Size large; form high and comparatively slender, less stout and bovine than in the succeeding genera. Muzzle hairy. Tail long and tufted. Mammæ 4. Large accessory hoofs present.

Skull proportionally long, its frontal region very convex upwards, the large horn-cores rising almost vertically above the posterior half of the orbits. Lachrymal fissures almost or quite obsolete. No anteorbital fossa. Premaxillæ not reaching to the nasals.

Horns medium or long, slightly but evenly divergent, nearly vertical basally, strongly curved backwards above; heavily ringed.

Female with horns similar to those of the male, but shorter, slenderer, and much smoother.

Of this genus, which contains some of the handsomest Antelopes in existence, we recognize three species, one of them (alas!) now extinct. The widely distributed Roan Antelope may be provisionally separated into four local subspecies, which require further elucidation.

• A. General colour greyish or pale brown. Horns not or but little longer than the head.
• a. Size smaller. Face without black markings.  110. H. leucophæus.
• b. Size larger. Face with strongly contrasted black and white markings.  111. H. equinus.
• a¹. General colour greyish; ears shorter. (S. Africa.)  Subsp. typica.
• b¹. General colour pale rufous; ears shorter. (E. Africa.)  Subsp. rufo-pallida.
• c¹. General colour browner; ears longer. (N.E. Africa.)  Subsp. bakeri.
• d¹. General colour more fulvous; ears still longer. (W. Africa.)  Subsp. gambiana.
• B. General colour black. Horns much longer than the head.  112. H. niger.

110. THE BLUE-BUCK.
HIPPOTRAGUS LEUCOPHÆUS (Pall.).
[PLATE LXXVI.]

Vernacular Name:—Blawe-bock of Dutch (Le Vaillant).

Size much less than in the two following species; height at withers from 45 inches (♂ in Paris) to 40 inches (♀ in Vienna). General colour bluish grey. Forehead brown; upper lip and a patch in front of the eye lighter than the general colour, but there are none of the marked black and white contrasts so prominent in H. equinus. Ears not so long or so pointed as in H. equinus, and without black tufts at their tips. Mane on nape of neck short, inconspicuous, directed forward; throat-mane almost or quite absent. Belly dull whitish, not contrasted with the sides. Limbs with an inconspicuous darker line down their anterior surfaces. Tail-tuft greyish, but little darker than the general colour.

Skull probably merely differing from that of H. equinus by its smaller size, but, so far as is known, no museum possesses an example of it.

Horns like those of H. equinus, but much smaller and more slender; perhaps rather longer in proportion to the size of the animal. Those of the Paris specimen (a male) measure 21½ inches in length round the curve and have 28 rings upon them. The pair in the British Museum are rather shorter.

The Blue-buck, like the Quagga (Equus quagga), belongs to the category of larger animals that have become extinct within the historic period. While the Square-lipped Rhinoceros (Rhinoceros simus) and the Mountain Zebra (Equus zebra) are still occasionally to be met with in one or two remote districts of South Africa, it would seem that the Blue-buck and the Quagga, as living creatures, have utterly perished from the face of the earth, and are only now represented by a few specimens in some of the principal museums of Europe.

Although the “Blaauwe-bok” or “Blau-bok” was long known to early travellers at the Cape under its vernacular names, the great Russian naturalist Pallas was the first to register it definitely in the ‘Annals of Science.’ Under the name Antilope leucophæa Pallas described it in 1766, in his ‘Miscellanea Zoologica,’ from a specimen in the Leyden Museum[1], and placed it as the first species of his genus Antilope. We have already, however (Book of Ant. III. p. 3), given the reasons why another species—the Black-buck of India—should be deemed to be the type of Pallas’s generic term Antilope, and in accordance with ordinary usage we employ Sundevall’s name Hippotragus for the present species and its allies.

The next author after Pallas to mention this Antelope appears to have been Allamand, who made various contributions to an edition of Buffon’s ‘Histoire Naturelle,’ issued in Holland by Schneider in 1766 and the following years. Allamand, however, carelessly confounded this South-African Antelope with the Gazella gutturosa of Siberia, of which the native name is Tzeiran or Dzéren (see Book of Ant. III. p. 84), and adopted the same name for it. Allamand’s figure of his “Tzeiran” was taken from a mounted specimen in the Cabinet d’Histoire Naturelle of J. C. Sylvius von Lennep, of Harlem, which on the death of the owner had passed by bequest to the Société Hollandaise des Sciences of that city. This specimen, when in skin, had been obtained from a dealer who did not know whence it came, but from the mode of its preparation it was believed by Allamand to have been brought from the Cape.

