Very large, heavily-built, bovine Antelopes, differing from the rest of the Tragelaphinæ in the presence of horns in both sexes. Horns longer than the face, arising well behind the orbits and directed backwards in the plane of the nasal bones, massive (in the male) and furnished with a strong but close spiral twist in the basal half; the anterior crest large, making a complete circuit of the horn and reappearing on its anterior surface near the middle when the horn is unworn, and always at some distance from the tip.

Hair on the forehead longer than on the rest of the head, and forming, in old males, a thick and stiff mat; hair on the nape forming a short mane reversed in the direction of the growth, the parting close to the withers. Throat furnished with a flap of loose skin, or dewlap, which bears a beardlike tuft of hairs.

Tail reaching to the hocks, covered with short hair, but tufted at the tip.

Female. Like the male, but slighter in build; without the thick frontal mat of hair; horns longer, thinner, less strongly crested, and usually much less twisted. Mammæ 4.

The two species of this genus may be shortly diagnosed as follows:—

• A. Ears narrow and pointed; neck brown like the body.  132. T. oryx.
• B. Ears large and expanded; neck black, with a white posterior margin.  133. T. derbianus.

132. THE ELAND.
TAUROTRAGUS ORYX (Pall.).
[PLATES XCVIII. & XCIX.]

Subspecies a. Taurotragus oryx tyficus.

Adult male, at the withers, about five feet ten inches or, according to some writers, sometimes considerably over six feet in height. Body, head, legs, and neck of a tolerably uniform tawny colour, but often assuming a slaty-grey hue in old age, owing to the rubbing off of the hair and the consequent exposure of the skin beneath. Frontal mat of hairs varying from yellowish brown to black, apparently becoming darker with age; nose generally ashy black; lips and chin white. Ears narrow and pointed, of a uniform greyish-fawn tint, with at most a small black patch on the lower rim in front. A narrow black spinal stripe, extending from the withers to the base of the tail. Tail-tuft and tuft of hair on the dewlap black. Legs like the body, but, on the inner side, sometimes with a greyish patch above the knee on the pasterns; back of the pasterns and narrow rim above the hoofs and false hoofs black.

Horns about 30 inches or more in length.

Female like the male in colouring, but smaller and more slightly built; the horns thinner, less strongly crested, and less twisted, but usually longer and (exceptionally) reaching a length of 34 inches.

The subspecies T. o. livingstonii is generally similar to the typical form, but has the coat of a ruddier fawn-colour, and is ornamented on each side of the body and hind-quarters with from about eight to eleven narrow vertical white stripes: these are strong upon the flanks and faint upon the haunches; they commence from the black spinal stripe and gradually fade away upon the belly and lower part of the thighs. In the typical form, moreover, there is a large black patch on the inner and posterior side of the fore leg above the knee. Horns reaching about 32 inches.

Female differing from the male in the same respects as in T. o. typicus.

The subspecies T. o. gigas is based on a pair of horns obtained by Heuglin on the White Nile, and distinguished by their large size, great length (35 inches), and strong corrugations. From Schweinfurth’s observations we learn that this form carries well-marked body-stripes throughout life, sometimes 15 in number. In these two respects it would seem to approach Taurotragus derbianus, but Schweinfurth says nothing about the black neck of the last species.

At the close of the long series of Antelopes we arrive at the largest and finest form of the whole group, and one, moreover, that might well become of great economical importance to mankind, if proper measures were taken for its acclimatization.

The “Eland,” as it is now universally called, was well known to the early settlers of the Cape, where it received its name from some fancied resemblance to the Elk (Alces machlis), which is the “Eland” of the Hollanders and the “Elenn” or “Elendthier” of the Germans. It must have been size, we suppose, more than any other point of similarity, that induced the Dutchmen to apply such an unsuitable name to this animal.

The old traveller Peter Kolben, about 1719, gave the first recognizable, though rather misleading, account of the Eland, which at that epoch was still found in the mountains near Capetown. In 1764 Buffon, in the twelfth volume of his ‘Histoire Naturelle,’ called it “Le Coudous,” or, at any rate, gave unmistakable figures of its horns under that name, which, we suppose, he had by some error transposed to it from the Kudu (Strepsiceros capensis). It was mainly upon Kolben’s Alces capensis and Buffon’s “Coudous” that Pallas, in his first essay on the genus Antilope (1776), based his Antilope oryx, alleging that it “seemed to be” the Antilope oryx of ancient authors! At the same time he states that he had examined a complete skeleton of this animal in the Museum of Prince William of Holland. Very unfortunately, in his second and amended list of the Antelopes, Pallas proposed to make a change in his former names by transferring the term “oryx” to another animal (the Antilope bezoartica of his first memoir) and assigning the new name “oreas” to the Eland. This change, however, we may say, has been generally acquiesced in, and the name oreas has been almost universally applied to the Eland, either specifically or generically, until modern days, when the zealous searchers after priority have resuscitated Pallas’s long-forgotten term “oryx.” This, indeed, seems certainly to be the earliest specific name applicable to the present animal and should, in strict justice, be adopted.

