We can scarcely be said to have passed out of these rocks, when we begin to find new conditions in the earth. It is here to be observed that the subsequent rocks are formed, in a great measure, of matters derived from the substance of those which went before, but contain also beds of limestone, which is to no small extent composed of an ingredient which has not hitherto appeared. Limestone is a carbonate of lime, a secondary compound, of which one of the ingredients, carbonic acid gas, presents the element carbon, a perfect novelty in our progress. Whence this substance? The question is the more interesting, from our knowing that carbon is the main ingredient in organic things. There is reason to believe that its primeval condition was that of a gas, confined in the interior of the earth, and diffused in the atmosphere. The atmosphere still contains about a two-thousandth part of carbonic acid gas, forming the grand store from which the substance of each year’s crop of herbage and grain is derived, passing from herbage and grain into animal substance, and from animals again rendered back to the atmosphere in their expired breath, so that its amount is never impaired. Knowing this, when we hear of carbon beginning to appear in the ascending series of rocks, we are unavoidably led to consider it as marking a time of some importance in the earth’s history, a new era of natural conditions, one in which organic life has probably played a part.
It is not easy to suppose that, at this period, carbon was adopted directly in its gaseous form into rocks; for, if so, why should it not have been taken into earlier ones also? But we know that plants take it in, and transform it into substance; and we also know that there are classes of animals (marine polypes) which are capable of appropriating it, in connexion with lime, (carbonate of lime,) from the waters of the ocean, provided it be there in solution; and this substance do these animals deposit in masses (coral reefs) equal in extent to many strata. It has even been suggested, on strong grounds of probability, that a class of limestone beds are simply these reefs subjected to subsequent heat and pressure.
The appearance, then, of limestone beds in the early part of the stratified series, may be presumed to be connected with the fact of the commencement of organic life upon our planet, and, indeed, a consequent and a symptom of it.
It may not be out of place here to remark, that carbon is presumed to exist largely in the interior of the earth, from the fact of such considerable quantities of it issuing at this day, in the form of carbonic acid gas, from fissures and springs. The primeval and subsequent history of this element is worthy of much attention, and we shall have to revert to it as a matter greatly concerning our subject. Delabeche estimates the quantity of carbonic acid gas locked up in every cubic yard of limestone, at 16,000 cubic feet. The quantity locked up in coal, in which it forms from 64 to 75 per cent., must also be enormous. If all this were disengaged in a gaseous form, the constitution of the atmosphere would undergo a change, of which the first effect would be the extinction of life in all land animals. But a large proportion of it must have at one time been in the atmosphere. The atmosphere would then, of course, be incapable of supporting life in land animals. It is important, however, to observe that such an atmosphere would not be inconsistent with a luxuriant land vegetation; for experiment has proved that plants will flourish in air containing one-twelfth of this gas, or 166 times more than the present charge of our atmosphere. The results which we observe are perfectly consistent with, and may be said to presuppose an atmosphere highly charged with this gas, from about the close of the primary non-fossiliferous rocks to the termination of the carboniferous series, for there we see vast deposits (coal) containing carbon as a large ingredient, while at the same time the leaves of the Stone Book present no record of the contemporaneous existence of land animals.
The hypothesis of the connexion of the first limestone beds with the commencement of organic life upon our planet is supported by the fact, that in these beds we find the first remains of the bodies of animated creatures. My hypothesis may indeed be unsound; but, whether or not, it is clear, taking organic remains as upon the whole a faithful chronicle, that the deposition of these limestone beds was coeval with the existence of the earliest, or all but the earliest, living creatures upon earth.
And what were those creatures? It might well be with a kind of awe that the uninstructed inquirer would wait for an answer to this question. But nature is simpler than man’s wit would make her, and behold, the interrogation only brings before us the unpretending forms of various zoophytes and polypes, together with a few single and double-valved shell-fish (mollusks), all of them creatures of the sea. It is rather surprising to find these before any vegetable forms, considering that vegetables appear to us as forming the necessary first link in the chain of nutrition; but it is probable that there were sea plants, and also some simpler forms of animal life, before this period, although of too slight a substance to leave any fossil trace of their existence.
The exact point in the ascending stratified series at which the first traces of organic life are to be found is not clearly determined. Dr. M’Culloch states that he found fossil orthocerata (a kind of shell-fish) so early as the gneiss tract of Loch Eribol, in Sutherland; but Messrs. Sedgwick and Murchison, on a subsequent search, could not verify the discovery. It has also been stated, that the gneiss and mica tract of Bohemia contains some seams of grawacke, in which are organic remains; but British geologists have not as yet attached much importance to this statement. We have to look a little higher in the series for indubitable traces of organic life.
Above the gneiss and mica slate system, or group of strata, is the Clay Slate and Grawacke Slate System; that is to say, it is higher in the order of supraposition, though very often it rests immediately on the primitive granite. The sub-groups of this system are in the following succession upwards:—1, hornblende slate; 2, chiastolite slate; 3, clay slate; 4, Snowdon rocks, (grawacke and conglomerates;) 5, Bala limestone; 6, Plynlymmon rocks, (grawacke and grawacke slates, with beds of conglomerates.) This system is largely developed in the west and north of England, and it has been well examined, partly because some of the slate beds are extensively quarried for domestic purposes. If we overlook the dubious statements respecting Sutherland and Bohemia, we have in this “system” the first appearances of life upon our planet. The animal remains are chiefly confined to the slate beds, those named from Bala, in Wales, being the most prolific. Zoophyta, polyparia, crinoidea, conchifera, and crustacea, [60] are the orders of the animal kingdom thus found in the earliest of earth’s sepulchres. The orders are distinguished without difficulty, from the general characters of the creatures whose remains are found; but it is only in this general character that they bear a general resemblance to any creatures now existing. When we come to consider specific characters, we see that a difference exists—that, in short, the species and even genera are no longer represented upon earth. More than this, it will be found that the earliest species comparatively soon gave place to others, and that they are not represented even in the next higher group of rocks. One important remark has been made, that a comparatively small variety of species is found in the older rocks, although of some particular ones the remains are very abundant; as, for instance, of a species of asaphus, which is found between the laminæ of some of the slate rocks of Wales, and the corresponding rocks of Normandy and Germany in enormous quantities.
