[66] In giving this description the red (Glyn) slates of North Wales are treated as belonging to the Caerfai series, though this correlation depends on lithological characters only at present.
The Menevian beds consist essentially of very fine, well laminated black and grey muds, which are of a texture favourable for the production of a somewhat regular jointing, causing the rock to break into small rectangular blocks. They are thin, not exceeding 600 feet in thickness, and indicate the incoming of the general deep-water phase of the Lower Palæozoic epoch. The Lingula Flags mark a local return to shallower water conditions, especially in the central portion. The total thickness is over 3,000 feet, of which the lower stage (locally the Maentwrog series) is over 500 feet, and consists of blackish muds, the middle (Festiniog stage[67]) is about 2,000 feet thick, and is composed chiefly of shallower water gritty flags, whilst the upper (Dolgelly) stage is of about the same thickness as the lower stage and has similar lithological characters.
[67] The term Festiniog has been used for the whole Lingula Flag series as well as for the middle stage. It will be well to use it with reference to the stage only.
The Tremadoc Slates are about 1,000 feet thick. They are divided into a lower and upper stage, of about equal thickness, and are essentially composed of iron-stained slates, with a considerable admixture of calcareous matter in some parts of South Wales, when they furnish the nearest approach to a limestone which has been found amongst the Welsh Cambrian strata. They were probably formed in a fairly deep sea.
Much pyroclastic rock and some lava flows are intercalated amongst the Welsh Cambrian sediments. Tuffs are formed in the lower beds of St David's, and lavas and ashes have been found amongst the Lingula Flags and Tremadoc Slates of North Wales, while the Lingula Flags of South Wales have furnished several bands of ash to the north of Haverfordwest. Much of the material of the grits and muds may be derived from volcanic rocks, though how far this is so cannot be stated in the absence of information obtained by detailed petrological examination of the rocks.
The various isolated outcrops of Cambrian strata amongst the counties of the Welsh borders and adjoining Midland counties indicate a great thinning of the Cambrian rocks in this direction.
The probable equivalents of the Caerfai rocks occur at Nuneaton, Comley, and on the flanks of the Wrekin and Malvern hills. The thin basal conglomerates are succeeded by quartzites, and sometimes red calcareous sandstones (Comley sandstone). These rocks are succeeded by thin arenaceous and calcareous beds which represent either the Solva or Menevian beds of Wales. The Lingula Flags are represented by the Malvern Shales of the Malvern area and the Stockingford Shales of Nuneaton, whilst the Tremadoc Slates have as their equivalents the Shineton Shales. The exact thicknesses of these deposits do not seem to have been recorded, but Prof. Lapworth observes that in central Shropshire "the Comley and Shineton groups which ... have a collective thickness of perhaps less than 3,000 feet, we have apparently a condensed epitome of the entire Cambrian system as at present generally defined."
The Cambrian rocks of the North-west Highlands consist of a thin conglomerate succeeded by grits and flags with shaley beds, and above these a mass of limestone, which may represent some of the Ordovician deposits as well as those of Cambrian age. Pending a complete description of the faunas of these rocks, it is sufficient to state that the only fauna which has hitherto been described in detail indicates the existence of Lowest Cambrian rocks. Further remarks will be made on this head when describing the character of the Cambrian faunas. The Cambrian rocks of the North-west Highlands are also very thin as compared with those of Wales, so that the Highland and Welsh borderland regions appear to have existed as a deeper sea area than that which is indicated by the Cambrian rocks of Wales, an inference which is to some extent borne out by study of the Cambrian rocks of extra-British areas, to which we may now turn.
The principal European developments of Cambrian rock are found in Scandinavia, Russia, Bohemia and Spain, and of these the Scandinavian one is by far the most fully developed, as there is a complete sequence in the rocks of that peninsula. They occur both in Norway and Sweden, but the Swedish exposures are the most interesting in most respects, especially those of Westrogothia and Scania. The rocks are of no great thickness, and consist essentially of black carbonaceous shales, with inconstant bands of impure black limestone composed almost entirely of the remains of trilobites or more rarely of brachiopods. These Alum Shales, as they are termed, rest unconformably upon Precambrian rocks, and have arenaceous and conglomeratic deposits at the base. In Russia the rocks are still further attenuated, and have not yielded the relics of so many faunas as have been found in the Scandinavian Cambrian rocks.