As regards its generic name the Eland has been equally unfortunate. Desmarest, in 1822, first proposed to use “Oreas” as a subgeneric term for this form; and Gray, in 1850, employed it as a genus, combining it with the specific term “canna,” so that the name of the Eland became Oreas canna. As will be seen by our list of synonyms, this name was generally adopted, and has been in constant use for the present species for the last twenty years. We have, however, shown that “oreas,” as a specific term, must give place to “oryx”; and in like manner “Oreas” cannot stand as a generic term for the present animal, because it has been previously employed in zoology as a genus of Lepidoptera (1806) and as a genus of Mollusca (1808), both of which antedate Desmarest’s use of it in 1822. Under these circumstances it is necessary to adopt the next given name, Taurotragus of Wagner, and the correct scientific name of the Eland, according to modern usage, will be Taurotragus oryx.

Having now stated at full length our reasons for the unwelcome but necessary change of name of this Antelope, we will resume our comments on its literary history.

In the Supplement to his ‘Histoire Naturelle,’ published in 1782, Buffon was able to give an improved account of the Eland. This was mostly copied from Allamand’s article inserted in Schneider’s edition of the ‘Histoire Naturelle,’ issued at Amsterdam in the previous year, and was accompanied by a perfectly recognizable figure of the whole animal under the name of “Le Canna,” adopted from its supposed Hottentot appellation. Shortly afterwards (1783) Vosmaer, in a number of his ‘Regnum Animale,’ published a full description and coloured figure of the Eland from a specimen then living in the Menagerie of the Prince of Orange in Holland, probably the same as that from which Allamand had taken his information. Sparrman, who visited the Cape about this period, also gave a good account of the structure and habits of this Antelope, as observed by him in the Alexandria and Somerset-East Divisions of the Colony. Paterson, in 1790, recorded having met with Elands in Caledon, as also in the Van-Ryndorp and Uitenhage Divisions a few years previously. Thunberg, another well-known traveller and naturalist (1795), found the Eland in Uniondale, and Lichtenstein in Calvinia, Aberdeen, and Middelburg (1803–4). Our countryman Burchell, as recorded in his ‘Travels,’ came across Elands in 1822, in Prieska, Herbert, and Britstown, and found them numerous in Hanover.

We may now pass on to the days of Harris, whose celebrated hunting-expedition into the interior took place in 1836 and 1837. Even at that date the Eland was pronounced to be extinct in the Cape Colony, but was met with in abundance on the banks of the Vaal River, where Harris feasted himself and his followers on its succulent meat.

By all classes in Africa, Harris writes, the flesh of the Eland is deservedly esteemed over that of any other animal:

It is right, however, to mention that other experienced authorities do not altogether agree with Harris’s pronouncement on this subject. Mr. Selous, for example, states his opinion that the flesh of the Eland has been “very much over-estimated,” and is “not to be compared in flavour with that of the Buffalo, Giraffe, Hippopotamus, and White Rhinoceros.” (De gustibus non est disputandum!)

Harris describes the favourite haunts of the Eland on the Vaal River in his days as follows:—

Since Harris issued his work in 1840 all the writers on the game-animals of Southern Africa have devoted more or less space to the Eland. Delagorgue, who published his travels in 1847, found this Antelope in plenty in Zululand. Methuen, in his ‘Life in the Wilderness’ (1848), describes its habits in the Kalahari Desert, and Livingstone (1857) alludes to the Eland as being able to exist without water, and states that one may see hundreds of them in places thirty or forty miles distant from that element. The Hon. W. H. Drummond, in his ‘Rough Notes on the Large Game of South Africa,’ has devoted a whole chapter to the pursuit of the Eland, which he met with on the Black and White Umvalosi Rivers, and in other districts, but not within the Colony itself, in which, according to Bryden, it became extinct between 1840 and 1850, having probably lingered longer in the waterless deserts of Bushman’s Land than in any other locality.

