The second ancestral stage of Man, as of all the higher animals and plants, is formed by a simple cell, that is, a little piece of protoplasm enclosing a kernel. There still exist large numbers of similar “single-celled organisms.” Among them the common, simple Amœbæ (vol. i. p. 188, Fig. 2) cannot have been essentially different from these progenitors. The form value of every Amœba is essentially the same as that still possessed by the egg of Man, and by the egg of all other animals. (Vol. i. p. 189, Fig. 3.) The naked egg-cells of Sponges, which creep about exactly like Amœbæ, cannot be distinguished from them. The egg-cell of Man, which like that of most other animals is surrounded by a membrane, resembles an enclosed Amœba. The first single-celled animals of this kind arose out of Monera by the differentiation of the inner kernel and the external protoplasm; they lived in the earlier Primordial period. An irrefutable proof that such single-celled primæval animals really existed as the direct ancestors of Man, is furnished according to the fundamental law of biogeny (vol. i. p. 309) by the fact that the human egg is nothing more than a simple cell. (Compare p. 124.)

Third Stage: Synamœbæ.

In order to form an approximate conception of the organisation of those ancestors of Man which first developed out of the single-celled Primæval animals, it is necessary to trace the changes undergone by the human egg in the beginning of its individual development. It is just here that ontogeny guides us with the greatest certainty on to the track of phylogeny. We have already seen that the egg of Man (in the same way as that of all other Mammals), after fructification has taken place, falls by self-division into a mass of simple and equi-formal Amœba-like cells (vol. i. p. 190, Fig. 4 D). All these divided globules are at first exactly like one another, naked cells containing a kernel, but without covering; in many animals they show movements like those of the Amœbæ. This ontogenetic stage of development which we called Morula (p. 125), on account of its mulberry shape, is a certain proof that in the early primordial period there existed ancestors of man which possessed the form value of a mass of homogeneous, loosely connected cells. They may be called a community of Amœbæ (Synamœbæ). (Compare p. 127.) They originated out of the single-celled Primæval animals of the second stage by repeated self-division and by the permanent union of the products of this division.

Fourth Stage: Ciliated Larva (Planæada).

In the course of the ontogenesis of most of the lower animals, and also in that of the lowest Vertebrate animals, the Lanceolate Animals, or Amphioxus, there first develops out of the Morula (Frontispiece, Fig. 3) a ciliated larva (planula). Those cells, lying on the surface of the homogeneous mass of cells, extend hair-like processes, or fringes of hairs, which by striking against the water keep the whole body rotating. The round many-celled body thus becomes differentiated, in that the external cells covered with cilia differ from the non-ciliated internal cells (Frontispiece, Fig. 4). In Man and in all other Vertebrate animals (with the exception of the Amphioxus), as well as in all Arthropoda, this stage of the ciliated larva has been lost, in the course of time, by abbreviated inheritance. There must, however, have existed ancestors of Man in the early Primordial period which possessed the form value of these ciliated larvæ (Planæa, p. 125). A certain proof of this is furnished by the Amphioxus, which is on the one hand related by blood to Man, but on the other has retained down to the present day the stage of the planula.

Fifth Stage: Primæval Stomach Animals (Gastræada).

In the course of the individual development of Amphioxus, as well as in the most different lower animals, there first arises out of the planula the extremely important form of larva which we have named stomach larva, or gastrula (p. 126; Frontispiece, Fig. 5, 6). According to the fundamental law of biogeny this gastrula proves the former existence of an independent form of primæval animal of the same structure, and this we have named primæval stomach animal, or Gastræa (pp. 127, 128). These Gastræada must have existed during the older Primordial period, and they must have also included the ancestors of man. A certain proof of this is furnished by the Amphioxus, which in spite of its blood relationship to Man still passes through the stage of the gastrula with a simple intestine and a double intestinal wall. (Compare Plate X. Fig. B 4.)

Sixth Stage: Gliding Worms (Turbellaria).