The Bohemian development is incomplete, owing apparently to an unconformity at the base of the overlying Ordovician rocks, while the Spanish deposits which seem fairly thick and composed largely of mechanical sediments have not been worked out in very great detail.
The American development of Cambrian rocks resembles the European one in many striking particulars, and as in the case of Europe, there are lateral variations in the lithological characters of the rocks, though in the opposite direction, the shallow-water deposits occurring on the east coast, and the deep-water deposits further west.
The general distribution of the different types of Cambrian strata in Europe and North America has been accounted for on the supposition that in Cambrian times a tract of land lay over much of the present site of the North Atlantic Ocean, and that the detritus of that land formed the shallow-water accumulations of Wales and the east of Canada, whilst further away from it were deposited the open-sea accumulations of Scandinavia and Russia on one side and of the more westerly regions of North America on the other, as indicated in Fig. 16.
The Cambrian Faunas. The Cambrian Period has been termed the age of trilobites, for they are the dominant forms of the time, but they are associated with many other forms of invertebrata; indeed all the great groups of this division are represented in the earliest Cambrian fauna. Dr C. D. Walcott records representatives of Spongiae, Hydrozoa, Echinodermata, Annelida, Brachiopoda, Lamellibranchiata, Gastropoda, Pteropoda, Crustacea and Trilobita as occurring in the Olenellus beds of North America and other groups are represented in the rocks of this age in the Old World. The Cambrian trilobites as a whole are of more generalised types than those of the later systems which furnish their remains, as indicated especially by the looseness of the body, and the large number of body rings in many of the genera, while the tail or pygidium was small and formed of only a few coalesced segments, as pointed out by Barrande. In the later trilobites the test is more compact, there are on the whole fewer body rings, as more of these have become fused into a tail which is therefore larger than that of the average tail of the Cambrian trilobite.
Taking the faunas in order, the oldest or Olenellus fauna has furnished a great variety of forms in the North-west Highlands of Scotland, Shropshire, Scandinavia, Esthonia, Sardinia, Canada, and Newfoundland, whilst representative species of the fauna have been recorded also from Worcestershire, Warwickshire, Pembrokeshire, India, China, and Australia.
The dominant form is the trilobite of the genus or group Olenellus, which contains a great variety of species referable to three or four divisions which have been ranked as separate genera by some writers. Associated with Olenellus are trilobites belonging to other genera, which are found in higher deposits, though there represented by different species.
Brachiopods are fairly abundant, especially those provided with a horny shell; of these, the genus Kutorgina is widely distributed.
The zoological relationships of several of the fossils of this horizon are as yet doubtful. The Archæocyathinæ show affinities with certain corals; a number of tests, included in the genus Hyolithes and its allies are doubtfully referred to the Pteropods, and the position of the genus Volborthella is uncertain. Special attention is directed to these doubtful relationships, as it is possible that a number of 'generalised forms' of organisms occur in these strata[68].
[68] For an account of the Olenellus fauna see Walcott, C. D., "The Fauna of the Lower Cambrian or Olenellus Zone," Tenth Annual Report of the Director of the United States Geological Survey, Washington, 1890. It is possible that some of the fossils mentioned in that report belong to strata above that containing Olenellus.
It should be noticed here that faunas have been discovered which are possibly of earlier date than the Olenellus fauna, as they do not correspond with it, or with those of newer strata. One, the Neobolus fauna of the Salt Range of India, occurs in beds below those with Olenellus, though it is not yet clear that Olenellus will not be eventually discovered associated with it, whilst the other, the Protolenus fauna of Canada, is of unknown age[69].
[69] For an account of the Neobolus beds see Noetling, F., "On the Cambrian Formation of the Eastern Salt Range," Records Geol. Survey, India, vol. XXVII. p. 71, and for the Protolenus fauna consult a paper by Matthew, G. F., "The Protolenus Fauna," Trans. New York Acad. of Science, 1895, vol. XIV. p. 101.