Ascending to the next group of rocks, we find the traces of life become more abundant, the number of species extended, and important additions made in certain vestiges of fuci, or sea-plants, and of fishes. This group of rocks has been called by English geologists, the Silurian System, because largely developed at the surface of a district of western England, formerly occupied by a people whom the Roman historians call Silures. It is a series of sandstones, limestones, and beds of shale (hardened mud), which are classed in the following sub-groups, beginning with the undermost:—1, Llandillo rocks, (darkish calcareous flagstones;) 2 and 3, two groups called Caradoc rocks; 4, Wenlock shale; 5, Wenlock limestone; 6, Lower Ludlow rocks, (shales and limestones;) 7, Aymestry limestone; 8, Upper Ludlow rocks, (shales and limestone, chiefly micaceous.) From the lowest beds upwards, there are polypiaria, though most prevalent in the Wenlock limestone; conchifera, a vast number of genera, but all of the order brachiopoda, (including terebratula, pentamerus, spirifer, orthis, leptæna;) mollusca, of several orders and many genera, (including turritella, orthoceras, nautilus, bellerophon;) crustacea, all of them trilobites, (including trinucleus, asaphus, calamene.) A little above the Llandillo rocks, there have been discovered certain convoluted forms, which are now established as annelids, or sea-worms, a tribe of creatures still existing, (nereidina and serpulina,) and which may often be found beneath stones on a sea-beach. One of these, figured by Mr. Murchison, is furnished with feet in vast numbers all along its body, like a centipede. The occurrence of annelids is important, on account of their character and status in the animal kingdom. They are red-blooded and hermaphrodite, and form a link of connexion between the annulosa (white-blooded worms) and a humble class of the vertebrata. [62] The Wenlock limestone is most remarkable amongst all the rocks of the Silurian system, for organic remains. Many slabs of it are wholly composed of corals, shells, and trilobites, held together by shale. It contains many genera of crinoidea and polypiaria, and it is thought that some beds of it are wholly the production of the latter creatures, or are, in other words, coral reefs transformed by heat and pressure into rocks. Remains of fishes, of a very minute size, have been detected by Mr. Philips in the Aymestry limestone, being apparently the first examples of vertebrated animals which breathed upon our planet. In the upper Ludlow rocks, remains of six genera of fish have been for a longer period known; they belong to the order of cartilaginous fishes, an order of mean organization and ferocious habits, of which the shark and sturgeon are living specimens. “Some were furnished with long palates, and squat, firmly-based teeth, well adapted for crushing the strong-cased zoophytes and shells of the period, fragments of which occur in the fœcal remains; some with teeth that, like the fossil sharks of the later formations, resemble lines of miniature pyramids, larger and smaller alternating; some with teeth sharp, thin, and so deeply serrated, that every individual tooth resembles a row of poniards set up against the walls of an armory; and these last, says Agassiz, furnished with weapons so murderous, must have been the pirates of the period. Some had their fins guarded with long spines, hooked like the beak of an eagle; some with spines of straighter and more slender form, and ribbed and furrowed longitudinally like columns; some were shielded by an armour of bony points, and some thickly covered with glistening scales.” [64]
The traces of fuci in this system are all but sufficient to allow of a distinction of genera. In some parts of North America, extensive though thin beds of them have been found. A distinguished French geologist, M. Brogniart, has shewn that all existing marine plants are classifiable with regard to the zones of climate; some being fitted for the torrid zone, some for the temperate, some for the frigid. And he establishes that the fuci of these early rocks speak of a torrid climate, although they may be found in what are now temperate regions; he also states that those of the higher rocks betoken, as we ascend, a gradually diminishing temperature.
We thus early begin to find proofs of the general uniformity of organic life over the surface of the earth, at the time when each particular system of rocks was formed. Species identical with the remains in the Wenlock limestone occur in the corresponding class of rocks in the Eifel, and partially in the Harz, Norway, Russia, and Brittany. The situations of the remains in Russia are fifteen hundred miles from the Wenlock beds; but at the distance of between six and seven thousand from those,—namely, in the vale of Mississippi, the same species are discovered. Uniformity in animal life over large geographical areas argues uniformity in the conditions of animal life; and hence arise some curious inferences. Species, in the same low class of animals, are now much more limited; for instance, the Red Sea gives different polypiaria, zoophytes, and shell-fish, from the Mediterranean. It is the opinion of M. Brogniart, that the uniformity which existed in the primeval times can only be attributed to the temperature arising from the internal heat, which had yet, as he supposes, been sufficiently great to overpower the ordinary meteorological influences, and spread a tropical clime all over the globe.
We advance to a new chapter in this marvellous history—the era of the Old Red Sandstone System. This term has been recently applied to a series of strata, of enormous thickness in the whole mass, largely developed in Herefordshire, Shropshire, Worcestershire, and South Wales; also in the counties of Fife, Forfar, Moray, Cromarty, and Caithness; and in Russia and North America, if not in many other parts of the world. The particular strata forming the system are somewhat different in different countries; but there is a general character to the extent of these being a mixture of flagstones, marly rocks, and sandstones, usually of a laminous structure, with conglomerates. There is also a schist shewing the presence of bitumen; a remarkable new ingredient, since it is a vegetable production. In the conglomerates, of great extent and thickness, which form, in at least one district, the basis or leading feature of the system, inclosing water-worn fragments of quartz and other rocks, we have evidence of the seas of that period having been subjected to a violent and long-continued agitation, probably from volcanic causes. The upper members of the series bear the appearance of having been deposited in comparatively tranquil seas. The English specimens of this system shew a remarkable freedom from those disturbances which result in the interjection of trap; and they are thus defective in mineral ores. In some parts of England the old red sandstone system has been stated as 10,000 feet in thickness.
In this era, the forms of life which existed in the Silurian are continued: we have the same orders of marine creatures, zoophyta, polypiaria, conchifera, crustacea; but to these are added numerous fishes, some of which are of most extraordinary and surprising forms. Several of the strata are crowded with remains of fish, shewing that the seas in which those beds were deposited had swarmed with that class of inhabitants. The investigation of this system is recent; but already [68] M. Agassiz has ascertained about twenty genera, and thrice the number of species. And it is remarkable that the Silurian fishes are here only represented in genera; the whole of the species of that era had already passed away. Even throughout the sub-groups of the system itself, the species are changed; and these are phenomena observed throughout all the subsequent systems or geological eras; apparently arguing that, during the deposition of all the rocks, a gradual change of physical conditions was constantly going on. A varying temperature, or even a varying depth of sea, would at present be attended with similar changes in marine life; and by analogy we are entitled to assume that such variations in the ancient seas might be amongst the causes of that constant change of genera and species in the inhabitants of those seas to which the organic contents of the rocks bear witness.
Some of the fossils of this system,—the cephalaspis, coccosteus, pterichthys, holoptychius—are, in form and structure, entirely different from any fishes now existing, only the sturgeon family having any trace of affinity to them in any respect. They seem to form a sort of connecting link between the crustacea and true fishes.