Finally, Mr. H. A. Bryden, writing in 1897, in his ‘Nature and Sport in South Africa,’ on the rapidly disappearing forms of South-African game, laments the noblest of all the Antelopes of the world as taking the lead in this sad progress. At the present time, he says, one must go far north into the parched and pathless recesses of the Upper Kalahari before the “vanishing Eland” can be reached, and “even in these unexplored wilds these rare creatures can nowadays be scarcely considered safe.” Mr. Bryden proceeds to describe the progress of its extermination now going on as follows:—

Thus we see that the typical brown unstriped Eland, which formerly pervaded the whole of the Cape Colony and the adjacent districts, and in 1652 (according to Van Riebeck) was found even on Table Mountain, is now, as nearly as possible, extinct; although its closely-allied white-striped brother, called Livingstone’s Eland, after the distinguished explorer and missionary, is still to be met with in the countries further north. As regards the points of difference between Livingstone’s Eland and the typical form, which we will now proceed to explain, we cannot do better than quote from Mr. Selous’s excellent article on the subject lately published in Mr. Rowland Ward’s ‘Great and Small Game of South Africa’:—

As these striped Elands grow up, Mr. Selous continues, they differ considerably one from another. Both bull and cow become of a bluish grey as their coats become thinner with age, and at a little distance the white stripes are often indistinguishable, although as long as there is any hair left they can always be seen on close inspection. Also the dark patches on the inner sides of the front legs become more faint with age, and in very old animals disappear altogether.

Mr. Selous also points out that intermediate forms are found between the two subspecies of Elands, and that, in fact, there is a complete passage through a long series of variations from one form to the other.

That this is the case is shown clearly by Mr. Selous’s own observations. He writes (in the same work) as follows:—

Mr. Bryden and other well-known authorities on the game-animals of South Africa entirely confirm Mr. Selous’s observations.

Under these circumstances it seems quite impossible to treat Livingstone’s Eland as a distinct species, but, as will be seen by our list of synonyms, we have placed it under a different heading, and have assigned most of the references to Elands north of the Zambesi to Taurotragus oryx livingstonii.

We will now, starting from the Zambesi, endeavour to trace the Eland into the most northern part of its range.

Peters, in his ‘Reise nach Mossambique,’ gives several localities for the Eland in the Portuguese Provinces on the Zambesi. Mr. R. Crawshay, who has devoted great attention to the Antelopes of Nyasaland, tells us that the Eland is widely distributed there, both on the hills and on the wooded plains at the foot of them, and gives many localities in which they are to be met with. As regards its markings, he informs us that it is subject to great variety in British East Africa, “both in colour and as regards the plainness or otherwise of the white stripes.” In a single troop, individuals may be seen varying from a light tawny yellow to the slaty blue of old age, while in some the stripes are clearly defined, in others faintly, and in others again they are not distinguishable at all.

In 1895 Sir Harry Johnston, K.C.B., F.Z.S., presented to the Zoological Society of London a remarkably fine pair of horns of Livingstone’s Eland, which are now suspended in their meeting-room at Hanover Square. The animal which carried them was shot in 1893 in Nyasaland between Zomba and Lake Chilwa. By the kind favour of that Society we are able to insert in these pages a copy of the figure of these horns (fig. 116, p. 205), which was published in the ‘Proceedings’ for 1895.

When we go further north into German East Africa, the Eland appears to be not quite so abundant, although Herr Matschie mentions it as found in Usagara, and it was seen by Neumann between the Pangani River and Irangi during his recent journey, besides having been met with in Ugogo by Speke and Grant in former years.

In British East Africa it would appear to be more prevalent again, although somewhat local. Mr. Jackson, in his volume on ‘Big Game,’ writes as follows concerning its range and habits in that country:—

Mr. S. L. Hinde has kindly favoured us with the following field-notes of his recent experience of the Eland in British East Africa:—

Count Teleki, as we are informed by Herr v. Höhnel in his narrative of the first expedition to Lake Rudolf, met with the Eland on the Likipia plateau, north of Mount Kenya, where, according to a letter addressed by v. Höhnel to Sclater, they encountered a herd of about 170. But we are not aware of any evidence of its being found further north in British East Africa or in any part of Somaliland. Here, therefore, we appear to have reached its furthest limits in this direction, but further west, in the Valley of the Nile, there is good evidence of the existence of this form of Antelope in much higher latitudes.

The famous explorer Baron von Heuglin was the first traveller who recorded the existence of an Eland in the districts of the Upper Nile, although v. Pruyssenaer (as Heuglin states) had previously recognized its occurrence on the Bahr el-Abiad and Bahr-el-Sobat. But Heuglin, having obtained a pair of Eland’s horns from the Upper White Nile, about 7° N. lat., referred them, on account of their large size, to a new species, “Boselaphus gigas.” He gives a figure of these horns and states that they measure 35 inches in length, and show a distance of 32 inches between the two points. We have thought it advisable to reproduce Heuglin’s figure of this remarkable pair of horns (see fig. 117, p. 208). In a subsequent work (‘Reise in das Geb. d. Weiss. Nil’) Heuglin adds that his Taurotragus gigas is found in pairs and singly in the forests of the Djur River and amongst the Arol negros.