The Olenellus beds are succeeded by beds containing the Paradoxides fauna, which have been found in North and South Wales, Shropshire, Scandinavia, Bohemia, Spain, and North and South America. Olenellus and its allies became extinct (or else so scarce that no relics of them have been discovered in the Paradoxides beds) before the commencement of the deposition of the strata containing the Paradoxides fauna, and few genera pass from the beds with the one fauna to that containing the other. The Paradoxides fauna existed for a considerable period, and the beds have been divided into a series of zones characterised by different species of Paradoxides, thus
Dr Hicks records the following zones in Pembrokeshire[70]:—
| Zone of | Paradoxides | Davidis | Menevian. | |
| " | " | Hicksii | ||
| " | " | Aurora | Solva. | |
| " | " | Solvensis | ||
| " | " | Harknessi |
[70] The order here as elsewhere is ascending, i.e. the newest deposit is placed at the top.
Dr Tullberg divides the Paradoxides beds of Scania into thirteen zones, though only a few of these are characterised by definite species of Paradoxides. The Olenellus beds have not yet been divided into zones, though this will probably be the outcome of further study[71].
[71] The Paradoxides fauna is described in the following works: Britain, Hicks, H. and Salter J. W., Quart. Journ. Geol. Soc., vol. XXIV. p. 510, XXV. p. 51, XXVII. p. 173, and Hicks, H. and Harkness, R., ibid. vol. XXVII. p. 384; Scandinavia, Angelin, N. P., Palæontologia Scandinavica; Brögger, W. C., Nyt Magazin for Naturvidenskaberne, vol. XXIV., Linnarsson, G., Sveriges Geologiska Undersökning, Ser. C. No. 35; Bohemia, Barrande, J., Système Silurien du centre de la Bohême; Spain, Prado, C. de, "Sur l'existence de la faune Primordiale dans la chaîne Cantabrique suivie de la description des Fossiles par MM. de Verneuil et Barrande," Bull. Soc. Geol. France, 2 Series, vol. XVII. p. 516; America, Walcott, C. D., Bull. U. S. Geol. Survey: "The Cambrian Faunas of North America," and Matthew, G. F., Trans. Roy. Soc. Canada, 1882 and succeeding years.
The strata with Paradoxides are succeeded by those with the Olenus fauna, characterised by the genus Olenus and a large number of allied genera or sub-genera as some prefer to term them. The genus Olenus (sensu stricto) is very abundant in the lower part of the series, whilst the allied forms are more abundant in the upper beds. The genus Paradoxides and its associates disappeared before the deposition of these strata containing Olenus and its allies, and indeed the complete change in the character of the faunas in Europe is very remarkable. The Olenus fauna has been found in North Wales, Pembrokeshire, Warwickshire, Worcestershire, and abroad in Scandinavia and Canada. It is interesting to note among the fossils of the Olenus beds the occurrence of a graptolite which is associated with Olenus in Scandinavia; this is the earliest recorded appearance of a group which is destined to play so important a role amongst the fossils of the succeeding system[72]. The following zones have been detected by Dr S. A. Tullberg amongst the Olenus beds of Scania:—
| Zone of | Acerocare ecorne. |
| " | Dictyograptus flabelliformis. |
| " | Cyclognathus micropygus. |
| " | Peltura scarabæoides. |
| " | Eurycare camuricorne. |
| " | Parabolina spinulosa. |
| " | Ceratopyge sp. |
| " | Olenus (proper). |
| " | Leperditia. |
| " | Agnostus pisiformis. |
[72] For descriptions of the Olenus fauna consult the following:—Wales, Belt, T., Geol. Mag. Dec. I. vol. V. p. 5, and Salter, J. W., Decades Geol. Survey, Decade II. Pl. IX. and Decade XI. Pl. VIII.; Scandinavia, Angelin, N. P., Palæontologia Scandinavica, and Brögger, W. C., Die Silurischen Etagen 2 und 3 im Kristianiagebiet und auf Eker; Canada, Matthew, G. F., "Illustrations of the Fauna of the St John Group, No. VI.," Trans. Roy. Soc. Canada, 1891.