Our countryman Pennant, in his ‘History of Quadrupeds,’ of which the first edition was published in 1781, gave a third original description of this species, which he called the “Blue Antelope.” It was taken, he tells us, from a skin bought at Amsterdam, and said to have been obtained from the Cape of Good Hope. Pennant fully recognized its identity with Buffon’s “Tzeiran,” and remarked on the use of this erroneous Asiatic name for it.

Sparrman, who arrived at the Cape in 1772 and subsequently made a long journey into the interior, tells us in his narrative[2] of having found a skin of the Blue-buck (which he identified with Pallas’s A. leucophæa and Pennant’s “Blue Antelope”) preserved at “Krakeel-rivier,” but he does not appear to have met with it alive himself.

But a rather later explorer of the Cape Colony, the well-known French naturalist François Le Vaillant, author of the ‘Oiseaux d’Afrique’ and many other ornithological works, was more fortunate. As Le Vaillant appears to have been the only traveller known to have himself obtained an example of this now extinct species, we will extract from the first volume of his ‘Voyage d’Afrique’ what he has written on this subject.

On December 18th, 1781, Le Vaillant left Capetown to explore the country to the east called “Hottentot Holland.” A few days later, after crossing the river Sonder-end, and passing through the valley of Soete-Melck, he arrived at a place called “Tiger-Hoek,” where he had appointed some Hottentots to meet him, and went on the chase in their company:—

After Le Vaillant’s time little further addition was made to the history of this Antelope until the publication (in 1811 and 1812) of Lichtenstein’s ‘Travels in Southern Africa,’ in which several allusions to it will be found. In the first of these Lichtenstein, on the way from Swellendam to Algoa Bay in December 1803, tells us that much game—Antelopes and Zebras—was met with in the mountains near the Buffalo-jagt River, “but the beautiful Blau-bok (Antilope leucophæa) is, as Barrow has correctly supposed, almost exterminated. In the year 1800 one was shot, of which the skin is now at Leyden, but since then no more have been seen.” In the second volume of his ‘Reise,’ when on the Dweika, between Stellenbosch and Graaf Reinet, in the following December, Lichtenstein informs us that game was plentiful in the inner valleys of the mountains, and continues:—“Here are still found the Zebra, the Bontebok, and the Reh-bok in comparative abundance, and even the Blau-bok (Antilope leucophæa, which is almost exterminated elsewhere, is said to occur occasionally.” In his celebrated article upon the genus Antilope, published in the ‘Magazin der Gesellschaft naturforschender Freunde zu Berlin,’ two years subsequently, Lichtenstein, after a description of this species from the specimen at Berlin, continues as follows:—“The skin which I describe is, unfortunately, the last that has been seen. Since 1799, when this specimen was shot, no more have been met with, and it is known that this Antelope was found only in the now well-populated district of Zwellendam, and nowhere else. Apparently this beautiful animal is now quite extinct.”

The animal having been thus exterminated towards the end of the last century, its very existence became a matter of doubt to some naturalists, who were inclined to consider the specimens of it left in our museums as small or immature individuals of the nearly allied Roan Antelope (H. equinus). This view was taken by Andrew Smith[3], de Blainville, Gray, and even Harris, who, one would think, might have learned better from the traditions on the subject prevalent among the Boers. But the accurate Sundevall was strongly against this opinion, and, after examining the specimens at Upsala, Stockholm, and Paris, said decidedly “Minimè animal fictum, ut credidit A. Smith.” Sundevall, however, failed to convert Gray on this subject, and Gray, although, as he tells us, he had examined the specimen at Paris, chose to unite this species to the Roan Antelope, and to call them both Hippotragus leucophæus.

The most recent authority to vindicate the claims of the Blue-buck to specific distinctness is Herr F. F. Kohl, of Vienna, who, in an article upon new and rare Antelopes in the Imperial Natural History Museum, published in 1866, after accurately describing the specimen in that collection and pointing out its distinctive characters from H. equinus, gave a full list of the various synonyms to be allocated to these two species.

There can be little doubt, therefore, that Hippotragus leucophæus must be regarded as an extinct animal, of which at the present time five mounted specimens only are known to exist. All these we have already alluded to, but we may repeat that they are to be found in the Museums of Paris, Leyden, Vienna, Stockholm, and Upsala.