The cephalaspis may be considered as making the smallest advance from the crustacean character; it very much resembles in form the asaphus of lower formations, having a longish tail-like body inserted within the cusp of a large crescent-shaped head, somewhat like a saddler’s cutting-knife. The body is covered with strong plates of bone, enamelled, and the head was protected on the upper side with one large plate, as with a buckler—hence the name, implying buckler-head. A range of small fins conveys the idea of its having been as weak in motion as it is strong in structure. The coccosteus may be said to mark the next advance to fish creation. The outline of its body is of the form of a short thick coffin, rounded, covered with strong bony plates, and terminating in a long tail, which seems to have been the sole organ of motion. It is very remarkable, that, while the tail establishes this creature among the vertebrata and the fishes, its mouth has been opened vertically, like those of the crustaceans, but which is contrary to the mode of vertebrata generally. This seems a pretty strong mark of the link character of the coccosteus between these two great departments of the animal kingdom. The pterichthys has also strong bony plates over its body, arranged much like those of a tortoise, and has a long tail; but its most remarkable feature, and that which has suggested its name, is a pair of long and narrow wing-like appendages attached to the shoulders, which the creature is supposed to have erected for its defence when attacked by an enemy.
The holoptychius is of a flat oval form, furnished with fins, and ending in a long tail; the whole body covered with strong plates which overlap each other, and the head forming only a slight rounded projection from the general figure. The specimens in the lower beds are not above the size of a flounder; but in the higher strata, to judge by the size of the scales or plates which have been found, the creature attained a comparatively monstrous size.
The other fishes of the system,—the osteolepis, glyptolepis, dipterus, &c., are, in general outline, much like fishes still existing, but their organization has, nevertheless, some striking peculiarities. They have been entirely covered with bony scales or plates, enamelled externally; their spines are tipped with bone, and, as one striking and unvarying feature, the tail is only finned on the lower side. The internal skeleton, of which no traces have been preserved, is presumed to have been cartilaginous. They therefore unite the character of cartilaginous fishes with a character peculiar to themselves, and in which we see pretty clear vestiges of the pre-existent crustaceous form.
With regard to the link character of these animals, some curious facts are mentioned. It appears that in the imperfect condition of the vertebral column, and the inferior situation of the mouth in the pterichthys, coccosteus, &c., there is an analogy to the form of the dorsal cord and position of the mouth in the embryo of perfect fishes. The one-sided form of the tail in the osteolepis &c. finds a similar analogy in the form of the tail in the embryo of the salmon. It is not premature to remark how broadly these facts seem to hint at a parity of law affecting the progress of general creation, and the progress of an individual fœtus of one of the more perfect animals.
It is equally ascertained of the types of being prevalent in the old red, as of those of the preceding system, that they are uniform in the corresponding strata of distant parts of the earth; for instance, Russia and North America.
In the old red sandstone, the marine plants, of which faint traces are observable in the Silurians, continue to appear. It would seem as if less change took place in the vegetation than in the animals of those early seas; and for this, as Mr. Miller has remarked, it is easy to imagine reasons. For example, an infusion of lime into the sea would destroy animal life, but be favourable to vegetation.
As yet there were no land animals or plants, and for this the presumable reason is, that no dry land as yet existed. We are not left to make this inference solely from the absence of land animals and plants; in the arrangement of the primary (stratified) rocks, we have further evidence of it. That these rocks were formed in a generally horizontal position, we are as well assured as that they were formed at the bottom of seas. But they are always found greatly inclined in position, tilted up against the slopes of the granitic masses which are beneath them in geological order, though often shooting up to a higher point in the atmosphere. No doubt can be entertained that these granitic masses, forming our principal mountain ranges, have been protruded from below, or, at least, thrust much further up, since the deposition of the primary rocks. The protrusion was what tilted up the primary rocks; and the inference is, of course, unavoidable, that these mountains have risen chiefly, at least, since the primary rocks were laid down. It is remarkable that, while the primary rocks thus incline towards granitic nuclei or axes, the strata higher in the series rest against these again, generally at a less inclination, or none at all, shewing that these strata were laid down after the swelling mountain eminences had, by their protrusion, tilted up the primary strata. And thus it may be said an era of local upthrowing of the primitive and (perhaps) central matter of our planet, is established as happening about the close of the primary strata, and beginning of the next ensuing system. It may be called the Era of the Oldest Mountains, or, more boldly, of the formation of the detached portions of dry land over the hitherto watery surface of the globe—an important part of the designs of Providence, for which the time was now apparently come. It may be remarked, that volcanic disturbances and protrusions of trap took place throughout the whole period of the deposition of the primary rocks; but they were upon a comparatively limited scale, and probably all took place under water. It was only now that the central granitic masses of the great mountain ranges were thrown up, carrying up with them broken edges of the primary strata; a process which seems to have had this difference from the other, that it was the effect of a more tremendous force exerted at a lower depth in the earth, and generally acting in lines pervading a considerable portion of the earth’s surface. We shall by-and-by see that the protrusion of some of the mountain ranges was not completed, or did not stop, at that period. There is no part of geological science more clear than that which refers to the ages of mountains. It is as certain that the Grampian mountains of Scotland are older than the Alps and Apennines, as it is that civilization had visited Italy, and had enabled her to subdue the world, while Scotland was the residence of “roving barbarians.” The Pyrenees, Carpathians, and other ranges of continental Europe, are all younger than the Grampians, or even the insignificant Mendip Hills of southern England. Stratification tells this tale as plainly as Livy tells the history of the Roman republic. It tells us—to use the words of Professor Philips—that at the time when the Grampians sent streams and detritus to straits where now the valleys of the Forth and Clyde meet, the greater part of Europe was a wide ocean.
The last three systems—called, in England, the Cumbrian, Silurian, and Devonian, and collectively the palæozoic rocks, from their containing the remains of the earliest inhabitants of the globe—are of vast thickness; in England, not much less than 30,000 feet, or nearly six miles. In other parts of the world, as we have seen, the earliest of these systems alone is of much greater depth—arguing an enormous profundity in the ocean in which they were formed.
We now enter upon a new great epoch in the history of our globe. There was now dry land. As a consequence of this fact, there was fresh water, for rain, instead of immediately returning to the sea, as formerly, was now gathered in channels of the earth, and became springs, rivers, and lakes. There was now a theatre for the existence of land plants and animals, and it remains to be inquired if these accordingly were produced.
The Secondary Rocks, in which our further researches are to be prosecuted, consist of a great and varied series, resting, generally unconformably, against flanks of the upturned primary rocks, sometimes themselves considerably inclined, at others, forming extensive basin-like beds, nearly horizontal; in many places, much broken up and shifted by disturbances from below. They have all been formed out of the materials of the older rocks, by virtue of the wearing power of air and water, which is still every day carrying down vast quantities of the elevated matter of the globe into the sea. But the separate strata are each much more distinct in the matter of its composition than might be expected. Some are siliceous or arenaceous (sandstones), composed mainly of fine grains from the quartz rocks—the most abundant of the primary strata. Others are argillaceous—clays, shales, &c., chiefly derived, probably, from the slate beds of the primary series. Others are calcareous, derived from the early limestone. As a general feature, they are softer and less crystalline than the primary rocks, as if they had endured less of both heat and pressure than the senior formation. There are beds (coal) formed solely of vegetable matter, and some others in which the main ingredient is particles of iron, (the iron black band.) The secondary rocks are quite as communicative with regard to their portion of the earth’s history as the primitive were.