The beds with Dictyograptus flabelliformis form a wonderfully constant horizon at or near the top of the Olenus beds. They are found in North Wales, the Border Counties between Wales and England, France, Scandinavia, Russia and Canada.
The passage fauna of the beds which are the equivalents of the Tremadoc Slates may be spoken of as the Ceratopyge fauna, for Ceratopyge forficula, a remarkable species of trilobite, characterises it in Scandinavia, and will probably be found elsewhere. Ceratopyge beds have been found in North and South Wales, Shropshire, Scandinavia, Bavaria and North America, and in each case the fauna is intermediate in character between that of the Cambrian and that of the Ordovician system, containing the loosely-formed trilobites of the former with the more compact ones of the latter. The genus Bryograptus, a many-branched graptolite, also appears to characterise this fauna[73].
[73] For accounts of the Tremadoc Slates Fauna in England and Wales see Ramsay, A. C., Geology of North Wales, Appendix; Hicks, H., Quart. Journ. Geol. Soc., vol. XXIX. p. 39; Callaway, C., ibid. vol. XXXIII. p. 652, whilst many of the foreign fossils are noticed in Brögger's Die Silurischen Etagen 2 und 3 and Barrande's Faune silurienne des Environs de Hof en Bavière.
The faunas of the Cambrian rocks have not been studied in sufficient detail, with reference to the physical surroundings of the organisms, to throw much light upon the conditions under which the strata were deposited, though the evidence obtained from an examination of the lithological characters of the deposits is generally corroborated by study of the organic contents.
THE ORDOVICIAN SYSTEM.
Classification. The Ordovician strata were originally divided into series by Sedgwick as follows:—
Upper Bala,
Middle Bala,
Lower Bala,
Arenig.
The Arenig series was at one time included by some writers with the Lower Bala under the name Llandeilo, but the word Llandeilo is now used in the sense of Sedgwick's Lower Bala. The Middle Bala is often spoken of as Caradoc, but the terms Bala and Caradoc are sometimes used interchangeably. As much confusion attaches to the use of the name Bala without explanation, the alternative titles have been largely adopted, and as the series are well defined there is no objection to their use, save that some expression is wanted equivalent to Upper Bala. The local term Ashgill shales was originally applied by Mr W. Talbot Aveline to beds of this age in Lakeland, and I have elsewhere suggested the use of this name for the whole series in that region; its use may well be extended to the series which is developed in many parts of Britain and the continent. The terms which will be used here, therefore, for the different series of the Ordovician system are the following:—
| Ashgill | Series | (= Upper | Bala) |
| Caradoc | " | (= Middle | " ) |
| Llandeilo | " | (= Lower | " ) |
| Arenig. | " | ||
Adopting a palæontological classification, we may speak of the Arenig and Llandeilo beds as those containing the Asaphus fauna, whilst the Caradoc and Ashgill beds possess the Trinucleus fauna; this is the terminology employed by Angelin for the equivalent strata of Sweden. It must be noted that here the names applied are not those of absolutely characteristic genera, as was the case with those adopted for naming the Cambrian faunas, for both Asaphus and Trinucleus range through the beds of the system; but whereas Asaphus is most abundant in the beds of the two lower series, Trinucleus occurs most frequently in those of the two upper series.
Description of the strata. The Ordovician rocks are found over large tracts in North and South Wales, in the counties on the Welsh border, in Lakeland and the outlying districts in the Southern Uplands of Scotland, and in detached areas in Ireland. There are three main types of deposit:—(i) the volcanic type, in which the ordinary sediments are associated with a large amount of contemporaneous volcanic matter, (ii) the black shale type, with a fauna consisting largely of graptolites, and (iii) the ordinary sedimentary type, in which we find alternations of grits, shales, and more or less impure limestones. We also find developments which are intermediate between any two or even all three of these types. The first type is characteristically developed in Caernarvonshire and Merionethshire, the second in the Dumfriesshire Uplands, and the third in the Girvan district of Ayrshire. The variation in the thickness of these three types of deposit is shown in the accompanying sections of the Caernarvon, Merioneth, Moffat and Girvan regions (see Fig. 17).