The human ancestors of the sixth stage which originated out of the Gastræada of the fifth stage, were low worms, which, of all the forms of worms known to us, were most closely allied to the Gliding Worms, or Turbellaria, or at least upon the whole possessed their form value. Like the Turbellaria of the present day, the whole surface of their body was covered with cilia, and they possessed a simple body of an oval shape, entirely without appendages. These acœlomatous worms did not as yet possess a true body-cavity (cœlom) nor blood. They originated in the early primordial period out of the Gastræada, by the formation of a middle germ-layer, or muscular layer, and also by the further differentiation of the internal parts into various organs; more especially the first formation of a nervous system, the simplest organs of sense, the simplest organs for secretion (kidneys) and generation (sexual organs). The proof that human ancestors existed of a similar formation, is to be looked for in the circumstance that comparative anatomy and ontogeny point to the lower acœlomatous Worms as the common primary form, not merely of all higher Worms, but also of the four higher tribes of animals. Now, of all the animals known to us, the Turbellaria, which possess neither a body-cavity nor blood, are most closely allied to these primæval acœlomatous Primary Worms.

Seventh Stage: Soft Worms (Scolecida).

Between the Turbellaria of the preceding stage and the Sack Worms of the next stage, we must necessarily assume at least one connecting intermediate stage. For the Tunicata, which of all known animals stand nearest to the eighth stage, and the Turbellaria which most resemble the sixth stage, indeed both belong to the lower division of the unsegmented Worms; but still these two divisions differ so much from one another in their organization, that we must necessarily assume the earlier existence of extinct intermediate forms between the two. These connecting links, of which no fossil remains exist, owing to the soft nature of their bodies, we may comprise as Soft Worms, or Scolecida. They developed out of the Turbellaria of the sixth stage by forming a true body-cavity (a cœlom) and blood in their interior. It is difficult to say which of the still living Cœlomati are nearest akin to these extinct Scolecida; it may be the Acorn-worms (Balanoglossus). The proof that even the direct ancestors of man belonged to these Scolecida, is furnished by the comparative anatomy and the ontogeny of Worms and of the Amphioxus. The form value of this stage must moreover have been represented by several very different intermediate stages, in the wide gap between Turbellaria and Tunicata.

Eighth Stage: Sack Worms (Himatega).

Under the name of Sack worms, or Himatega, we here allude in the eighth place to those Cœlomati, out of which the most ancient skull-less Vertebrata were directly developed. Among the Cœlomati of the present day, the Ascidians are the nearest relatives of these exceedingly remarkable Worms, which connect the widely differing classes of Invertebrate and Vertebrate animals. That the ancestors of man really existed during the primordial period in the form of these Himatega, is distinctly proved by the exceedingly remarkable and important agreement presented by the ontogeny of the Amphioxus and the Ascidia. (Compare Plates XII. and XIII., also pp. 152, 200, etc.) From this fact the earlier existence of Sack Worms may be inferred; they of all known worms were most closely related to our recent Tunicates, especially to the freely swimming young forms or larvæ of the simple Sea-squirts (Ascidia, Phallusia). They originated out of the worms of the seventh stage by the formation of a dorsal nerve-marrow (medulla tube), and by the formation of the spinal rod (chorda dorsalis) which lies below it. It is just the position of this central spinal rod, or axial skeleton, between the dorsal marrow on the dorsal side, and the intestinal canal on the ventral side, which is most characteristic of all Vertebrate animals, including man, but also of the larvæ of the Ascidia. The form value of this stage nearly corresponds with that which the larvæ of the simple Sea-squirts possess at the time when they show the beginning of the dorsal marrow and spinal rod. (Plate XII. Fig. A 5: compare the explanation of these figures in the Appendix.)

SECOND HALF OF THE SERIES OF HUMAN ANCESTORS. VERTEBRATE ANIMAL ANCESTORS OF MAN
(Vertebrata).

Ninth Stage: Skull-less Animals (Acrania).

The series of human ancestors, which in accordance with their whole organisation we have to consider as Vertebrate animals, begins with the Skull-less animals, or Acrania, of whose nature the still living Lancelet (Amphioxus lanceolatus, Plate XII. B, XIII. B) gives us a faint idea. Since this little animal in its earliest embryonal state entirely agrees with the Ascidia, and in its further development shows itself to be a true Vertebrate animal, it forms a direct transition from the Vertebrata to the Invertebrata. Even if the human ancestors of the ninth stage in many respects differed from the Amphioxus—the last surviving representative of the Skull-less animals—yet they must have resembled it in its most essential characteristics, in the absence of head, skull, and brain. Skull-less animals of such structure—out of which animals with skulls developed at a later period—lived during the primordial period, and originated out of the Himatega of the eighth stage by the formation of the metamera, or body segments, as also by the further differentiation of all organs, especially the more perfect development of the dorsal nerve-marrow and the spinal rod lying below it. Probably the separation of the two sexes (gonochorism) also began at this stage, whereas all the previously mentioned invertebrate ancestors (apart from the 3—4 first neutral stages) exhibited the condition of hermaphrodites (hermaphroditism). (Compare vol. i. p. 196.) The certain proof of the former existence of these skull-less and brain-less ancestors of man, is furnished by the comparative anatomy and the ontogeny of the Amphioxus and of the Craniota.