The first, or lowest, group of the secondary rocks is called the Carboniferous Formation, from the remarkable feature of its numerous interspersed beds of coal. It commences with the beds of the mountain limestone, which, in some situations, as in Derbyshire and Ireland, are of great thickness, being alternated with chert (a siliceous sandstone), sandstones, shales, and beds of coal, generally of the harder and less bituminous kind (anthracite), the whole being covered in some places by the millstone grit, a siliceous conglomerate composed of the detritus of the primary rocks. The mountain limestone, attaining in England to a depth of eight hundred yards, greatly exceeds in volume any of the primary limestone beds, and shews an enormous addition of power to the causes formerly suggested as having produced this substance. In fact, remains of corals, crinoidea, and shells, are so abundant in it, as to compose three-fourths of the mass in some parts. Above the mountain limestone commence the more conspicuous coal beds, alternating with sandstones, shales, beds of limestone, and ironstone. Coal is altogether composed of the matter of a terrestrial vegetation, transmuted by pressure. Some fresh-water shells have been found in it, but few of marine origin, and no remains of those zoophytes and crinoidea so abundant in the mountain limestone and other rocks. Coal beds exist in Europe, Asia, and America, and have hitherto been esteemed as the most valuable of mineral productions, from the important services which the substance renders in manufactures and in domestic economy. It is to be remarked, that there are some local variations in the arrangement of coal beds. In France, they rest immediately on the granite and other primary rocks, the intermediate strata not having been found at those places. In America, the kind called anthracite occurs among the slate beds, and this species also abounds more in the mountain limestone than with us. These last circumstances only shew that different parts of the earth’s surface did not all witness the same events of a certain fixed series exactly at the same time. There had been an exhibition of dry land about the site of America, a little earlier than in Europe.
Some features of the condition of the earth during the deposition of the carboniferous group, are made out with a clearness which must satisfy most minds. First we are told of a time when carbonate of lime was formed in vast abundance at the bottoms of profound seas, accompanied by an unusually large population of corals and encrinites; while in some parts of the earth there were patches of dry land, covered with a luxuriant vegetation. Next we have a comparatively brief period of volcanic disturbance, (when the conglomerate was formed.) Then the causes favourable to the so abundant production of limestone, and the large population of marine acrita, decline, and we find the masses of dry land increase in number and extent, and begin to bear an amount of forest vegetation, far exceeding that of the most sheltered tropical spots of the present surface. The climate, even in the latitude of Baffin’s Bay, was torrid, and perhaps the atmosphere contained a larger charge of carbonic acid gas (the material of vegetation) than it now does. The forests or thickets of the period, included no species of plants now known upon earth. They mainly consisted of gigantic shrubs, which are either not represented by any existing types, or are akin to kinds which are now only found in small and lowly forms. That these forests grew upon a Polynesia, or multitude of small islands, is considered probable, from similar vegetation being now found in such situations within the tropics. With regard to the circumstances under which the masses of vegetable matter were transformed into successive coal strata, geologists are divided. From examples seen at the present day, at the mouths of such rivers as the Mississippi, which traverse extensive sylvan regions, and from other circumstances to be adverted to, it is held likely by some that the vegetable matter, the rubbish of decayed forests, was carried by rivers into estuaries, and there accumulated in vast natural rafts, until it sunk to the bottom, where an overlayer of sand or mud would prepare it for becoming a stratum of coal. Others conceive that the vegetation first went into the condition of a peat moss, that a sink in the level then exposed it to be overrun by the sea, and covered with a layer of sand or mud; that a subsequent uprise made the mud dry land, and fitted it to bear a new forest, which afterwards, like its predecessor, became a bed of peat; that, in short, by repetitions of this process, the alternate layers of coal, sandstone, and shale, constituting the carboniferous group, were formed. It is favourable to this last view that marine fossils are scarcely found in the body of the coal itself, though abundant in the shale layers above and below it; also that in several places erect stems of trees are found with their roots still fixed in the shale beds, and crossing the sandstone beds at almost right angles, shewing that these, at least, had not been drifted from their original situations. On the other hand, it is not easy to admit such repeated risings and sinkings of surface as would be required, on this hypothesis, to form a series of coal strata. Perhaps we may most safely rest at present with the supposition that coal has been formed under both classes of circumstances, though in the latter only as an exception to the former.
Upwards of three hundred species of plants have been ascertained to exist in the coal formation; but it is not necessary to suppose that the whole contained in that system are now, or ever will be distinguished. Experiments shew that some great classes of plants become decomposed in water in a much less space of time than others, and it is remarkable that those which decompose soonest, are of the classes found most rare, or not at all, in the coal strata. It is consequently to be inferred that there may have been grasses and mosses at this era, and many species of trees, the remains of which had lost all trace of organic form before their substance sunk into the mass of which coal was formed. In speaking, therefore, of the vegetation of this period, we must bear in mind that it may have comprehended forms of which we have no memorial.
Supposing, nevertheless, that, in the main, the ascertained vegetation of the coal system is that which grew at the time of its formation, it is interesting to find that the terrestrial botany of our globe begins with classes of comparatively simple forms and structure. In the ranks of the vegetable kingdom, the lowest place is taken by plants of cellular tissue, and which have no flowers, (cryptogamia,) as lichens, mosses, fungi, ferns, sea-weeds. Above these stand plants of vascular tissue, and bearing flowers, in which again there are two great subdivisions; first, plants having one seed-lobe, (monocotyledons,) and in which the new matter is added within, (endogenous,) of which the cane and palm are examples; second, plants having two seed-lobes, (dicotyledons,) and in which the new matter is added on the outside under the bark, (exogenous,) of which the pine, elm, oak, and most of the British forest-trees are examples; these subdivisions also ranking in the order in which they are here stated. Now it is clear that a predominance of these forms in succession marked the successive epochs developed by fossil geology; the simple abounding first, and the complex afterwards.
Two-thirds of the plants of the carboniferous era are of the cellular or cryptogamic kind, a proportion which would probably be much increased if we knew the whole Flora of that era. The ascertained dicotyledons, or higher-class plants, are comparatively few in this formation; but it will be found that they constantly increased as the globe grew older.
The master-form or type of the era was the fern, or breckan, of which about one hundred and thirty species have already been ascertained as entering into the composition of coal. [84a] The fern is a plant which thrives best in warm, shaded, and moist situations. In tropical countries, where these conditions abound, there are many more species than in temperate climes, and some of these are arborescent, or of a tree-like size and luxuriance. [84b] The ferns of the coal strata have been of this magnitude, and that without regard to the parts of the earth where they are found. In the coal of Baffin’s Bay, of Newcastle, and of the torrid zone alike, are the fossil ferns arborescent, shewing clearly that, in that era, the present tropical temperature, or one even higher, existed in very high latitudes.