The North Welsh area gives two different developments of the Ordovician strata, one of which is much less volcanic than the other. In the Merioneth-Caernarvon area, two great masses of volcanic rock form the Aran and Arenig hills of Merioneth and the Snowdonian group of Caernarvon. The former are of Arenig, the latter of Caradoc age. The Merionethshire volcanic rocks consist of a great thickness of lavas and ashes of intermediate composition (anderites), associated with sandy and muddy sediments of no great vertical depth. The Llandeilo beds of this area are chiefly of the nature of black shales, while the Caradoc series is represented by volcanic lavas and ashes of acid composition (felsites) with a few thin interbedded sediments. A calcareous ash forming the summit of Snowdon is of importance as being on the same horizon as a limestone (the Bala limestone) found in the other North Welsh area. The Ashgill series is not represented in Snowdonia.
In the other North Welsh tract, around Bala Lake, the volcanic matter is much less conspicuous. The Arenig rocks are not seen nearer than the Arenig mountains which form the western boundary of this second tract. The Llandeilo beds consist of shaley deposits with a well-marked limestone, the Llandeilo limestone, in the centre, whilst the Caradoc beds consist chiefly of muddy sediments with some thin ashes and a limestone, the Bala limestone, at the top. The Ashgill series contains a basal limestone, the Rhiwlas limestone, succeeded by shales, and another thin limestone called the Hirnant limestone at the summit.
In South Wales the Arenig beds[74] are chiefly composed of slates, and are divisible into an upper and lower group. The total thickness is about 2000 feet. The Llandeilo beds contain three series:—
| Upper Llandeilo Slates | 1000 |
| Llandeilo Limestone | 200 |
| Lower Llandeilo Slates | 800. |
[74] A remarkable fauna, fairly well represented in Britain and exceedingly well developed on the continent, exists in the Uppermost Arenig and Lower Llandeilo beds, and it is well separated from the dominant Arenig fauna below and Llandeilo fauna above. To the beds which contain it Dr Hicks has given the name Llanvirn series.
The Caradoc beds consist of black graptolitic shales of no great thickness, succeeded by an impure limestone on the horizon of the Bala limestone, while the Ashgill series like that of North Wales is separated into upper and lower limestone stages with an intervening stage composed of shales.
The deposits of the Welsh borderland are well developed in Shropshire, where there is practically a repetition of the Caernarvon-Merioneth development, with variations in detail. The Arenig and Caradoc volcanic rocks are not so thick as those of the Welsh district, but are nevertheless of considerable importance[75].
[75] For information concerning these beds see Lapworth, C. and Watts, W. W., "The Geology of South Shropshire," Proc. Geol. Assoc., vol. XIII. p. 297.
In the hilly region of Cumberland, Westmorland, and the adjoining parts of Yorkshire the succession differs from that of any of the Welsh regions, for the great period of volcanicity was during the formation of the Llandeilo rocks, and there were merely sporadic outbursts in Arenig and Caradoc times. The Arenig rocks consist of black shales with interstratified beds of coarser sediment, and some thin lavas and ashes of intermediate type. The Llandeilo series is represented by a very great thickness of volcanic rocks, varying in composition from basic to acid lavas, with associated pyroclastic rocks. The rocks of the Caradoc period largely consist of impure limestone with associated argillaceous rocks, and contemporaneous volcanic rocks of acid character. A marked unconformity is found locally in the centre of these. The Ashgill series consists of a basal limestone with shales above, and there is evidence that volcanic activity had not become extinct during the deposition of the rocks of this series.