Tenth Stage: Single-nostriled Animals (Monorrhina).

Out of the Skull-less ancestors of man there arose in the first place animals with skulls, or Craniota, of the most imperfect nature. The lowest stage of all still living Craniota is occupied by the class of round-mouthed animals, or Cyclostoma, namely, the Hag (Myxinoidea) and Lampreys (Petromyzontia). From the internal organization of these single-nostriled animals, or Monorrhina, we can form an approximate idea of the nature of the human ancestors of the tenth stage. In the former, as also in the latter, skull and brain must have been of the simplest form, and many important organs, as for example, the swimming bladder, the sympathetic nerve, the spleen, the jaw skeleton, and both pairs of legs, may probably as yet not have existed. However, the pouch gills and the round sucking mouth of the Cyclostoma must probably be looked upon as purely adaptive characteristics, which did not exist in the corresponding stage of ancestors. The single-nostriled animals originated during the primordial period out of the skull-less animals by the anterior end of the dorsal marrow developing into the brain, and the anterior end of the dorsal chord into the skull. The certain proof that such single-nostriled and jawless ancestors of man did exist, is found in the “comparative anatomy of the Myxinoidea.

”

Eleventh Stage: Primæval Fish (Selachii.).

Of all known Vertebrate animals, the ancestors of the Primæval Fish probably showed most resemblance to the still living Sharks (Squalacei). They originated out of the single-nostriled animals by the division of the single nostril into two lateral halves, by the formation of a sympathetic nervous system, a jaw skeleton, a swimming bladder, and two pairs of legs (breast fins or fore-legs, and ventral fins or hind-legs). The internal organisation of this stage may probably, upon the whole, have corresponded to the lowest species of Sharks known to us; the swimming bladder was however more strongly developed; in the case of the latter it exists only as a rudimentary organ. They lived as early as the Silurian period, as is proved by the fossil remains of sharks (teeth and fin spines) from the Silurian strata. A certain proof that the Silurian ancestors of man and of all the other double-nostriled animals were nearest akin to the Selachii, is furnished by the comparative anatomy of the latter; it shows that the relations of organisation in all Amphirrhina can be derived from those of the Selachii.

Twelfth Stage: Mud Fish (Dipneusta).

Our twelfth ancestral stage is formed by Vertebrate animals which probably possessed a remote resemblance to the still living Salamander fish (Ceratodus, Protopterus, Lepidosiren, p. 212). They originated out of the Primæval fish (probably at the beginning of the palæolithic, or primary period) by adaptation to life on land, and by the transformation of the swimming bladder into an air-breathing lung, and of the nasal cavity (which now opened into the cavity of the mouth) into air passages. The series of the ancestors of man which breathed air through lungs began at this stage. Their organisation may probably in many respects have agreed with that of the still living Ceratodus and Protopterus, but at the same time may have been very different. They probably lived at the beginning of the Devonian period. Their existence is proved by comparative anatomy, which shows the Dipneusta to be an intermediate stage between the Selachii and Amphibia.

Thirteenth Stage: Gilled Amphibians (Sozobranchia).

Out of those Mud Fish, which we considered the primary forms of all the Vertebrata which breathe through lungs, there developed the class of Amphibia as the main line (pp. 205, 216). Here began the five-toed formation of the foot (the Pentadactyla), which was thence transmitted to the higher Vertebrata, and finally also to Man. The gilled Amphibians must be looked upon as our most ancient ancestors of the class of Amphibia; besides possessing lungs they retained throughout life regular gills, like the still living Proteus and Axolotl (p. 218). They originated out of the Dipneusta by the transformation of the paddling fins into five-toed legs, and also by the more perfect differentiation of various organs, especially of the vertebral column. In any case they existed about the middle of the palæolithic, or primary period, possibly even before the Coal period; for fossil Amphibia are found in coal. The proof that similar gilled Amphibians were our direct ancestors, is given by the comparative anatomy and the ontogeny of Amphibia and Mammals.

Fourteenth Stage: Tailed Amphibians (Sozura).

