers with a strap-shaped corolla. Flowers yellow.
Receptacle pitted.  241

241. Pappus of scales. Involucral bracts united at the base. Leaves prickly.
(See 37.)  Berkheya Ehrh.

Pappus of bristles. Involucral bracts free. Shrubs. Heads solitary.
(See 138.).  Eremothamnus O. Hoffm.

242. Flowers yellow.  243

Flowers white, red, or blue.  244

243. Pappus of scales. Involucral bracts united at the base. Leaves prickly.
(See 37.)  Berkheya Ehrh.

Pappus of bristles. Involucral bracts free.—Species 7. Central Africa.
(Including Autunesia O. Hoffm. and Newtonia O. Hoffm., under Vernonia
Schreb.)  Gongrothamnus Steetz

244. Receptacle with scales between the flowers. Leaves linear or lanceolate.  245

Receptacle glabrous, rarely ciliate, bristly, or with toothed borders to the pits, without scales between the flowers. Style-branches rather long and pointed. [Tribe VERNONIEAE.]  247

245. Heads solitary. Scales between the flowers membranous. Style-branches rather long and pointed. Pappus of unequally broad scales.—Species
1. West Africa (Congo).  Dewildemania O. Hoffm.

Heads in corymbs. Scales between the flowers with a coloured appendage.
Style-branches short, blunt or somewhat pointed. Pappus of scales
united into a toothed cup.  246

246. Outer involucral bracts longer than the inner. Corolla slightly irregular.
Leaves linear, sessile, glabrous.—Species 1. Southern West Africa
(Angola).  Omphalopappus O. Hoffm.

Outer involucral bracts shorter than the inner. Corolla regular. Leaves lanceolate, short-stalked, short-haired.—Species 1. Southern West
Africa (Angola).  Gossweilera S. Moore

247. Heads in dense glomerules or heads, few-flowered. Involucral bracts in several rows of two bracts each. Corolla 5-cleft. Fruits 10-ribbed.
Pappus of 1-2 rows of bristles or narrow scales. Herbs.—Species 5.
Tropics. Used medicinally.  Elephantopus L.

Heads not in dense glomerules or heads. [Subtribe VERNONINAE.]  248

248. Pappus wanting.  249

Pappus present.  251

249. Heads 1-4-flowered, in corymbs. Involucre oblong. Fruits with an indistinctly cup-shaped border at the top. Shrubs. (See 136.)  Apodocephala Bak.

Heads many-flowered. Involucre campanulate or hemispherical. Herbs.  250

250. Fruits truncate at the top, 4-5-ribbed.—Species 5. Tropical and
South Africa and Egypt.  Ethulia L.

Fruits rounded at the top.—Species 10. Central Africa.  Gutenbergia Schultz

251. Pappus ear- or cup-shaped, entire or nearly so. Herbs.  252

Pappus formed of scales or bristles.  253

252. Pappus auricle-like. Heads in cymes. Leaves linear.—Species 1.
East Africa.  Hoehnelia Schweinf.

Pappus cup-shaped. Heads solitary or in fascicles.—Species 1. Tropics.  Sparganophorus Vaill.

253. Pappus of one row of scales and sometimes some bristles within them.  254

Pappus only of bristles or of several rows of bristles with some small scales outside them.  257

254. Pappus-scales 5, long and narrow. Fruits 5-ribbed. Corolla white.
Involucral bracts in two rows. Heads in panicles. Shrubs.—Species 1.
West Africa (Congo).  Msuata O. Hoffm.

Pappus-scales short. Herbs.  255

255. Pappus-scales 5. Fruits 4-ribbed, glabrous. Corolla violet. Receptacle pitted; pits with toothed borders. Heads many-flowered, in glomerules.—Species
3. Central Africa.  Ageratina O. Hoffm.