Passing on to Scotland, the graptolitic type is admirably shown in the southern Uplands of the neighbourhood of Moffat, Dumfriesshire. The base of the Ordovician system has not been found, but the lowest series seems to be represented by shales with a graptolite possibly of Arenig age. Above this are volcanic beds succeeded by a group of black shales known as the Moffat shales. They are only about six hundred feet in thickness, and yet represent much of the Ordovician and part of the Silurian strata as developed elsewhere. The beds belonging to the Ordovician system are divided into two series, the Glenkiln shales below and the Hartfell shales above. The former consist of intensely black muds with few fossils save graptolites, and a deposit of chert at the base which is composed of radiolaria. The graptolites of the black shales are Upper Llandeilo forms, but the thin deposit of radiolarian chert may represent the rest of the Llandeilo period and part of the Arenig period also. The Hartfell shales are also usually black graptolite shales with lighter deposits nearly barren of organic remains; they represent the Caradoc and Ashgill series and pass conformably into the deposits of Silurian age[76]. The ordinary sedimentary type of Ordovician rocks is found in Ayrshire, though a few thin graptolitic seams are intercalated with the conglomerates and shelly sands, clays and limestones of the region, which is therefore peculiarly valuable as affording a means of comparison of the shelly type with the graptolitic type of Ordovician deposits. The Arenig series consists of black shales with graptolites, and these rocks are succeeded by a volcanic group which is probably of Llandeilo age. Above these volcanic beds, as in Dumfriesshire, we find three great divisions, two of which are of Ordovician, the third of Silurian age. The Ordovician divisions are respectively termed the Barr series, which is the equivalent of the Glenkiln shales, and the Ardmillan series above, equivalent to the Hartfell shales[77].
[76] The Moffat beds are described in a paper by Prof. Lapworth entitled "The Moffat Series" in the Quarterly Journal of the Geological Society, vol. XXXIV. p. 239. This paper, which is a masterpiece of detailed work, has furnished a clue to many problems. Few students will be able to follow the numerous details, and for general information concerning the beds they are recommended to read another paper by the same author "On the Ballantrae Rocks of South Scotland," Geol. Mag. Dec. III. vol. VI. p. 20. An account of the radiolarian cherts by Dr G. J. Hinde will be found in the Annals and Magazine of Natural History for July, 1890, p. 40.
[77] See Lapworth, C., "The Girvan Succession," Quart. Journ. Geol. Soc., vol. XXXVIII. p. 537, and also the paper on the Ballantrae Rocks referred to in the preceding footnote. The latter paper should be carefully read by all students of the stratigraphy of the Lower Palæozoic Rocks.
It is interesting to find that in the north of Ireland the rocks generally coincide in characters with those which are found along the same line of strike in Great Britain; thus, the Girvan type appears in Londonderry, Tyrone and Fermanagh, the Moffat type in County Down, and the Lake District type in the counties of Dublin and Kildare.
On the continent the volcanic material which plays so important a part in the constitution of the Ordovician accumulations of Britain is practically absent, and the strata are largely composed of accumulations of shale and limestone with occasional coarser deposits. In Scandinavia, the Arenig beds consist of limestones with a few shales, the Llandeilo deposits are largely calcareous, those of Caradoc age are partly calcareous and towards the top usually argillaceous, while the equivalents of the British Ashgill series are calcareous at the base and argillaceous at the summit. In Russia the calcareous matter preponderates over the argillaceous material.
Ordovician strata are also found in Belgium, France, Bohemia, and other places, and are largely composed of mechanical sediments of varying degrees of fineness mixed occasionally with some calcareous matter.
The variation in the characters of the Ordovician strata of Britain points to accumulation in a fairly deep sea, usually at some distance from the land, but dotted over with volcanoes which often rose above the water, causing the addition of much volcanic material to the ordinary sediments, and the existence of minor unconformities at different horizons along their flanks. As these unconformities are not always associated with volcanic material it is obvious that uplifts must have occurred occasionally during the deposition of the rocks; one important uplift is indicated by the occurrence of an unconformity in the Arenig rocks of Wales, while another is seen amongst the Caradoc rocks of the Welsh borders. On the whole, however, the period was one of slow subsidence, the deposition of material generally keeping pace with this subsidence, and accordingly there is a great uniformity of characters amongst the strata over wide areas. The probable continuation through the Ordovician period of the tract of land over the present site of the N. Atlantic ocean which as we have reason to suppose existed during Cambrian times, is indicated by similar changes of lithological character amongst the strata when traced from Britain eastward to Russia in both Cambrian and Ordovician times, and the continuance of these conditions over the American area is also indicated by study of the variations amongst the American Ordovician deposits.