In the swamps and ditches of England there grows a plant called the horse-tail (equisetum), having a succulent, erect, jointed stem, with slender leaves, and a scaly catkin at the top. A second large section of the plants of the carboniferous era were of this kind (equisetaceæ), but, like the fern, reaching the magnitudes of trees. While existing equiseta rarely exceed three feet in height, and the stems are generally under half an inch in diameter, their kindred, entombed in the coal beds, seem to have been generally fourteen or fifteen feet high, with stems from six inches to a foot in thickness. Arborescent plants of this family, like the arborescent ferns, now grow only in tropical countries, and their being found in the coal beds in all latitudes is consequently held as an additional proof, that at this era a warm climate was extended much farther to the north than at present. It is to be remarked that plants of this kind (forming two genera, the most abundant of which is the calamites) are only represented on the present surface by plants of the same family: the species which flourished at this era gradually lessen in number as we advance upwards in the series of rocks, and disappear before we arrive at the tertiary formation.
The club-moss family (lycopodiaceæ) are other plants of the present surface, usually seen in a lowly and creeping form in temperate latitudes, but presenting species which rise to a greater magnitude within the tropics. Many specimens of this family are found in the coal beds; it is thought they have contributed more to the substance of the coal than any other family. But, like the ferns and equisetaceæ, they rise to a prodigious magnitude. The lepidodendra (so the fossil genus is called) have probably been from sixty-five to eighty feet in height, having at their base a diameter of about three feet, while their leaves measured twenty inches in length. In the forests of the coal era, the lepidodendra would enjoy the rank of firs in our forests, affording shade to the only less stately ferns and calamites. The internal structure of the stem, and the character of the seed-vessels, shew them to have been a link between single-lobed and double-lobed plants, a fact worthy of note, as it favours the idea that, in vegetable, as well as animal creation, a progress has been observed, in conformity with advancing conditions. It is also curious to find a missing link of so much importance in a genus of plants which has long ceased to have a living place upon earth.
The other leading plants of the coal era are without representatives on the present surface, and their characters are in general less clearly ascertained. Amongst the most remarkable are—the sigillaria, of which large stems are very abundant, shewing that the interior has been soft, and the exterior fluted with separate leaves inserted in vertical rows along the flutings—and the stigmaria, plants apparently calculated to flourish in marshes or pools, having a short, thick, fleshy stem, with a dome-shaped top, from which sprung branches of from twenty to thirty feet long. Amongst monocotyledons were some palms, (flabellaria and næggerathia,) besides a few not distinctly assignable to any class.
The dicotyledons of the coal are comparatively few, though on the present surface they are the most numerous sub-class. Besides some of doubtful affinity, (annularia, asterophyllites, &c.,) there were a few of the pine family, which seem to have been the highest class of trees of this era, and are only as yet found in isolated cases, and in sandstone beds. The first discovered lay in the Craigleith quarry, near Edinburgh, and consisted of a stem about two feet thick, and forty-seven feet in length. Others have since been found, both in the same situation, and at Newcastle. Leaves and fruit being wanting, an ingenious mode of detecting the nature of these trees was hit upon by Mr. Witham of Lartington. Taking thin polished cross slices of the stem, and subjecting them to the microscope, he detected the structure of the wood to be that of a cone-bearing tree, by the presence of certain “reticulations” which distinguish that family, in addition to the usual radiating and concentric lines. That particular tree was concluded to be an araucaria, a species now found in Norfolk Island, in the South Sea, and in a few other remote situations. The coniferæ of this era form the dawn of dicotyledenous trees, of which they may be said to be the simplest type, and to which, it has already been noticed, the lepidodendra are a link from the monocotyledons. The concentric rings of the Craigleith and other coniferæ of this era have been mentioned. It is interesting to find in these a record of the changing seasons of those early ages, when as yet there were no human beings to observe time or tide. They are clearly traced; but it is observed that they are more slightly marked than is the case with their family at the present day, as if the changes of temperature had been within a narrower range.
Such was the vegetation of the carbonigenous era, composed of forms at the bottom of the botanical scale, flowerless, fruitless, but luxuriant and abundant beyond what the most favoured spots on earth can now shew. The rigidity of the leaves of its plants, and the absence of fleshy fruits and farinaceous seeds, unfitted it to afford nutriment to animals; and, monotonous in its forms, and destitute of brilliant colouring, its sward probably unenlivened by any of the smaller flowering herbs, its shades uncheered by the hum of insects, or the music of birds, it must have been but a sombre scene to a human visitant. But neither man nor any other animals were then in existence to look for such uses or such beauties in this vegetation. It was serving other and equally important ends, clearing (probably) the atmosphere of matter noxious to animal life, and storing up mineral masses which were in long subsequent ages to prove of the greatest service to the human race, even to the extent of favouring the progress of its civilization.
The animal remains of this era are not numerous, in comparison with those which go before, or those which come after. The mountain limestone, indeed, deposited at the commencement of it, abounds unusually in polypiaria and crinoidea; but when we ascend to the coal-beds themselves, the case is altered, and these marine remains altogether disappear. We have then only a limited variety of conchifers and shell mollusks, with fragments of a few species of fishes, and these are rarely or never found in the coal seams, but in the shales alternating with them. Some of the fishes are of a sauroid character, that is, partake of the nature of the lizard, a genus of the reptilia, a land class of animals, so that we may be said here to have the first approach to a kind of animals calculated to breathe the atmosphere. Such is the Megalichthys Hibbertii, found by Dr. Hibbert Ware, in a limestone bed of fresh-water origin, underneath the coal at Burdiehouse, near Edinburgh. Others of the same kind have been found in the coal measures in Yorkshire, and in the low coal shales at Manchester. This is no more than might be expected, as collections of fresh water now existed, and it is presumable that they would be peopled. The chief other fishes of the coal era are named palæothrissum, palæoniscus, diperdus.
Coal strata are nearly confined to the group termed the carboniferous formation. Thin beds are not unknown afterwards, but they occur only as a rare exception. It is therefore thought that the most important of the conditions which allowed of so abundant a terrestrial vegetation, had ceased about the time when this formation was closed. The high temperature was not one of the conditions which terminated, for there are evidences of it afterwards; but probably the superabundance of carbonic acid gas supposed to have existed during this era was expended before its close. There can be little doubt that the infusion of a large dose of this gas into the atmosphere at the present day would be attended by precisely the same circumstances as in the time of the carboniferous formation. Land animal life would not have a place on earth; vegetation would be enormous; and coal strata would be formed from the vast accumulations of woody matter, which would gather in every sea, near the mouths of great rivers. On the exhaustion of the superabundance of carbonic acid gas, the coal formation would cease, and the earth might again become a suitable theatre of being for land animals.