Our amphibious ancestors which retained their gills throughout life, were replaced at a later period by other Amphibia, which, by metamorphosis, lost the gills which they had possessed in early life, but retained the tail, as in the case of the salamanders and newts of the present day. (Compare p. 218.) They originated out of the gilled Amphibians by accustoming themselves in early life to breathe only through gills, and later in life only through lungs. They probably existed even in the second half of the primary, namely, during the Permian period, but possibly even during the Coal period. The proof of their existence lies in the fact that tailed Amphibians form a necessary intermediate link between the preceding and succeeding stages.

Fifteenth Stage: Primæval Amniota (Protamnia).

The name Protamnion we have given to the primary form of the three higher classes of Vertebrate animals, out of which the Proreptilia and the Promammalia developed as two diverging branches (p. 222). It originated out of unknown tailed Amphibia by the complete loss of the gills, by the formation of the amnion, of the cochlea, and of the round window in the auditory organ, and of the organs of tears. It probably originated in the beginning of the mesolithic or secondary period, perhaps even towards the end of the primary, in the Permian period. The certain proof that it once existed lies in the comparative anatomy and the ontogeny of the Amniota; for all Reptiles, Birds, and Mammals, including Man, agree in so many important characteristics that they must, with full assurance, be admitted to be the descendants of a single common primary form, namely, of the Protamnion.

Sixteenth Stage: Primary Mammals (Promammalia).

We now find ourselves more at home with our ancestors. From the sixteenth up to the twenty-second stage they all belong to the large and well known class of Mammals, the confines of which we ourselves have as yet not transgressed. The common, long since extinct and unknown primary forms of all Mammalia, which we have named Promammalia, were at all events, of all still living animals, of the class most closely related to the Beaked animals, or Ornithostoma (Ornithorhynchus, Echidna, p. 233). They differed from the latter, however, by the teeth present in their jaws. The formation of the beak in the Beaked animals of the present day must be looked upon as an adaptive characteristic which developed at a later period. The Promammalia arose out of the Protamnia (probably only at the beginning of the secondary period, namely, in the Trias) by various advances in their internal organisation, as also by the transformation of the epidermal scales into hairs, and by the formation of a mammary gland which furnished milk for the nourishment of the young ones. The certain proof that the Promammalia—inasmuch as they are the common primary forms of all Mammals—also belong to our ancestors, lies in the comparative anatomy and the ontogeny of Mammalia and Man.

Seventeenth Stage: Pouched Animals (Marsupialia).

The three sub-classes of Mammalia—as we have already seen—stand in such a relation to one another that the Marsupials, both as regards their anatomy and their ontogeny and phylogeny, form the direct transition from the Monotrema to Placental animals (p. 247). Consequently, human ancestors must also have existed among Marsupials. They originated out of the Monotrema—which include the primary Mammalia, or Promammalia—by the division of the cloaca into the rectum and the urogenital sinus, by the formation of a nipple on the mammary gland, and by the partial suppression of the clavicles. The oldest Marsupials at all events existed as early as the Jura period (perhaps even in the Trias); during the Chalk period they passed through a series of stages preparing the way for the origin of Placentalia. The certain proof of our derivation from Marsupials—nearly akin to the still living opossum and kangaroo in their essential inner structure—is furnished by the comparative anatomy and the ontogeny of Mammalia.

Eighteenth Stage: Semi-apes (Prosimiæ).

The small group of Semi-apes, as we have already seen, is one of the most important and most interesting orders of Mammalia. It contains the direct primary forms of Genuine Apes, and thus also of Man. Our Semi-ape ancestors probably possessed only a very faint external resemblance to the still living, short-footed Semi-apes (Brachytarsi), especially the Maki, Indri, and Lori (p. 256). They originated (probably at the beginning of the Cenolithic, or Tertiary period) out of Marsupials of Rat-like appearance by the formation of a placenta, the loss of the marsupium and the marsupial bones, and by the higher development of the commissures of the brain. The certain proof that Genuine Apes, and hence also our own race, are the direct descendants of Semi-apes, is to be found in the comparative anatomy and the ontogeny of Placental animals.

Nineteenth Stage: Tailed Apes (Menocerca).

Of the two classes of Genuine Apes which developed out of the Semi-apes, it is only the narrow-nosed, or Catarrhini, which are closely related by blood to Man. Our older ancestors from this group probably resembled the still living Nose-apes and Holy-apes (Semnopithecus), which possess jaws and narrow noses like Man, but have a long tail, and their bodies densely covered with hair (p. 271). The Tailed Apes with narrow noses (Catarrhini Menocerci) originated out of Semi-apes by the transformation of the jaw, and by the claws on their toes becoming changed into nails; this probably took place as early as the older Tertiary period. The certain proof of our derivation from Tailed Catarrhini is to be found in the comparative anatomy and the ontogeny of Apes and of Man.

