PLATE 26.
STRUCTURE OF THE MOUTH AND THORAX.
N.B.—The same letters of reference are used for the parts of the mouth
throughout this plate. All the organs here represented have been cleaned
by boiling in potash, and consist exclusively of the external membrane.
- Fig. 1, Mouth of Balanus perforatus, viewed from vertically above,
with the first pair of cirri x x in their proper position,
cut off close above their basal articulations.
- a, outer maxilla;
a′, ditto, cut closely off.
- b, maxilla.
- c, mandibles.
- d, palpus;
d′ ditto, cut closely off.
- e, labrum, crest of.
- x x, first pair of cirri, cut off.
- 2, Supra-œsophageal cavity of the mouth of Balanus improvisus,
torn open, with the palpi, mandibles, and maxillæ removed,
exhibiting the inner face of the labrum, laid flat, and the
inner faces of the outer maxillæ; scale as in fig. 1.
- a, outer maxillæ;
a′′, inner and lower lobe of ditto.
- d′, point of attachment to the torn-off palpus.
- e, labrum, crest off;
e′, central notch, with graduated teeth on both sides.
- f, triangular thickened portion of the inner fold of the
labrum;
f′, rib or bar of membrane, thickened to support
the surrounding very thin membrane of the supra-œsophageal
cavity.
- g, opening of the œsophagus.
- h, forked bar of thickened membrane, for same purpose
as f′.
- 3, Mouth of Coronula balænaris, divided vertically in a transverse
line, with the outer and inner maxillæ and mandible on
the left side removed, with the labrum, the other mandible
and both palpi viewed on the inside, and with the
œsophagus adherent in its proper position.
- c, Mandible, upper free segment;
c1, second or middle segment;
c2, supposed third or basal segment, forming the
basal margin of the mouth, but not separated by any articulation
from the labrum.
- d d, palpi;
d′, aperture, through which the inside of this
palpus can be seen, caused by the mandible having been cut
off; the membrane on the under side of this aperture is
thickened, and affords the chief support to the palpus;
d′′, thin membrane, apparently part of the palpus, connecting
the attached basal end of the palpus to the externo-lateral
surface of the mandible.
- e′, central notch in the crest of the labrum;
e′′ e′′, basal margin of the labrum, to which the membrane of the body
is joined.
- f, thickened portion of membrane, part of the inner fold of
the labrum, corresponding with f in fig. 2.
- g, orifice of the œsophagus, with the front part of the supra-œsophageal
cavity cut away;
g′, lower, expanded, or bell-shaped, end of the œsophagus, in its
natural condition.
- i, k, concavities for the attachment of muscles running to
the mandibles.
- m, cut off edge of membrane forming the side of the mouth,
which cut edge can be followed up above the aperture d′,
where the upper basal end of the palpus has been cut off,
and so onwards by g, to the mandible on the opposite
side of the mouth.
- 4, Mouth of Coronula balænaris, viewed from the outside,
in front, with the inner maxilla, mandible, and palpus on
the left side of the figure, and nearly the whole labrum, and
the greater part of the palpus on the right side, all cut
away. The maxilla, mandible, and basal portion of the palpus
on the right hand, are pulled out of their proper positions;
for the toothed edge of the mandible ought to have
stood higher, and nearly in a line between and behind the
outer maxillæ, and then the basal margin e′′ of the lateral
portion of the labrum would be raised, and curling round,
would stand nearly where the letter b′ is now placed.
- a a, outer maxilla, upper free lobe;
a′′, lower and inner free lobe;
a1, supposed second or middle segment;
a2, supposed third or basal segment;
a′′′, basal articulation of mouth, separating it from the
ventral surface of the thorax.
- b, inner maxilla;
b′, apodeme of do.
- c, mandible, upper free segment;
c1, second segment;
c2, third and basal segment, but not separated by an articulation
from the labrum.
- d, palpus;
d′′, membrane uniting the basal end of the palpus, where attached
to the labrum, to the side of the mandible.
- e′′, basal margin of the labrum, on one side, displaced and
pulled down.
- n, orifice of the olfactory pouch.
- 5, Balanus amaryllis: mandible, upper free segment, seen from
the side that faces the labrum;
p, an arched line, where the thickened membrane of the upper
free part terminates, and is united by thin membrane to the near
side of the palpus;
q, ligamentous bands for the attachment of muscles.
- 6, Balanus amaryllis: palpus, on the same scale as the mandible,
fig. 5, seen from the outside, so that the further or under
face is the side which, towards the right hand, is united
to the near face of the mandible, fig. 5;
r, long bristles, springing from a protuberance near the extremity;
these bristles, in many species, form a long single row, parallel
to the basal margin;
s, upper row of shorter bristles;
t, row of very short bristles, arising from the further side
and curling over the crest of the labrum.
- 7, Balanus amaryllis: maxilla on twice the scale of the mandible
and palpus;
b, upper free segment;
b′, apodeme;
b1, thickened portion of membrane, perhaps answering
to the second segment of the mandibles;
b2, thin membrane, extending down to the basal edge of the mouth,
possibly answering to the basal segment of the mandibles;
or perhaps the thickened membrane forming the protuberance
c2 in fig. 3, may be considered as the basal
segment of the maxillæ.
- u, step-formed projection at lower angle of maxilla.
- 8, Thorax of Coronula diadema: outer membrane of the five posterior
segments laid almost flat, and viewed externally as a
transparent object.
- 2 to 6, the second to the sixth cirrus, cut off a little above
their basal articulations.
- a, basal curved end of the probosciformed penis.
- b, anus;
b′, the membrane surrounding the anus, probably
consisting of a rudiment of the abdomen.
- c, rectum, seen through on the under side.
- d, basal articulation of the sixth cirrus;
d′, do. of the fifth cirrus;
d2, do. of the second cirrus.
- e e′, posterior thoracic segment, carrying the sixth pair of
cirri; the dorso-lateral portion e is separated, in this one
segment, from the corresponding portion e′ by a narrow
slip of thinner membrane, which may be a part of the
abdomen let in.
- e2 e2, thoracic segment, supporting the second pair of cirri.
- f, f3, folds or articulations between the adjoining segments.
- g, swollen, punctured portions of membrane, not differing
homologically from the rest of the segments.
- h, posterior portion of the prosoma, or the specially enlarged
part of the thorax; it carries the first pair of cirri; homologically
h forms part of the second segment of the
thorax of the archetype Crustacean.