The Ordovician Faunas. The Ordovician period has justly been termed the Period of Graptolites, which are the dominant forms of the time, and continue in abundance throughout the period. The abundance of graptolites in black shales associated with few other organisms has often been noted. It appears to be due to a large extent to the slow accumulation of the graptolitic deposits, allowing an abundance of these creatures to be showered upon the ocean floor, after death, for the evidence derived from detailed examination of their structure points to their existence as floating organisms. The tests of other creatures largely calcareous may well have been dissolved before reaching the sea-floor. In support of the view that these black shales are abysmal deposits may be noted the singular persistence of their lithological characters over wide areas, their replacement by much greater thicknesses of normal sediments along the ancient coast-lines, the frequent occurrence together of blind trilobites with those having abnormally large eyes when these creatures are associated with graptolites in the black shales, and lastly the interstratification of the black shales with radiolarian cherts similar to the modern abysmal radiolarian oozes. If this be so, we ought to find graptolites in marine deposits of all kinds, and indeed they are found there, though largely masked by the mass of sediment and the hosts of other included fossils, so that their discovery is rendered much more difficult than when they occur in the black shales,—a state of things which is familiar in the case of other pelagic organisms as Globigerinæ, radiolaria, and pteropods, whose tests abound in the abysmal deposits and are comparatively rare in those of terrigenous origin[78].
[78] The importance of the graptolites as indices of the geological age will be seen by perusal of Prof. Lapworth's paper "On the Geological Distribution of the Rhabdophora," Ann. and Mag. Nat. Hist., Ser. 5, vol. III. (1897).
The characters of the Ordovician trilobites have already been noticed. These organisms are abundant, and occur in sediments of all kinds. Of other groups, the significance of the radiolaria has been referred to above. Corals occasionally form reef-like masses of limestone as in the limestones of the Caradoc epoch; the echinoderms are well represented, cystids being locally abundant; of the crustacea, many remains of tests of phyllocarida have been recorded; the brachiopods are very abundant, and of the mollusca, lamellibranchs, gastropods and cephalopods all occur with frequency though none of these groups is very prevalent. Certain forms have been referred to pteropods though with doubt, and other shells seem to be referable to the heteropods. The existence of vertebrates during Ordovician times is not, in the opinion of many geologists, proved, though remains of fishes have been recorded from the Ordovician strata of North America; but it is desirable that more evidence of this occurrence should be given[79].
[79] Walcott, C. D., "Preliminary Notes on the Discovery of a Vertebrate Fauna in Silurian (Ordovician) Strata," Bulletin Geol. Soc. America, vol. III. p. 153.
The distribution of the Ordovician faunas like that of the sediments points to the prevalence of open ocean conditions over wide areas during the period, with occasional approaches to land, which was often of a volcanic nature. Around this land clustered the ordinary invertebrates, building up coral-reefs and shell-banks, whilst away in the open oceans the graptolites floated, almost alone, and sank to the ocean floor after death.
THE SILURIAN SYSTEM AND THE CHANGES WHICH OCCURRED IN BRITAIN AT THE CLOSE OF SILURIAN TIMES.