The termination of the carboniferous formation is marked by symptoms of volcanic violence, which some geologists have considered to denote the close of one system of things and the beginning of another. Coal beds generally lie in basins, as if following the curve of the bottom of seas. But there is no such basin which is not broken up into pieces, some of which have been tossed up on edge, others allowed to sink, causing the ends of strata to be in some instances many yards, and in a few several hundred feet, removed from the corresponding ends of neighbouring fragments. These are held to be results of volcanic movements below, the operation of which is further seen in numerous upbursts and intrusions of volcanic rock (trap). That these disturbances took place about the close of the formation, and not later, is shewn in the fact of the next higher group of strata being comparatively undisturbed. Other symptoms of this time of violence are seen in the beds of conglomerate which occur amongst the first strata above the coal. These, as usual, consist of fragments of the elder rocks, more or less worn from being tumbled about in agitated water, and laid down in a mud paste, afterwards hardened. Volcanic disturbances break up the rocks; the pieces are worn in seas; and a deposit of conglomerate is the consequence. Of porphyry, there are some such pieces in the conglomerate of Devonshire, three or four tons in weight. It is to be admitted for strict truth that, in some parts of Europe, the carboniferous formation is followed by superior deposits, without the appearance of such disturbances between their respective periods; but apparently this case belongs to the class of exceptions already noticed. [93] That disturbance was general, is supported by the further and important fact of the destruction of many forms of organic being previously flourishing, particularly of the vegetable kingdom.
The next volume of the rock series refers to an era distinguished by an event of no less importance than the commencement of land animals. The New Red Sandstone System is subdivided into groups, some of which are wanting in some places; they are pretty fully developed in the north of England, in the following ascending order:—1. Lower red sandstone; 2. Magnesian limestone; 3. Red and white sandstones and conglomerate; 4. Variegated marls. Between the third and fourth there is, in Germany, another group, called the Muschelkalk, a word expressing a limestone full of shells.
The first group, containing the conglomerates already adverted to, seems to have been produced during the time of disturbance which occurred so generally after the carbonigenous era. This new era is distinguished by a paucity of organic remains, as might partly be expected from the appearances of disturbance, and the red tint of the rocks, the latter being communicated by a solution of oxide of iron, a substance unfavourable to animal life.
The second group is a limestone with an infusion of magnesia. It is developed less generally than some others, but occurs conspicuously in England and Germany. Its place, above the red sandstone, shews the recurrence of circumstances favourable to animal life, and we accordingly find in it not only zoophytes, conchifera, and a few tribes of fish, but some faint traces of land plants, and a new and startling appearance—a reptile of saurian (lizard) character, analogous to the now existing family called monitors. Remains of this creature are found in cupriferous (copper-bearing) slate connected with the mountain limestone, at Mansfield and Glucksbrunn, in Germany, which may be taken as evidence that dry land existed in that age near those places. The magnesia limestone is also remarkable as the last rock in which appears the leptæna, or producta, a conchifer of numerous species which makes a conspicuous appearance in all previous seas. It is likewise to be observed, that the fishes of this age, to the genera of which the names palæoniscus, catopterus, platysomus, &c., have been applied, vanish, and henceforth appear no more.
The third group, chiefly sandstones, variously coloured according to the amount and nature of the metallic oxide infused into them, shews a recurrence of agitation, and a consequent diminution of the amount of animal life. In the upper part, however, of this group, there are abundant symptoms of a revival of proper conditions for such life. There are marl beds, the origin of which substance in decomposed shells is obvious; and in Germany, though not in England, here occurs the muschelkalk, containing numerous organic remains, (generally different from those of the magnesian limestone,) and noted for the specimens of land animals, which it is the first to present in any considerable abundance to our notice.
These animals are of the vertebrate sub-kingdom, but of its lowest class next after fishes,—namely, reptiles,—a portion of the terrestrial tribes whose imperfect respiratory system perhaps fitted them for enduring an atmosphere not yet quite suitable for birds or mammifers. [97] The specimens found in the muschelkalk are allied to the crocodile and lizard tribes of the present day, but in the latter instance are upon a scale of magnitude as much superior to present forms as the lepidodendron of the coal era was superior to the dwarf club-mosses of our time. These saurians also combine some peculiarities of structure of a most extraordinary character.
The animal to which the name ichthyosaurus has been given, was as long as a young whale, and it was fitted for living in the water, though breathing the atmosphere. It had the vertebral column and general bodily form of a fish, but to that were added the head and breast-bone of a lizard, and the paddles of the whale tribes. The beak, moreover, was that of a porpoise, and the teeth were those of a crocodile. It must have been a most destructive creature to the fish of those early seas.
The plesiosaurus was of similar bulk, with a turtle-like body and paddles, shewing that the sea was its element, but with a long serpent-like neck, terminating in a saurian head, calculated to reach prey at a considerable distance. These two animals, of which many varieties have been discovered, constituting distinct species, are supposed to have lived in the shallow borders of the seas of this and subsequent formations, devouring immense quantities of the finny tribes. It was at first thought that no creatures approaching them in character now inhabit the earth; but latterly Mr. Darwin has discovered, in the reptile-peopled Galapagos Islands, in the South Sea, a marine saurian from three to four feet long.
The megalosaurus was an enormous lizard—a land creature, also carnivorous. The pterodactyle was another lizard, but furnished with wings to pursue its prey in the air, and varying in size between a cormorant and a snipe. Crocodiles abounded, and some of these were herbivorous. Such was the iguanodon, a creature of the character of the iguana of the Ganges, but reaching a hundred feet in length, or twenty times that of its modern representative.
There were also numerous tortoises, some of them reaching a great size; and Professor Owen has found in Warwickshire some remains of an animal of the batrachian order, [99] to which, from the peculiar form of the teeth, he has given the name of labyrinthidon. Thus, three of Cuvier’s four orders of reptilia (sauria, chelonia, and batrachia) are represented in this formation, the serpent order (ophidia) being alone wanting.
The variegated marl beds which constitute the uppermost group of the formation, present two additional genera of huge saurians,—the phytosaurus and mastodonsaurus.
It is in the upper beds of the red sandstone that beds of salt first occur. These are sometimes of such thickness, that the mine from which the material has been excavated looks like a lofty church. We see in the present world no circumstances calculated to produce the formation of a bed of rock salt; yet it is not difficult to understand how such strata were formed in an age marked by ultra-tropical heat and frequent volcanic disturbances. An estuary, cut off by an upthrow of trap, or a change of level, and left to dry up under the heat of the sun, would quickly become the bed of a dense layer of rock salt. A second shift of level, or some other volcanic disturbance, connecting it again with the sea, would expose this stratum to being covered over with a layer of sand or mud, destined in time to form the next stratum of rock above it.
The plants of this era are few and unobtrusive. Equiseta, calamites, ferns, Voltzia, and a few of the other families found so abundantly in the preceding formation, here present themselves, but in diminished size and quantity.