The well-known traveller Schweinfurth, in his ’Im Herzen von Afrika,’ also alludes more than once to the existence of the Eland on the upper confluents of the White Nile. In the first place, he met with it in Bongoland, where he says that it resorts to the drier slopes of the hills during the rains, and descends to the valleys in the winter months. In the second volume of his narrative, Schweinfurth mentions it again, and gives two figures of the horns of what he calls the Central-African Eland, of two very different forms. Schweinfurth states that the skin is plainly striped, and that this is certainly no mark of youth, because he has seen very old examples which had about fifteen narrow parallel stripes, about a finger in breadth, on both sides. It is quite evident, therefore, that the Eland of this part of Africa belongs to the striped form. It may be identical with T. o. livingstonii, but, as Heuglin has given it a name, we will allow him the benefit of the doubt for the present, and will call this northern striped form Taurotragus oryx gigas until further investigations have been made.

Before closing our systematic account of the Eland we must say a few words respecting some curious horns which were at first ascribed to a new and unknown Antelope, but are now generally admitted to be nothing more than abnormal horns of the cow Eland. These horns were first brought before the scientific world by Dr. Günther, who exhibited a pair of them at a meeting of the Zoological Society in 1889, and stated that they had been obtained on the frontiers of Natal. Dr. Günther’s opinion was that they belonged to an unknown Antelope of the Tragelaphine group, but under the uncertainty as to what form they were most nearly allied, he proposed to designate the presumed species Antilope triangularis. Through the courtesy of the Zoological Society of London, and with Dr. Günther’s kind permission, we are able to reproduce here the illustration of these horns (fig. 118, p. 209) which accompanied Dr. Günther’s paper in the Society’s ‘Proceedings.’

Writing of these horns in 1891, Mr. Lydekker was so convinced of their essential difference from those of any other known Antelope that he proposed to raise the animal that bore them to generic rank under the name “Doratoceros.”

Some years subsequently, in 1896, Sclater obtained, on loan, a fine pair of horns of nearly similar character from Mr. Justice Hopley, of Kimberley, and, after comparing them with the typical pair of Antilope triangularis in the British Museum, came to the conclusion that they must have belonged to the same species of Antelope. Mr. Justice Hopley’s pair were not quite so long, rather more incurved backwards, and less broadly spread; they were also smoother at the base, showing but slight traces of corrugations. When exhibiting these horns to the Zoological Society, Mr. Sclater stated that he could see nothing whatever to negative the opinion, already prevailing amongst other naturalists, that these horns were abnormal horns of the cow Eland, which had grown into a lengthened form without making the ordinary twist usually observable in that species and in other Tragelaphs. It is right to add that Mr. Lydekker himself is now also of the same opinion, and has stated (‘Horns and Hoofs,’ p. 260) that these horns “are almost certainly abnormal specimens of those of a cow Eland.”

As we have already stated, living examples of the Eland were received in Holland from the Cape as long ago as about 1783, when they were described by Vosmaer and others as being in the menagerie of the Prince of Orange. In England the first examples of this species of which we can find any record were those which constituted the herd in the celebrated menagerie of Edward, 13th Earl of Derby, President of the Zoological Society of London. There is, unfortunately, little information available as to the origin and history of this celebrated herd, but from some notes published by Lord Derby in the first volume of the ‘Gleanings,’ we learn that the first specimens received were obtained for him by Mr. Burke from the Cape in November 1842, and consisted of two males and a female. The female first bred in August 1843, and produced young in 1844, 1845, and 1846, at which date Lord Derby remarked that he had in his possession four males and two females of this Antelope. At the dispersal of the Derby Menagerie by auction in October 1851 the Knowsley herd consisted of two males and three females. These passed into the Gardens of the Zoological Society of London, having been selected out of the whole stock by the Council of the Society in virtue of a bequest by Lord Derby to that Society of any group of animals in his collection that they might prefer.