Pappus-scales more than 5. Receptacle not pitted.  256

256. Heads one-flowered, in corymbs. Involucral bracts in two rows. Fruits cylindrical, hairy. Pappus-scales united below. Leaves linear, with parallel veins.—Species 10. South Africa.  Corymbium L.

Heads several-flowered. Involucral bracts in several rows. Fruits
3-5-angled, glandular. Pappus-scales free or nearly so.—Species
4. Central Africa.  Herderia Cass.

257. Pappus of caducous, usually one-ranked bristles.  258

Pappus of persistent, several-ranked bristles or of bristles and scales.  260

258. Pappus surrounded by a wavy ring. Fruits 5-ribbed. Heads 4-5-flowered, in corymbs. Shrubs.—Species 1. East Africa.  Volkensia O. Hoffm.

Pappus not surrounded by a wavy ring. Herbs or undershrubs.  259

259. Outer involucral bracts leaf-like, much larger than the inner. Fruits blunt, 8-10-ribbed.—Species 1. West Africa (Congo).  Centratherum Cass.

Outer involucral bracts, like the inner, scale-like. Fruits 4-5-angled.—Species
30. Central and South Africa. (Including Bothriocline Oliv. and Stephanolepis S. Moore)  Erlangea Schultz

260. Leaves unarmed.—Species 330. Tropical and South Africa. Some of the species yield wood or medicaments. (Including Bechium DC.,
Cyanopis Blume, and Decaneurum DC.) (Plate 150.)  Vernonia Schreb.

Leaves prickly.  261

261. Heads axillary. Stem branched, woody at the base. Fruits hairy.—Species
2. South Africa.  Hoplophyllum DC.

Heads terminal. Stem not branched, herbaceous throughout. Fruits many-ribbed.—Species 2. West Africa.  Aedesia O. Hoffm.

262. (238.) Style-branches of the hermaphrodite flowers rather long, semi-cylindrical, usually blunt; the lower part bearing scarcely projecting and not recurved marginal rows of stigmatic papillae, the upper part subequally clothed with hairs on both sides. Flowers all hermaphrodite, red, blue or whitish. [Tribe EUPATORIEAE.]  263

Style-branches of the hermaphrodite flowers rather flat, bearing conspicuous marginal rows of stigmatic papillae sometimes confluent in the middle, and above them either a crown of rather long hairs, with or without shorter ones, or an appendage clothed with dense hairs on the outer face, scantily haired or not hairy on the inner; more rarely style-branches almost uniformly clothed with hairs, but then rows of stigmatic papillae curved outwards or confluent in the middle, or flowers yellow or partly female.  266

263. Anthers without an apical appendage. Fruits angular, glandular, without prominent ribs. Pappus of 3-5 bristles. Herbs. Leaves opposite.
Heads in panicles.—Species 2. Tropical and South Africa. Used medicinally.  Adenostemma Forst.

Anthers with an apical appendage. Pappus of numerous bristles or of scales. [Subtribe AGERATINAE.]  264

264. Pappus of 5-10 scales. Herbs. Leaves opposite. Heads in panicles.—Species
1. Used as an ornamental or medicinal plant.  Ageratum L.

Pappus of numerous bristles.  265

265. Involucral bracts 4-5. Heads few-flowered, in panicles. Leaves opposite.—Species
10. Tropical and South Africa. Some are used medicinally.
(Willugbaeya Neck.)  Mikania Willd.

Involucral bracts 6 or more.—Species 6. Four species indigenous to
North and Central Africa, the other two naturalized in Madagascar and the Mascarenes. Some are used as ornamental or medicinal plants.  Eupatorium L.