Twentieth Stage: Man-like Apes (Anthropoides).

Of all still living Apes the large tail-less, narrow-nosed Apes, namely, the Orang and Gibbon in Asia, the Gorilla and Chimpanzee in Africa, are most nearly akin to Man. It is probable that these Man-like Apes, or Anthropoides, originated during the Mid-tertiary period, namely, in the Miocene period. They developed out of the Tailed Catarrhini of the preceding stage—with which they essentially agree—by the loss of the tail, the partial loss of the hairy covering and by the excessive development of that portion of the brain just above the facial portion of the skull. There do not exist direct human ancestors among the Anthropoides of the present day, but they certainly existed among the unknown extinct Human Apes of the Miocene period. The certain proof of their former existence is furnished by the comparative anatomy of Man-like Apes and of Man.

Twenty-first Stage: Ape-like Men (Pithecanthropi).

Although the preceding ancestral stage is already so nearly akin to genuine Men that we scarcely require to assume an intermediate connecting stage, still we can look upon the speechless Primæval Men (Alali) as this intermediate link. These Ape-like men, or Pithecanthropi, very probably existed towards the end of the Tertiary period. They originated out of the Man-like Apes, or Anthropoides, by becoming completely habituated to an upright walk, and by the corresponding stronger differentiation of both pairs of legs. The fore hand of the Anthropoides became the human hand, their hinder hand became a foot for walking. Although these Ape-like Men must not merely by the external formation of their bodies, but also by their internal mental development, have been much more akin to real Men than the Man-like Apes could have been, yet they did not possess the real and chief characteristic of man, namely, the articulate human language of words, the corresponding development of a higher consciousness, and the formation of ideas. The certain proof that such Primæval Men without the power of speech, or Ape-like Men, must have preceded men possessing speech, is the result arrived at by an inquiring mind from comparative philology (from the “comparative anatomy” of language), and especially from the history of the development of language in every child (“glottal ontogenesis”) as well as in every nation (“glottal phylogenesis”).

Twenty-second Stage: Men (Homines).

Genuine Men developed out of the Ape-like Men of the preceding stage by the gradual development of the animal language of sounds into a connected or articulate language, of words. The development of this function, of course, went hand in hand with the development of its organs, namely, the higher differentiation of the larynx and the brain. The transition from speechless Ape-like Men to Genuine or Talking Men probably took place at the beginning of the Quaternary period, namely, in the Diluvial period, but possibly even at an earlier date, in the more recent Tertiary. As, according to the unanimous opinion of most eminent philologists, all human languages are not derived from a common primæval language, we must assume a polyphyletic origin of language, and in accordance with this a polyphyletic transition from speechless Ape-like Men to Genuine Men.

CHAPTER XXIII.

MIGRATION AND DISTRIBUTION OF MANKIND. HUMAN SPECIES AND HUMAN RACES.

The rich treasure of knowledge we possess in the comparative anatomy and the history of the development of Vertebrate animals, enables us even now to establish the most important outlines of the human pedigree in the way we have done in the last chapter. One must, however, not expect to be able to survey satisfactorily in every detail the history or phylogeny of the human species which will henceforth form the basis of Anthropology, and of all other sciences. The complete development of this most important science—of which we can only lay the first foundation—must remain reserved for the more accurate and extensive investigations of a future time. This applies also to those more special questions of human phylogeny at which it is desirable before concluding to take a cursory glance, namely, the question of the time and place of the origin of the human race, as also of the different species and races into which it has differentiated.