This seems to be the proper place to advert to certain memorials of a peculiar and unexpected character respecting these early ages in the sandstones. So low as the bottom of the carboniferous system, slabs are found marked over a great extent of surface with that peculiar corrugation or wrinkling which the receding tide leaves upon a sandy beach when the sea is but slightly agitated; and not only are these ripple-marks, as they are called, found on the surfaces, but casts of them are found on the under sides of slabs lying above. The phenomena suggests the time when the sand ultimately formed into these stone slabs, was part of the beach of a sea of the carbonigenous era; when, left wavy by one tide, it was covered over with a thin layer of fresh sand by the next, and so on, precisely as such circumstances might be expected to take place at the present day. Sandstone surfaces, ripple-marked, are found throughout the subsequent formations: in those of the new red, at more than one place in England, they further bear impressions of rain-drops which have fallen upon them—the rain, of course, of the inconceivably remote age in which the sandstones were formed. In the Greensill sandstone, near Shrewsbury, it has even been possible to tell from what direction the shower came which impressed the sandy surface, the rims of the marks being somewhat raised on one side, exactly as might be expected from a slanting shower falling at this day upon one of our beaches. These facts have the same sort of interest as the season rings of the Craigleith conifers, as speaking of a parity between some of the familiar processes of nature in those early ages and our own.
In the new red sandstone, impressions still more important in the inferences to which they tend, have been observed,—namely, the footmarks of various animals. In a quarry of this formation, at Corncockle Muir, in Dumfriesshire, where the slabs incline at an angle of thirty-eight degrees, the vestiges of an animal supposed to have been a tortoise are distinctly traced up and down the slope, as if the creature had had occasion to pass backwards and forwards in that direction only, possibly in its daily visits to the sea. Some slabs similarly impressed, in the Stourton quarries in Cheshire, are further marked with a shower of rain which we know must have fallen afterwards, for its little hollows are impressed in the footmarks also, though more slightly than on the rest of the surface, the comparative hardness of a trodden place having apparently prevented so deep an impression being made. At Hessberg, in Saxony, the vestiges of four distinct animals have been traced, one of them a web-footed animal of small size, considered as a congener of the crocodile; another, whose footsteps having a resemblance to an impression of a swelled human hand, has caused it to be named the cheirotherium. The footsteps of the cheirotherium have been found also in the Stourton quarries above mentioned. Professor Owen, who stands at the head of comparative anatomy in the present day, has expressed his belief that this last animal was the same batrachian of which he has found fragments in the new red sandstone of Warwickshire. At Runcorn, near Manchester, and elsewhere, have been discovered the tracks of an animal which Mr. Owen calls the rynchosaurus, uniting with the body of a reptile the beak and feet of a bird, and which clearly had been a link between these two classes.
If geologists shall ultimately give their approbation to the inferences made from a recent discovery in America, we shall have the addition of perfect birds, though probably of a low type, to the animal forms of this era. It is stated to be in quarries of this rock, in the valley of Connecticut, that footprints have been found, apparently produced by birds of the order grallæ, or waders. “The footsteps appear in regular succession on the continuous track of an animal, in the act of walking or running, with the right and left foot always in their relative places. The distance of the intervals between each footstep on the same track is occasionally varied, but to no greater amount than may be explained by the bird having altered its pace. Many tracks of different individuals and different species are often found crossing each other, and crowded, like impressions of feet upon the shores of a muddy stream, where ducks and geese resort.” [103] Some of these prints indicate small animals, but others denote birds of what would now be an unusually large size. One animal, having a foot fifteen inches in length, (one-half more than that of the ostrich,) and a stride of from four to six feet, has been appropriately entitled, ornithichnites giganteus.
The chronicles of this period consist of a series of beds, mostly calcareous, taking their general name (Oolite System) from a conspicuous member of them—the oolite—a limestone composed of an aggregation of small round grains or spherules, and so called from its fancied resemblance to a cluster of eggs, or the roe of a fish. This texture of stone is novel and striking. It is supposed to be of chemical origin, each spherule being an aggregation of particles round a central nucleus. The oolite system is largely developed in England, France, Westphalia, and Northern Italy; it appears in Northern India and Africa, and patches of it exist in Scotland, and in the vale of the Mississippi. It may of course be yet discovered in many other parts of the world.
The series, as shewn in the neighbourhood of Bath, is (beginning with the lowest) as follows:—1. Lias, a set of strata variously composed of limestone, clay, marl, and shale, clay being predominant; 2. Lower oolitic formation, including, besides the great oolite bed of central England, fullers’ earth beds, forest marble, and cornbrash; 3. Middle oolitic formation, composed of two sub-groups, the Oxford clay and coral rag, the latter being a mere layer of the works of the coral polype; 4. Upper oolitic formation, including what are called Kimmeridge clay and Portland oolite. In Yorkshire there is an additional group above the lias, and in Sutherlandshire there is another group above that again. In the wealds (moorlands) of Kent and Sussex, there is, in like manner, above the fourth of the Bath series, another additional group, to which the name of the Wealden has been given, from its situation, and which, composed of sandstones and clays, is subdivided into Purbeck beds, Hastings sand, and Weald clay.
There are no particular appearances of disturbance between the close of the new red sandstone and the beginning of the oolite system, as far as has been observed in England. Yet there is a great change in the materials of the rocks of the two formations, shewing that while the bottoms of the seas of the one period had been chiefly arenaceous, those of the other were chiefly clayey and limy. And there is an equal difference between the two periods in respect of both botany and zoology. While the new red sandstone shews comparatively scanty traces of organic creation, those in the oolite are extremely abundant, particularly in the department of animals, and more particularly still of sea mollusca, which, it has been observed, are always the more conspicuous in proportion to the predominance of calcareous rocks. It is also remarkable that the animals of the oolitic system are entirely different in species from those of the preceding age, and that these species cease before the next. In this system we likewise find that uniformity over great space which has been remarked of the Faunas of earlier formations. “In the equivalent deposits in the Himalaya Mountains, at Fernando Po, in the region north of the Cape of Good Hope, and in the Run of Cutch, and other parts of Hindostan, fossils have been discovered, which, as far as English naturalists who have seen them can determine, are undistinguishable from certain oolite and lias fossils of Europe.” [108a]
The dry land of this age presented cycadeæ, “a beautiful class of plants between the palms and conifers, having a tall, straight trunk, terminating in a magnificent crown of foliage.” [108b] There were tree ferns, but in smaller proportion than in former ages; also equisetaceæ, lilia, and conifers. The vegetation was generally analogous to that of the Cape of Good Hope and Australia, which seems to argue a climate (we must remember, a universal climate) between the tropical and temperate. It was, however, sufficiently luxuriant in some instances to produce thin seams of coal, for such are found in the oolite formation of both Yorkshire and Sutherland. The sea, as for ages before, contained algæ, of which, however, only a few species have been preserved to our day. The lower classes of the inhabitants of the ocean were unprecedentedly abundant. The polypiaria were in such abundance as to form whole strata of themselves. The crinoidea and echinites were also extremely numerous. Shell mollusks, in hundreds of new species, occupied the bottoms of the seas of those ages, while of the swimming shell-fish, ammonites and belemnites, there were also many scores of varieties. The belemnite here calls for some particular notice. It commences in the oolite, and terminates in the next formation. It is an elongated, conical shell, terminating in a point, and having, at the larger end, a cavity for the residence of the animal, with a series of air-chambers below. The animal, placed in the upper cavity, could raise or depress itself in the water at pleasure by a pneumatic operation upon the entral air tube pervading its shell. Its tentacula, sent abroad over the summit of the shell, searched the sea for prey. The creature had an ink-bag, with which it could muddle the water around it, to protect itself from more powerful animals, and, strange to say, this has been found so well preserved that an artist has used it in one instance as a paint, wherewith to delineate the belemnite itself.