Finally, however, we are glad to be able to add that, although our National Collection does not contain a complete example of this species, yet it possesses a frontlet and horns which, after careful comparison, we have no hesitation in referring to H. leucophæus. The horns (fig. 88, p. 11) are just 20 inches in length and 6·1 in basal circumference; they have the characteristic ridging and curvature of the horns of the male, and are obviously adult, but their size is less than the horns of the female Roan Antelope. The frontlet on which they are borne measures 3·85 inches between the orbits. The exact origin of this frontlet is not known, but it has been long in the Museum.

Our illustration of the Blue-buck (Plate LXXVI.) was put upon stone by Mr. Smit many years ago, from a water-colour sketch by Mr. Wolf, which is now before us. This sketch was drawn by Mr. Wolf under Sir Victor Brooke’s directions, probably from the specimen at Paris, which we believe Sir Victor examined more than once, but we regret to say that there is no certainty on this point. It should be mentioned, however, that the elongation of the hairs on the neck shown in the Plate is probably rather exaggerated, as this species, we are told by Sundevall, had only a very short “neck-mane.”

January, 1899.

111. THE ROAN ANTELOPE.
HIPPOTRAGUS EQUINUS (DESM.).
[PLATES LXXVII. & LXXVIIL]

Subspecies a. H. e. typicus.

Subspecies b. H. e. rufo-pallidus.

Subspecies c. H. e. bakeri.

Subspecies d. H. e. gambianus.

Size very large, an adult male standing 56 inches high at the withers. General colour greyish, browner in the two northern subspecies. Top and sides of face black, contrasting markedly with the white muzzle and lips and with a prominent patch just in front of the eyes. On the lower half of this patch the hairs are elongated into a brush. Behind the eyes a second less conspicuous white patch is present. The black, however, is only developed in the adult, young specimens having the face nearly uniform with the body. Ears very long, narrow, pointed, their tips pencilled with black. Mane well developed, brown, directed backward, except just on the withers, where there is a tendency for it to be whorled. Throat-mane long and prominent. Belly white, its definition laterally rather variable. Limbs brownish fawn, black patches occasionally present on the outer sides of the shoulders and forearms. Tail reaching to the hocks, its brush black.

Skull-dimensions of an adult male (of subspecies H. e. bakeri):—Basal length 16 inches, greatest breadth 6·75, muzzle to orbit 10·3.

Horns stout and strong, cylindrical, heavily ridged, evenly divergent, curved backward; comparatively short for the size of the animal, good specimens being only from 26 to 30 inches in length, and the longest recorded only 33.

Female similar to the male, but the horns more slender, smoother, less heavily ridged, and less strongly curved backward, and neck and body less robust.

It is a well-known and generally accepted fact amongst naturalists that animals which have a wide distribution have also a special tendency to vary, and that if specimens of them from different parts of their ranges are compared, such specimens are usually found not to agree exactly, but to be distinguishable by differential characters more or less evident. When these characters are easily observable and definable their possessors are usually referred to different species, which are supposed to “represent” one another in their respective areas, and are hence often called “representative species.” When the distinguishing characters are slight and less easily recognizable it has recently become the practice, especially among American naturalists, to designate their possessors as “subspecies,” and, in order to indicate this, to add a third “subspecific” name to the ordinary generic and specific terms. This plan we have already adopted in some cases in the present work. But there are many cases in which, either from imperfect evidence or from an insufficient supply of specimens, it is very difficult to decide whether a “local form,” as it may be termed, is better treated of as a species or as a subspecies. And in the present instance we have one of these cases before us. The Roan Antelope is very widely distributed in Africa. From the Cape Colony it extends all up the eastern side of the continent to British East Africa and Sennaar, and is also found on the west coast in Senegal, Togoland, Nigeria, and Angola. Specimens from all these countries present a very general resemblance, and have been considered by most authorities to be identical. On the contrary, other writers have regarded the local forms as distinct, and have separated them under different specific names. We confess that we have not been able (mainly, no doubt, from lack of sufficient specimens to consult) to come to a satisfactory conclusion on this subject; but, for the present, we think it a more prudent course to treat the local forms of this species found in the different districts of Africa as only of subspecific rank, and to class them all under the one specific head as Hippotragus equinus.

The Roan Antelope received its specific name as long ago as 1804, when a short description of it was published by Desmarest in the twenty-fourth volume of the first edition of the ‘Dictionnaire d’Histoire Naturelle,’ taken from a specimen in the Paris Museum. Desmarest designated it by the French name “Antilope Osanne,” but added Geoffrey’s MS. scientific name “Antilope equina” which must, therefore, be attributed to the former author, as having first published it. Desmarest states that the exact locality of this specimen was unknown, but we think it may be safely assumed to have been from the Cape. Desmarest’s description is not very accurate, but Desmoulins, who wrote the article “Antilope” in the subsequently issued ‘Dictionnaire Classique d’Histoire Naturelle,’ added a figure of the head of Geoffrey’s type, which seems to prove that it could have been of no other than the present species.