Classification. The Silurian system was originally divided by its founder, Sir R. I. Murchison, into three series, as follows:—
| Ludlow | Series |
| Wenlock | " |
| Llandovery | " |
The term May Hill, proposed by Sedgwick, is sometimes used as synonymous with Llandovery. This classification omits a somewhat important set of beds intercalated between those of the Llandovery and Wenlock series known as the Tarannon shales, and in Britain if we were to classify afresh, it would be more convenient to include some of the beds formerly referred to the Ludlow in the Wenlock. I shall, however, adopt the old and well-established classification, adding the term Tarannon to Llandovery, and speaking of the Llandovery-Tarannon series. The nature of the two classifications is shown in the following table:
| Stages. | Old Classification. | New Classification. | Palæontological Classification. |
|||
| 1 Upper Ludlow | Ludlow | Downtonian | Fauna with Encrinurus | |||
| 2 Aymestry Limestone | ||||||
| 3 Lower Ludlow | Salopian | |||||
| 4 Wenlock Limestone | Wenlock | |||||
| 5 Wenlock Shale | ||||||
| 6 Woolhope Limestone | ||||||
| 7 Tarannon Shales | Valentian | Fauna with Harpes | ||||
| 8 Upper Llandovery | Llandovery | |||||
| 9 Lower Llandovery | ||||||
Description of the strata. Lithologically the Silurian deposits of Britain form a continuation of those of the Ordovician period, with a local interruption due to the elevation of portions of Wales and the Welsh borders at the close of Ordovician times. Elsewhere we find a predominance of shales passing into grits at the top of the system, the change indicating the incoming of the shallow-water phase before the commencement of the second continental period. Particular stress is laid upon the predominant shaley character of the beds, for, on account of the richness and variety of the faunas of the calcareous rocks, greater attention is naturally paid to them in geological works, and the student may get a false idea of their relative importance. An attempt is made below (Fig. 18) to give a general idea of the variations in lithological characters of the Silurian rocks in different parts of Britain.
The Silurian strata are mostly found in the same localities as those which furnish exposures of the rocks of Ordovician age.
The development in the typical Silurian region of the Welsh borders is characterised by the abundance of calcareous matter which is found there as compared with that which exists in the other British localities.
The Llandovery strata are sandy, often conglomeratic, with a fair amount of calcareous matter in places. The arenaceous nature is undoubtedly due to the proximity of land caused by local upheaval at the end of Ordovician times, and the Upper Llandovery rocks sometimes rest unconformably on the Lower ones, at other times on Ordovician, Cambrian, or even Precambrian rocks. The Tarannon shales are light green shales with intercalated grits. The Wenlock series consists of a group of shales separating a lower, very inconstant, earthy limestone from an upper, more constant, thicker and purer limestone. The latter, the Wenlock limestone, is composed of fragments and perfect specimens of various fossils, and the fragmentary nature of many of the shells indicates the occurrence of wave-action and probable formation in shallow water, in some places against coral-reefs.
The Lower Ludlow beds consist of sandy shales; they are separated from the Upper Ludlow beds by an impure limestone, the Aymestry limestone. The Upper Ludlow beds consist mainly of grits and flags, often coloured red towards the summit.
In North Wales the Llandovery beds occasionally present the shelly arenaceous types of deposit as near Llangollen, at other times as near Conway, Corwen, and in Anglesey, the graptolitic shale type. They also rest unconformably upon the Ordovician rocks in this area. The Tarannon shales resemble those of the border county. The Wenlock series consists essentially of shales, while the Ludlow development differs from that of the borders in its greater thickness and the absence of any calcareous band in the centre. In Central Wales the graptolitic type of the Llandovery-Tarannon series is found, but the graptolite-bearing shales of the Llandovery epoch are thin beds occurring between grits and flags no doubt deposited in shallow water, and this division of the series is of very great thickness.
In South Wales the Silurian rocks are very similar to those of the Welsh borders, save that the calcareous deposits are fewer and thinner.
The Lake District Silurian strata generally resemble those of North Wales. The Llandovery-Tarannon rocks are of the graptolite-shale type, intercalated with fine grits in the case of the beds of Tarannon age. The Wenlock beds consist of shales, and the Ludlow beds of gritty shales beneath, and massive flags and grits at the summit. These Ludlow beds are here of great thickness (certainly not less than 7000 feet) and were obviously accumulated for the most part in shallow water.
The Llandovery-Tarannon rocks of Southern Scotland show the two types which prevailed in the Moffat and Girvan areas in later Ordovician times. The Llandovery beds of Moffat are known as the Birkhill shales, and are very thin. The representatives of the Tarannon shales, however, the Gala beds, consist mainly of grits, and attain a great thickness. In the Girvan area, the Llandovery beds are of the shelly type. Here as at Moffat and in the Lake District there is perfect conformity between the beds of Ordovician and those of Silurian age, and accordingly it is instructive to note the completeness of the palæontological break, especially in the Moffat district. The higher Silurian beds of Southern Scotland present a general resemblance to those of North Wales and the Lake District[80].