In the first place, the period of the earth’s history, within which the slow and gradual transmutation of the most man-like apes into the most ape-like men took place, can of course not be determined by years, nor even by centuries. This much can, however, with full assurance be maintained, for reasons given in the last chapter, that Man is derived from Placental animals. Now, as fossil remains of these Placentalia are found only in the tertiary rocks, the human race can at the earliest have developed only within the Tertiary period out of perfected man-like apes. What seems most probable is that this most important process in the history of terrestrial creation occurred towards the end of the Tertiary period, that is in the Pliocene, perhaps even in the Miocene period, but possibly also not until the beginning of the Diluvial period. At all events Man, as such, lived in central Europe as early as the Diluvial period, contemporaneously with many large, long since extinct mammals, especially with the diluvial elephant, or mammoth (Elephas primigenius), the woolly-haired rhinoceros (Rhinoceros tichorrhinus), the giant deer (Cervus euryceros), the cave bear (Ursus spelæus), the cave hyæna (Hyæna spelæa), the cave lion (Felis spelæus), etc. The results brought to light by recent geology and archæology as to these fossil men and their animal contemporaries of the diluvial period, are of the greatest interest. But as a closer examination of them would occupy too much of my limited space, I must confine myself here to setting forth their great general importance, and refer for particulars to the numerous writings which have recently been published on the Primæval History of Man, more especially to the excellent works of Charles Lyell,(30) Carl Vogt,(27) Friedrich Rolle,(28) John Lubbock,(44) L. Büchner,(43) etc.

The numerous and interesting discoveries presented to us by these extensive investigations of late years on the primæval history of the human race, place the important fact (long since probable for many other reasons) beyond a doubt, that the human race, as such, has existed for more than twenty thousand years. But it is also probable that more than a hundred thousand years, perhaps many hundred thousands of years, have elapsed since its first appearance; and, in contrast to this, it must seem very absurd that our calendars still represent the “Creation of the World, according to Calvisius,” to have taken place 5821 years ago.

Now, whether we reckon the period during which the human race, as such, has existed and diffused itself over the earth, as twenty thousand, a hundred thousand, or many hundred thousands of years, the lapse of time is in any case immensely small in comparison with the inconceivable length of time which was requisite for the gradual development of the long chain of human ancestors. This is evident even from the small thickness of all Diluvial deposits in comparison with the Tertiary, and of these again in comparison with the preceding deposits. (Compare p. 22.) But the infinitely long series of slowly and gradually developing animal forms from the simplest Moneron to the Amphioxus, from this to the Primæval Fish, from the Primæval Fish to the first Mammal, and again, from the latter to Man, also require for their historical development a succession of periods probably comprising many thousands of millions of years. (Compare vol. i. p. 129.)

Those processes of development which led to the origin of the most Ape-like Men out of the most Man-like Apes must be looked for in the two adaptational changes which, above all others, are distinctive of Man, namely, upright walk and articulate speech. These two physiological functions necessarily originated together with two corresponding morphological transmutations, with which they stand in the closest correlation, namely, the differentiation of the two pairs of limbs and the differentiation of the larynx. The important perfecting of these organs and their functions must have necessarily and powerfully reacted upon the differentiation of the brain and the mental activities dependent upon it, and thus have paved the way for the endless career in which Man has since progressively developed, and in which he has far outstripped his animal ancestors. (Gen. Morph. ii. p. 430.)

The first and earliest of these three great processes in the development of the human organism probably was the higher differentiation and the perfecting of the extremities which was effected by the habit of an upright walk. By the fore feet more and more exclusively adopting and retaining the function of grasping and handling, and the hinder feet more and more exclusively the function of standing and walking, there was developed that contrast between the hand and foot which is indeed not exclusively characteristic of man, but which is much more strongly developed in him than in the apes most like men. This differentiation of the fore and hinder extremities was, however, not merely most advantageous for their own development and perfecting, but it was followed at the same time by a whole series of very important changes in other parts of the body. The whole vertebral column, and more especially the girdle of the pelvis and shoulders, as also the muscles belonging to them, thereby experienced those changes which distinguish the human body from that of the most man-like apes. These transmutations were probably accomplished long before the origin of articulate speech; and the human race thus existed for long, with an upright walk and the characteristic human form of body connected with it, before the actual development of human language, which would have completed the second and the more important part of human development. We may therefore distinguish a special (21st) stage in the series of our human ancestors, namely, Speechless Man (Alalus), or Ape-man (Pithecanthropus), whose body was indeed formed exactly like that of Man in all essential characteristics, but who did not as yet possess articulate speech.

The origin of articulate language, and the higher differentiation and perfecting of the larynx connected with it, must be looked upon as only a later, and the most important stage in the process of the development of Man. It was, doubtless, this process which above all others helped to create the deep chasm between man and animal, and which also first caused the most important progress in the mental activity and the perfecting of the brain connected with it. There indeed exists in very many animals a language for communicating sensations, desires, and thoughts, partly a language of gestures, partly a language of feeling or touch, partly a language of cries or sounds, but a real language of words or ideas, a so-called “articulate” language, which by abstraction changes sounds into words, and words into sentences, belongs, as far as we know, exclusively to Man.