The crustacea discovered in this formation are less numerous. There are many fishes, some of which (acrodus, psammodus, &c.,) are presumed from remains of their palatal bones, to have been of the gigantic cartilaginous class, now represented by such as the cestraceon. It has been considered by Professor Owen as worthy of notice, that, the cestraceon being an inhabitant of the Australian seas, we have, in both the botany and ichthyology of this period, an analogy to that continent. The pycnodontes, (thick-toothed,) and lepidoides, (having thick scales,) are other families described by M. Agassiz as extensively prevalent. In the shallow waters of the oolitic formation, the ichthyosaurus, plesiosaurus, and other huge saurian carnivora of the preceding age, plied, in increased numbers, their destructive vocation. [110] To them were added new genera, the cetiosaurus, mososaurus, and some others, all of similar character and habits.
Land reptiles abounded, including species of the pterodactyle of the preceding age—tortoises, trionyces, crocodilians—and the pliosaurus, a creature which appears to have formed a link between the plesiosaurus and the crocodile. We know of at least six species of the flying saurian, the pterodactyle, in this formation.
Now, for the first time, we find remains of insects, an order of animals not well calculated for fossil preservation, and which are therefore amongst the rarest of the animal tribes found in rocks, though they are the most numerous of all living families. A single libellula (dragon-fly) was found in the Stonesfield slate, a member of the lower oolitic group quarried near Oxford; and this was for several years the only specimen known to exist so early; but now many species have been found in a corresponding rock at Solenhofen, in Germany. It is remarkable that the remains of insects are found most plentifully near the remains of pterodactyles, to which undoubtedly they served as prey.
The first glimpse of the highest class of the vertebrate sub-kingdom—mammalia—is obtained from the Stonesfield slate, where there has been found the jaw-bone of a quadruped evidently insectivorous, and inferred, from peculiarities in the structure of that small fragment, to have belonged to the marsupial family, (pouched animals). It may be observed, although no specimens of so high a class of animals as mammalia are found earlier, such may nevertheless have existed: the defect may be in our not having found them; but, other things considered, the probability is that heretofore there were no mammifers. It is an interesting circumstance that the first mammifers found should have belonged to the marsupialia, when the place of that order in the scale of creation is considered. In the imperfect structure of their brain, deficient in the organs connecting the two hemispheres—and in the mode of gestation, which is only in small part uterine—this family is clearly a link between the oviparous vertebrata (birds, reptiles, and fishes) and the higher mammifers. This is further established by their possessing a faint development of two canals passing from near the anus to the external surface of the viscera, which are fully possessed in reptiles and fishes, for the purpose of supplying aerated water to the blood circulating in particular vessels, but which are unneeded by mammifers. Such rudiments of organs in certain species which do not require them in any degree, are common in both the animal and vegetable kingdoms, but are always most conspicuous in families approaching in character to those classes to which the full organs are proper. This subject will be more particularly adverted to in the sequel.
The highest part of the oolitic formation presents some phenomena of an unusual and interesting character, which demand special notice. Immediately above the upper oolitic group in Buckinghamshire, in the vicinity of Weymouth, and other situations, there is a thin stratum, usually called by workmen the dirt-bed, which appears, from incontestable evidence, to have been a soil, formed, like soils of the present day, in the course of time, upon a surface which had previously been the bottom of the sea. The dirt-bed contains exuviæ of tropical trees, accumulated through time, as the forest shed its honours on the spot where it grew, and became itself decayed. Near Weymouth there is a piece of this stratum, in which stumps of trees remain rooted, mostly erect or slightly inclined, and from one to three feet high; while trunks of the same forest, also silicified, lie imbedded on the surface of the soil in which they grew.
Above this bed lie those which have been called the Wealden, from their full development in the Weald of Sussex; and these as incontestably argue that the dry land forming the dirt-bed had next afterwards become the area of brackish estuaries, or lakes partially connected with the sea; for the Wealden strata contain exuviæ of fresh-water tribes, besides those of the great saurians and chelonia. The area of this estuary comprehends the whole south-east province of England. A geologist thus confidently narrates the subsequent events: “Much calcareous matter was first deposited [in this estuary], and in it were entombed myriads of shells, apparently analogous to those of the vivipara. Then came a thick envelope of sand, sometimes interstratified with mud; and, finally, muddy matter prevailed. The solid surface beneath the waters would appear to have suffered a long continued and gradual depression, which was as gradually filled, or nearly so, with transported matter; in the end, however, after a depression of several hundred feet, the sea again entered upon the area, not suddenly or violently—for the Wealden rocks pass gradually into the superincumbent cretaceous series—but so quietly, that the mud containing the remains of terrestrial and fresh-water creatures was tranquilly covered up by sands replete with marine exuviæ.” [114] A subsequent depression of the same area, to the depth of at least three hundred fathoms, is believed to have taken place, to admit of the deposition of the cretaceous beds lying above.
From the scattered way in which remains of the larger terrestrial animals occur in the Wealden, and the intermixture of pebbles of the special appearance of those worn in rivers, it is also inferred that the estuary which once covered the south-east part of England was the mouth of a river of that far-descending class of which the Mississippi and Amazon are examples. What part of the earth’s surface presented the dry land through which that and other similar rivers flowed, no one can tell for certain. It has been surmised, that the particular one here spoken of may have flowed from a point not nearer than the site of the present Newfoundland. Professor Philips has suggested, from the analogy of the mineral composition, that anciently elevated coal strata may have composed the dry land from which the sandy matters of these strata were washed. Such a deposit as the Wealden almost necessarily implies a local, not a general condition; yet it has been thought that similar strata and remains exist in the Pays de Bray, near Beauvais. This leads to the supposition that there may have been, in that age, a series of river-receiving estuaries along the border of some such great ocean as the Atlantic, of which that of modern Sussex is only an example.