266. (262.) Style-branches of the hermaphrodite flowers bearing in the lower part marginal, finally reflexed rows of stigmatic papillae, in the upper part on the outer, more rarely also the inner surface, short subequal hairs. [Tribe ASTEREAE.]  267

Style-branches of the hermaphrodite flowers stigmatose at the margins or on the inner surface, bearing in the upper part or at the top only a crown of rather long hairs sometimes joined above or below or in both directions by shorter ones.  301

267. Female or neuter marginal flowers with a bell-, tube-, thread-, or very shortly strap-shaped corolla not overtopping the involucre.  268

Female or neuter marginal flowers with a distinctly strap-shaped corolla overtopping the involucre, or wanting.  276

268. Pappus of scales or few short bristles, or wanting. Corolla of the marginal flowers bell-, tube-, or thread-shaped. Herbs. [Subtribe GRANGEINAE.]  269

Pappus of long bristles. Corolla of the marginal flowers thread- or strap-shaped. [Subtribe CONYZINAE.]  272

269. Pappus wanting, rarely some minute bristles on the central fruits. Fruits compressed.—Species 6. Tropical and South Africa. Some species yield condiments and medicaments.  Dichrocephala DC.

Pappus present.  270

270. Pappus formed of bristles. Fruits subterete.—Species 1. Central
Africa.  Microtrichia DC.

Pappus cup-shaped or consisting of scales.  271

271. Receptacle with scales between the flowers.—Species 1. Central Africa and Egypt.  Ceruana Forsk.

Receptacle without scales between the flowers.—Species 3. Tropics to
Egypt. Used medicinally.  Grangea Adans.

272. Fruits turgid. Involucral bracts in many rows, membranous. Heads in panicles. Shrubs.—Species 13. Tropical and South Africa. Some species are used as vegetables or salad.  Microglossa DC.

Fruits compressed.  273

273. Female flowers in one row. Pappus-bristles deciduous. Shrubs or undershrubs. Leaves linear. Heads solitary at the ends of the branches.—Species
6. South and North Africa. (Leptothamnus DC.)  Nolletia Cass.

Female flowers in two or more rows.  274

274. Corolla of the marginal flowers strap-shaped, but shorter than the style, or thread-shaped, yellowish or whitish.—Species 80. Some of them yield condiments, medicaments, or insect-poison. (Marsea Adans., including Webbia Schultz).  Conyza Less.

Corolla of the marginal flowers strap-shaped, longer than the style. Involucral bracts in two rows. Herbs or undershrubs.  275

275. Ray-flowers yellow. Heads in corymbs.—Species 35. Southern and tropical Africa.  Nidorella Cass.

Ray-flowers red or white. (See 232.)  Erigeron L.

276. (267.) Ray-flowers yellow, sometimes reddish when old, or wanting.
Pappus of bristles. [Subtribe SOLIDAGININAE.]  277

Ray-flowers white, blue, or red.  289

277. Heads with all the flowers hermaphrodite; the inner flowers sometimes sterile.  278

Heads with the inner flowers hermaphrodite and surrounded by one row of female or neuter marginal flowers.  286

278. Involucral bracts in one row or in two very unequal rows (the outer of much smaller bracts). Fruits 5-10-ribbed. Herbs.  279

Involucral bracts in two subequal rows or in 3 or more rows.  280

279. Hairy appendages of the style-branches short. Pappus of bristles.
Leaves radical.—Species 1. West Africa.  Psednotrichia Hiern

Hairy appendages of the style-branches long. Pappus of hairs. Leaves alternate.—Species 20. Tropical and South-west Africa. (Including
Crassocephalum Moench, under Senecio L.)  Gynura Cass.

280. Pappus-bristles in 1 row. Shrubs. Leaves linear. Heads in leafy corymbs.  281

Pappus-bristles in 2 or more rows.  282

281. Fruits turgid, 5-ribbed.—Species 1. South Africa (Orange River Colony).  Pentheriella O. Hoffm. & Muschler

Fruits compressed.—Species 15. South Africa. Some are used as ornamental plants.  Chrysocoma L.