The origin of human language must, more than anything else, have had an ennobling and transforming influence upon the mental life of Man, and consequently upon his brain. The higher differentiation and perfecting of the brain and mental life as its highest function developed in direct correlation with its expression by means of speech. Hence, the highest authorities in comparative philology justly see in the development of human speech the most important process which distinguishes Man from his animal ancestors. This has been especially set forth by August Schleicher, in his treatise “On the Importance of Speech for the Natural History of Man.”(34) In this relation we see one of the closest connections between comparative zoology and comparative philology; and here the theory of development assigns to the latter the task of following the origin of language step by step. This task, as interesting as it is important, has of late years been successfully undertaken by many inquirers, but more especially by Wilhelm Bleek, who has been occupied for seventeen years in South Africa with the study of the languages of the lowest races of men, and hence has been enabled to solve the question. August Schleicher more especially discusses, in accordance with the theory of selection, how the various forms of speech, like all other organic forms and functions, have developed by the process of natural selection, and have divided into many species and dialects.

I have no space here to follow the process of the formation of language, and must refer in regard to this to the above-mentioned important work of Wilhelm Bleek, “On the Origin of Language.”(35) But we have still to mention one of the most important results of comparative philology, which is of the highest importance to the genealogy of the human species, that is, that human language was probably of a multiple, or polyphyletic origin. Human speech, as such, did not develop probably until the genus of Speech-less or Primæval Man, or Ape Man, had separated into several kinds or species. In each of these human species, and perhaps even in the different sub-species and varieties of this species, language developed freely and independently of the others. At least Schleicher, one of the first authorities on the subject, maintains that “even the beginnings of language—in sounds as well as in regard to ideas and views which were reflected in sounds, and further, in regard to their capability of development—must have been different. For it is positively impossible to trace all languages to one and the same primæval language. An impartial investigation rather shows that there are as many primæval languages as there are races.”(34) In like manner, Friederich Müller(41) and other eminent linguists assume a free and independent origin of the families of languages and their primæval stocks. It is well known, however, that the boundaries of these tribes of languages and their ramifications are by no means always the boundaries of the different human species, or the so-called “races,” distinguished by us on account of their bodily character istics. This, as well as the complicated relations of the mixture of races, and the various forms of hybrids, is the great difficulty lying in the way of tracing the human pedigree in its individual branches, species, races, varieties, etc.

In spite of these great and serious difficulties, we cannot here refrain from taking one more cursory glance at the ramification of the human pedigree, and at the same time considering, from the point of view of the theory of descent, the much discussed question of the monophyletic or polyphyletic origin of the human race, and its species or races. As is well known, two great parties have for a long time been at war with each other upon this question; the monophylists (or monogenists) maintain the unity of origin and the blood relationship of all races of men. The polyphylists (or polygenists), on the other hand, are of opinion that the different races of men are of independent origin. According to our previous genealogical investigations we cannot doubt that, at least in a wide sense, the monophyletic opinion is the right one. For even supposing that the transmutation of Man-like Apes into Men had taken place several times, yet those Apes themselves would again be allied by the one pedigree common to the whole order of Apes. The question therefore would always be merely about a nearer or remoter degree of blood relationship. In a narrower sense, on the other hand, the polyphylist’s opinion would probably be right, inasmuch as the different primæval languages have developed quite independently of one another. Hence, if the origin of an articulate language is considered as the real and principal act of humanification, and the species of the human race are distinguished according to the roots of their language, it might be said that the different races of men had originated, independently of one another, by different branches of primæval, speechless men directly springing from apes, and forming their own primæval language. Still they would of course be connected further up or lower down at their root, and thus all would finally be derived from a common primæval stock.

While we hold the latter of these convictions, and while we for many reasons believe that the different species of speechless primæval men were all derived from a common ape-like human form, we do not of course mean to say that all men are descended from one pair. This latter supposition, which our modern Indo-Germanic culture has taken from the Semitic myth of the Mosaic history of creation, is by no means tenable. The whole of the celebrated dispute, as to whether the human race is descended from a single pair or not, rests upon a completely false way of putting the question. It is just as senseless as the dispute as to whether all sporting dogs or all race-horses are descended from a single pair. We might with equal justice ask whether all Germans or all Englishmen are “descended from a single pair,” etc. A “first human pair,” or “a first man,” has in fact never existed, any more than there ever existed a first pair or a first individual of Englishmen, Germans, race-horses, or sporting dogs. The origin of a new species, of course, always results from an existing species, by a long chain of many different individuals sharing the slow process of transformation. Supposing that we had all the different pairs of Human Apes and Ape-like Men before us—which belong to the true ancestors of the human race—it would even then be quite impossible (without doing so most arbitrarily) to call any one of these pairs of ape-like men “the first pair.” As little can we derive each of the twelve races or species of men, which we shall consider directly, from a “first pair.”