The original stock of the Zoological Society’s herd of Elands consisted, therefore, of these five animals received by the Society in December 1851. Of these an old female had been born at Knowsley in 1846, and the other individuals, two males and two females, had been imported by Lord Derby in 1850. These animals throve well in their new quarters and began to increase rapidly. As will be seen by the list given in Wolf and Sclater’s ‘Zoological Sketches’ (vol. i.), two calves were born in 1853, three in 1854, four in 1855, and four in 1856. The first additions made to the original stock were a female presented by the late Sir George Grey in April 1859, and a male received in exchange from Viscount Hill in November of the same year. Since the date of its first institution, the Zoological Society’s herd of Elands has never failed, although occasionally reduced to somewhat small dimensions. Nearly every year one or more Eland-calves have been born in the Gardens, and care has been taken to lose no opportunity of introducing fresh blood whenever the occasion has offered. At the present moment, however, we regret to say, in consequence of the great difficulties now prevailing in obtaining living examples of the larger Antelopes of Africa, the Eland is represented in the Society’s Antelope-House by only two specimens, namely, a male, about six years old, bred in the Jardin d’Acclimatation of Paris, and received on October 12, 1898, and a young female, purchased of Herr Reiche, of Alfeld, in April 1899. The latter is more rufous in colouring and shows slight traces of stripes, which, however, she may probably lose when quite adult.

From these two specimens our illustration of Taurotragus oryx typicus (Plate XCVIII.) has been prepared by Mr. Smit.

Besides the Zoological Society’s animals, the only herd of Elands that we are aware of now existing in this country is that belonging to the Duke of Bedford, the President of the Zoological Society of London, which is kept in the beautiful Park at Woburn, along with a splendid series of Deer and other Ungulates. Through the kindness of His Grace we have been furnished with the following particulars concerning this herd, which now consists of fourteen individuals. Three of these are adult females, two of which were purchased from dealers, and the third from the Zoological Society of London, in whose Gardens it was bred. The adult male was purchased of Herr Reiche, of Alfeld. Five young males and two females have been bred at Woburn up to the end of 1899. Three calves, one male and two females, have been born at Woburn since the commencement of the present year.

Allusion has already been made to the Elands possessed by the late Roland, Viscount Hill, who, about the year 1861, possessed a fine herd of these animals. When visited by Sclater about that date, Lord Hill’s stock consisted of three males and seven females, which were kept at his Lordship’s residence Hawkstone, in Shropshire. They were the produce of individuals principally purchased by him from the Zoological Society, and were kept in grazing paddocks in Hawkstone Park. Unfortunately, a few years later Lord Hill lost his interest in these animals and got rid of them.

About the same period John, 2nd Marquis of Breadalbane, likewise purchased a herd of Elands, which, however, we believe, was not maintained long after the Marquis’s death in 1862.

In almost all the Zoological Gardens of the Continent also the Eland is a well-known object of interest, and in many of them, until the last few years, has thriven well and produced its kind; but, as already mentioned, the supply of Elands from abroad has recently much decreased, and at the present time there is a great difficulty in keeping our herds of Elands in Europe up to the mark by the necessary introduction of fresh blood.

One of the chief ornaments of the Mammal-Gallery in the British Museum is the mounted pair of Livingstone’s Elands obtained by Mr. F. C. Selous in Mashonaland in 1883. The male (as Mr. Selous informs us) was shot near Sadza’s Kraal, west of Marandalla’s, a station on the main road from Salisbury to Umtali, in July of that year, and the female near Salisbury in the following October. The male stands 67¾ inches high at the withers, and carries a pair of horns 22½ inches in length in a straight line; the female is 57½ inches in height, and has horns 27 inches in length. In both these animals the lateral stripes are well defined, and there are no black patches above the knee on the fore leg of the male, though in the female the patches are slightly visible. These specimens are fair representatives of Taurotragus oryx livingstonii, and have been figured as such in our illustration (Plate XCIX.), prepared by Smit. But it is right to add that it appears that the skins have apparently shrunk slightly in drying, as in his measurements of the male specimen in question, lately given in the ‘Great and Small Game of Africa’ (p. 426), Mr. Selous states that the height of this animal, “taken on the naked carcase after the skin had been removed,” was 69 inches. Moreover, in former days there were probably still larger specimens, as such reliable authorities as Barrow and Harris agree in stating that the old male Elands were known to attain a height of 6½ feet at the withers.

There are also in the National Collection other skins and skulls of the Eland obtained by Mr. Selous, and a number of other specimens, amongst which we may specify a skull and horns of a female from Nyasaland, presented by Sir Harry Johnston, and a skull and horns from the district of Kilimanjaro, presented by Mr. F. J. Jackson in 1892. We may remark that examples of the Eland of the White Nile (the problematical T. o. gigas) are much wanted to complete the series in the National Collection, besides which specimens from other definite localities in Eastern Africa and Angola would be very acceptable.

April, 1900.