282. Pappus-bristles in 2 very unequal rows, the outer of very short, sometimes scale-like bristles. Fruits compressed.  283

Pappus-bristles in 2 subequal rows or in 3 or more rows.  284

283. Outer pappus-bristles scale-like. Hairy appendages of the style-branches linear, obtuse. Herbs. Leaves opposite.—Species 2. Southern
West Africa. (Adenogonum Welw.)  Engleria O. Hoffm.

Outer pappus-bristles hair-like. Hairy appendages of the style-branches lanceolate. Shrubs.—Species 5. South Africa.  Fresenia DC.

284. Stem woody, shrubby.—Species 55. South Africa to Damaraland.  Pteronia L.

Stem herbaceous. Fruits compressed. Pappus-bristles in 2 or 3 rows.  285

285. Leaves decurrent. Flowers yellow. Heads in corymbs. Species 2.
South Africa. (Under Chrysocoma L.)  Heteromma Benth.

Leaves not decurrent. (See 232.)  Aster L.

286. (277.) Marginal rows of stigmatic papillae confluent at the rounded apex of the style-branches. Flower-heads large. Involucre broad, of many rows of bracts. Pappus-bristles in 2-3 rows. Herbs.—Species 2. South
Africa.  Alciope DC.

Marginal rows of stigmatic papillae not confluent at the apex of the style-branches. Heads small or middle-sized. Involucre oblong or campanulate.  287

287. Involucral bracts subequal, in 2 rows. Pappus-bristles numerous, unequal. Heads in panicles. Shrubs. Leaves marked with pellucid dots.—Species 1. Madagascar.  Glycideras Cass.

Involucral bracts unequal, in 3 or more rows.  288

288. Involucre broad-campanulate. Fruits 4-5-ribbed. Pappus-bristles in one row, intermixed with some shorter ones. Shrubs.—Species 3.
Madagascar.  Rochonia DC.

Involucre narrow-campanulate or oblong. Fruits 8-12-ribbed. Herbs.
Species 2. North Africa. Used as ornamental or medicinal plants.
“Goldenrod.”  Solidago L.

289. (276.) Pappus indistinct or wanting. Herbs.  290

Pappus of the central fruits formed of bristles or of scales and bristles.
[Subtribe ASTERINAE.]  292

290. Receptacle with scales between the flowers. Appendages of the style-branches linear, hairy all round. Involucral bracts membranous.
Leaves dissected.—Species 1. Tropics.  Chrysanthellum Rich.

Receptacle glabrous. Appendages of the style-branches triangular or lanceolate, hairy on the outer face only. [Subtribe BELLIDINAE.]  291

291. Involucral bracts scarious at the margin.—Species 2. Central Africa.  Brachycome Cass.

Involucral bracts herbaceous throughout. Leaves undivided.—Species
5. North Africa. Some are used as ornamental or medicinal plants.
“Daisy.”  Bellis L.

292. Inner fruits with a pappus of one-ranked feathery bristles, outer without
a pappus. Ray-flowers blue. Herbs.—Species 1. South Africa.
Used as an ornamental plant.  Charieis Cass.

Inner and outer fruits provided with a pappus.  293

293. Pappus of the inner fruits consisting of scales and bristles, that of the outer of scales only. Receptacle usually scaly. Heads solitary, terminating the branches. Herbs.—Species 12. South Africa.  Amellus L.

Pappus of all fruits consisting of bristles or of scales and bristles.  294

294. Pappus of 3-5 bristles intermixed with as many minute scales. Ray-flowers white. Heads solitary on leafless scapes. Herbs.—Species 1.
North Africa.  Bellium L.

Pappus of many bristles sometimes surrounded by some minute scales.  295

295. Pappus-bristles feathery. Herbs or undershrubs. (See 225.)  Mairia Nees

Pappus-bristles not feathery.  296

296. Fruits compressed.  297

Fruits not compressed. Heads in corymbs.  300

297. Fruits with 3-4 nerves on each side. Gummiferous shrubs or trees.—Species
4. Island of St. Helena.  Commidendron DC.