The first European explorer in South Africa to meet with the Roan Antelope in its native wilds appears to have been Samuel Daniell, who visited the Cape about the commencement of the present century under the patronage of Lieut.-General Francis Dundas, at that time Acting Governor. In his ‘African Scenery and Animals’ (of which the original folio series was issued in parts in 1804 and the following years) Daniell figured what was, there is little doubt, an example of this Antelope under the name of “The Tackhaitse” (no. 24), and informs us, in the accompanying letterpress, that he met with two of these animals near Latakoo (or Kuruman) in Bechuanaland, where “they are usually found grazing on the edge of the Karroo Plains near the foot of the hills in small herds of five or six.” Upon Daniell’s “Tackhaitse” Schinz founded his Capra œthiopica, Goldfuss his Capra barbata, and Fischer his Antilope truteri; but all these names are happily subsequent in date to the specific term usually adopted for this Antelope, and need not concern us further.

After Daniell the next traveller to meet with the Roan Antelope appears to have been Dr. Burchell, who was at the Cape from 1811 to 1815. In his ‘List of Quadrupeds presented to the British Museum,’ as part of the results of this memorable expedition, Burchell records a male of Antilope equina, “shot at the Little Klibbolikhónni Fountain in the Transgariepine” (now Orange Free State) in December 1812. In Hamilton Smith’s fourth volume of the Mammals of Griffith’s ‘Animal Kingdom’ a full description is given of this specimen (of which a pair of horns now alone remains in the National Collection), accompanied by a good uncoloured figure of it drawn by Thomas Landseer.

Sir Andrew Smith, whose journeys in the Cape districts took place from 1834 to 1836, published a coloured figure of this Antelope in 1840, in his ‘Illustrations of the Zoology of South Africa,’ and gives us the following account of its distribution in those days:—

The well-known sportsman and naturalist Sir William Cornwallis Harris, whose expedition through the interior of the Colony up to the Tropic of Capricorn took place in 1836 and 1837, writes in his usual charming style of this favourite object of the hunter’s pursuit[4]:—

The Roan Antelope appears never to have existed south of the Orange River, and in more recent days, we fear, has retired much further into the interior than the localities specified by Andrew Smith and Cornwallis Harris. Messrs. Nicolls and Eglinton, in their ‘Sportsman in South Africa,’ tell us that it is “now very rarely found on the upper and lower banks of the Botletle River about the Mababé Flats, Great Makari-kari Salt-pans, and Chobe districts, while in the less frequented portions of Matabeleland it is still fairly common, and although once numerous in Mashonaland, is now only to be found there in the low country towards the east coast.” Mr. Selous also states that it is “tolerably plentiful” in parts of Mashonaland, and that he found a good many in the Manica country, north of the Zambesi. Mr. W. L. Sclater informs us that on the western side of South Africa it is still to be found in plenty in Damaraland and Ovampoland.

In the Transvaal, Dr. Percy Rendall, writing in 1895, states that a few of these fine animals were still to be found on the Oliphants River. Herr Reiche, of Alfeld, informs us that, in his yearly importations of animals from the Transvaal since 1887, he has received no less than eight living examples of this Antelope, which have been disposed of to various Continental Gardens. These, however, may have been obtained in the adjoining Portuguese border-country to the north of that Republic. But Mr. Kirby, in his ‘Haunts of Wild Game,’ tells us that, “although very rare,” the Roan Antelope is still to be found in the north-eastern parts of the Transvaal. “There are a few on the high stony ridges across the Mehlamhali and about Maripi’s Berg and the Oliphants River, but nowhere in large numbers.” In 1891 he shot two fine bulls on the Nuanetsi, but they were wanderers.

It was across the Limpopo within the borders of Mashonaland that Mr. John Millais came across this beautiful Antelope in 1893. Its head forms one of the subjects of the cover of his enchanting volume ‘A Breath from the Veldt,’ the pages of which contain several excellent sketches of this splendid animal and much information on its habits. It was near Eland’s Fontein, between the Rivers Bubye and Nuanetsi, that Mr. Millais obtained his first specimen of the Roan Antelope, of which he writes as follows:—

Again, Mr. Millais writes:—