The difficulties met with in classifying the different races or species of men are quite the same as those which we discover in classifying animal and vegetable species. In both cases forms apparently quite different are connected with one another by a chain of intermediate forms of transition. In both cases the dispute as to what is a kind or a species, what a race or a variety, can never be determined. Since Blumenbach’s time, as is well known, it has been thought that mankind may be divided into five races or varieties, namely: (1) the Ethiopian, or black race (African negro); (2) the Malayan, or brown race (Malays, Polynesians, and Australians); (3) the Mongolian, or yellow race (the principal inhabitants of Asia and the Esquimaux of North America); (4) the Americans, or red race (the aborigines of America); and (5) the Caucasian, or white race (Europeans, north Africans, and south-western Asiatics). All of these five races of men, according to the Jewish legend of creation, are said to have been descended from “a single pair”—Adam and Eve,—and in accordance with this are said to be varieties of one kind or species. If, however, we compare them without prejudice, there can be no doubt that the differences of these five races are as great and even greater than the “specific differences” by which zoologists and botanists distinguish recognised “good” animal and vegetable species (“bonæ species”). The excellent palæontologist Quenstedt is right in maintaining that, “if Negroes and Caucasians were snails, zoologists would universally agree that they represented two very excellent species, which could never have originated from one pair by gradual divergence.”

The characteristics by which the races of men are gradually distinguished are partly taken from the formation of the hair, partly from the colour of the skin, and partly from the formation of the skull. In regard to the last character, two extremes are distinguished, namely, long heads and short heads. In long-headed men (Dolichocephali), whose strongest development is found in Negroes and Australians, the skull is extended, narrow, and compressed on the right and left. In short-headed men (Brachycephali), on the other hand, the skull is compressed in an exactly opposite manner, from the front to the back, is short and broad, which is especially striking in the case of the Mongolians. Medium-headed men (Mesocephali), standing between the two extremes, predominate especially among Americans. In every one of these three groups we find men with slanting teeth (Prognathi), whose jaws, like those of the animal snout, strongly project, and whose front teeth therefore slope in front, and men with straight teeth (Orthognathi), whose jaws project but little, and whose front teeth stand perpendicularly. During the last ten years a great deal of time and trouble have been devoted to the careful examination and measurement of the forms of skulls, which have, however, not been rewarded by corresponding results. For within a single species, as for example within the Mediterranean species, the form of the skull may vary so much that both extremes are met with in the same species. Much better starting-points for the classification of of the human species are furnished by the nature of the hair and speech, because they are much more strictly hereditary than the form of the skull.

Comparative philology seems especially to be becoming an authority in this matter. In the latest great work on the races of men, which Friederich Müller has published in his excellent “Ethnography,”(42) he justly places language in the fore-ground. Next to it the nature of the hair of the head is of great importance; for although it is in itself of course only a subordinate morphological character, yet it seems to be strictly transmitted within the race. Of the twelve species of men distinguished on the following table (p. 308), the four lower species are characterised by the woolly nature of the hair of their heads; every hair is flattened like a tape, and thus its section is oval. These four species of woolly-haired men (Ulotrichi) we may reduce into two groups—tuft-haired and fleecy-haired. The hair on the head of tuft-haired men (Lophocomi), Papuans and Hottentots, grows in unequally divided small tufts. The woolly hair of fleecy-haired men (Eriocomi), on the other hand, in Caffres and Negroes, grows equally all over the skin of the head. All Ulotrichi, or woolly-haired men, have slanting teeth and long heads, and the colour of their skin, hair, and eyes is always very dark. All are inhabitants of the Southern Hemisphere; it is only in Africa that they come north of the equator. They are on the whole at a much lower stage of development, and more like apes, than most of the Lissotrichi, or straight-haired men. The Ulotrichi are incapable of a true inner culture and of a higher mental development, even under the favourable conditions of adaptation now offered to them in the United States of North America. No woolly-haired nation has ever had an important “history.”

PEDIGREE OF TWELVE SPECIES OF MEN