Fruits with 1-2 nerves on each side or without nerves. Herbs, undershrubs, or non-gummiferous shrubs.  298

298. Stem woody, at least at the base, rarely herbaceous throughout and then much branched at the base. Involucral bracts scarious at the edge, usually in several rows. Marginal flowers in 1-2 rows, with a strap-shaped corolla. Hairy appendages of the style-branches lanceolate.
Pappus-bristles in one row, rarely surrounded by a second of much shorter bristles. (See 231.)  Felicia Cass.

Stem herbaceous, not much branched at the base.  299

299. Involucral bracts membranous, in 2 rows. Marginal flowers usually in several rows, the outer with a narrow-linear corolla, the inner sometimes with a tubular one. Hairy appendages of the style-branches usually short, triangular. Pappus-bristles in 1 row, rarely in 2 rows. (See
232.)  Erigeron L.

Involucral bracts herbaceous or scarious at the edges, in 3 or more rows.
Marginal flowers in 1-2 rows, with a more or less strap-shaped, usually oblong corolla. Hairy appendages of the style-branches lanceolate.
Pappus-bristles in 2 or 3 rows. (See 232.)  Aster L.

300. Involucral bracts in 2 rows. Receptacle convex. Marginal flowers in 1 row, with a white, 3-toothed corolla. Fruits hairy. Pappus-bristles in one row. Downy undershrubs.—Species 1. Madagascar.  Henricia Cass.

Involucral bracts in 3 or more rows. Receptacle flat. Marginal flowers in 2 rows, with an entire corolla. Fruits glabrous. Pappus-bristles in
2 rows. Trees with blackish hairs.—Species 1. Island of St. Helena.  Melanodendron DC.

301. (266.) Pappus consisting of thin, hair-like, simple or toothed, but not feathery bristles, rarely outer fruits without a pappus. Receptacle without scales between the flowers. [Tribe SENECIONEAE.]  302

Pappus consisting of strong, awn-like or feathery bristles, or of scales sometimes united into a crown, or wanting.  316

302. Involucral bracts united below, one-ranked. [Subtribe OTHONNINAE.]  303

Involucral bracts free, at least at and after the time of flowering. [Subtribe
SENECIONINAE.]  306

303. Involucral bracts slightly united at the base. Inner disc-flowers sterile.
Marginal flowers with a strap-shaped corolla. Fruits hairy. Inner disc-fruits without a pappus, the other fruits with a pappus of numerous interwoven hairs. Perennial, nearly stem-less herbs. Leaves pinnately dissected. Heads solitary on a long scape. (See 236.)  Ruckeria DC.

Involucral bracts obviously united below. Disc-flowers all fertile.
Pappus on all fruits. Heads on a short scape or a branched stem.  304

304. Stem woody, at least at the base. Heads stalked. Marginal flowers with a strap-shaped corolla. Fruits 10-ribbed. Pappus of several rows of caducous bristles.—Species 40. South and Central Africa.
Some species yield a resin.  Euryops Cass.

Stem herbaceous throughout.  305

305. Stem annual, branched. Heads small. Corolla-lobes with a strong midnerve. Fruit 5-ribbed. Pappus of few caducous bristles.—Species
3. Central and South Africa.  Oligothrix DC.

Stem perennial, short and scape-like or branched; in the latter case pappus-bristles persistent. Heads medium-sized, solitary. Corolla-lobes without a strong midnerve.—Species 2. East Africa.  Werneria H. B. & Kunth

306. Female marginal flowers with a tubular or filiform corolla.  307

Female marginal flowers with a strap-shaped corolla or wanting.  311

307. Involucral bracts in 3 or more rows, imbricate, with scarious borders.
Female marginal flowers in several rows. Fruits without ribs. Heads solitary or in glomerules, yellow-flowered. (See 183.)  Phagnalon Cass.

Involucral bracts in 1-2 rows.