Leaves crowded at the end of lateral branchlets or remote on vigorous shoots, linear-oblong, lanceolate, oval or obovate, acute or acuminate or sometimes contracted into a short broad point at apex, cuneate or occasionally rounded at base, entire, with slightly thickened margins, or rarely coarsely dentate, coated when they unfold with rufous tomentum, especially on the lower surface, or pubescent or sometimes nearly glabrous, and at maturity thick and firm, dark green and lustrous above, pale and often villose below, principally along the broad midrib and on the primary veins, 2′—5′ long and ½′—3′ wide; turning early in autumn bright scarlet on the upper surface only; petioles slender or stout, terete or wing-margined, ciliate, ¼′—1½′ in length, and often bright red. Flowers appearing in early spring when the leaves are about one third grown on slender pubescent or tomentose peduncles ½′—1½′ long, staminate in many-flowered dense or lax compound heads, pistillate in 2 to several-flowered clusters, sessile in the axils of conspicuous often foliaceous bracts, and furnished with 2 smaller acute hairy bractlets; calyx of the staminate flower disciform; petals thick, ovate-oblong, acute, rounded at apex, erect or slightly spreading, early deciduous; stamens exserted in the staminate flower, shorter than the petals in the pistillate flower; stigma stout, exserted, reflexed above the middle, 0 in the staminate flower. Fruit ripening in October, 1—3 from each flower-cluster, ovoid, ⅓′—⅔′ long, dark blue, with thin acrid flesh; stone light brown, ovoid, rounded at base, pointed at apex, terete or more or less flattened, and 10—12-ribbed, with narrow indistinct pale ribs rounded on the back.
A tree, with thick hard roots and few rootlets, often surrounded by root-sprouts, occasionally 100° or rarely 125° high, with a trunk sometimes 5° in diameter, numerous slender pendulous tough flexible branches forming a head sometimes short, cylindric and flat-topped, sometimes low and broad, or on trees crowded in the forest narrow, pyramidal or conic, and sometimes inversely conic and broad and flat at the top, and branchlets when they first appear light green to orange color, and in their first winter nearly glabrous or pale or rufous-pubescent, light red-brown marked by minute scattered pale lenticels and by small lunate leaf-scars displaying the ends of 3 conspicuous groups of fibro-vascular bundles, later becoming darker and developing short stout spur-like lateral branchlets; generally in the northern and extreme southern states much smaller, and rarely more than 50°-60° tall. Winter-buds obtuse, ¼′ long, with ovate acute apiculate dark red puberulous imbricated scales, those of the inner ranks accrescent, bright-colored at maturity, and marking the base of the branchlet with obscure ring-like scars. Bark of the trunk ¾′—1½′ thick, light brown often tinged with red, and deeply fissured, the surface of the ridges covered with small irregularly shaped scales. Wood heavy, soft, strong, very tough, not durable, light yellow or nearly white, with thick lighter colored sapwood of 80—100 layers of annual growth; used for the hubs of wheels, rollers in glass factories, ox-yokes, wharf-piles, and sometimes for the soles of shoes.
Distribution. Borders of swamps in wet imperfectly drained soil, and often especially southward on high wooded mountain slopes; valley of the Kennebec River, Maine, to southern Ontario, central Michigan, southeastern Missouri and eastern Oklahoma, and southward to northern Florida, and to the valley of the Brazos River, Texas; of its largest size on the southern Appalachian Mountains.
Occasionally cultivated as an ornamental tree in the eastern states, but difficult to transplant except when very young. The first tree in the eastern states to assume autumn colors of the leaves.
Leaves oblanceolate, oblong, elliptic or rarely ovate, acute or acuminate or occasionally rounded at the narrow apex, cuneate or rounded at the gradually narrowed base, and entire, when they unfold silky-villose above and hoary-tomentose beneath, soon becoming glabrous, dark yellow-green and lustrous on the upper surface, paler and sometimes glaucous on the lower surface, 2′—4′ long and ¾′—1′ wide, with a prominent midrib and numerous slender veins; petioles stout, ¼′—½′ in length. Flowers appearing when the leaves are nearly fully grown; staminate on slender villose pedicels, in many-flowered loose clusters on slender hairy peduncles 1′—1½′ in length; pistillate in pairs on rather stouter peduncles usually about 1′ long; calyx of the staminate flower disciform; petals oblong-ovate, rounded at apex, white, erect or slightly spreading, early deciduous. Fruit solitary or in pairs, on peduncles 1′—1½′ in length, oval or ellipsoid, dark blue, lustrous, about ⅓′ long, with acrid pulp; stone oval, compressed, narrowed at the ends, and prominently ribbed.
A tree, rarely more than 30° high, with a slender trunk gradually tapering upward from a swollen and much enlarged base, small spreading branches forming a narrow pyramidal or round-topped head, branchlets slightly villose when they first appear, soon glabrous, bright reddish brown in their first winter, becoming darker the following year, and numerous erect thick roots rising above the surface of the water. Winter-buds acute, dark red-brown, puberulous, and about ⅛′ long, the inner scales hoary-tomentose. Bark about 1′ thick, deeply furrowed, gray to very dark reddish brown.
Distribution. Small Pine-barren ponds of the coastal plain from North Carolina to central and eastern Florida, southern Alabama and Mississippi, and western Louisiana (near Lake Charles, Calcasieu Parish).
Leaves oblong, oval or obovate, acute, rounded or rarely obtuse, and apiculate at apex, gradually or abruptly cuneate or sometimes rounded at base, and entire, covered on the lower surface when they unfold with thick hoary tomentum and on the upper surface with short scattered pale hairs, and at maturity thick and firm, dark green, lustrous and slightly pilose above, pale below, 4′—6′ long and 2′—2½′ wide, with a stout midrib, 9 or 10 pairs of primary veins covered on the lower side with rufous pubescence or often nearly glabrous, and obscure reticulate veinlets; petioles stout, grooved, ½′—1′ in length. Flowers appearing in March and April; staminate in capitate clusters on slender hairy peduncles ½′ long, bibracteolate near the middle, and developed from the axils of the inner scales of the terminal bud, covered with long pale hairs on the outer surface of the short obscurely 5-toothed cup-shaped calyx and on the oblong petals rounded at apex; filaments longer than the petals; anthers oval and conspicuously tuberculate-roughened; pistillate solitary, 1/16′ long, on short stout woolly peduncles from the axils of bud-scales, and furnished at apex with 2 acute hairy bractlets; calyx coated, like the minute rounded spreading petals, with hoary tomentum; stamens included, with short filaments, and small mostly fertile anthers; style stout, exserted, reflexed from near the base. Fruit bright or dull red, on slender tomentose stems enlarged at apex and ½′—⅔′ long, ripening in July and August, and sometimes persistent on the branches until after the falling of the leaves, oblong or obovoid, 1′—1½′ in length, tipped with the thickened and pointed remnants of the style; flesh thick, juicy, very acid; stone oblong, compressed, narrowed at the ends, rounded at base, acute at apex, with walls produced into 10 or 12 broad thin papery white wings, about 1′ long, and 1 or rarely 2-seeded.
A tree, rarely 60°—70° high, with 1 or several stems occasionally 2° in diameter, spreading branches forming a narrow round-topped head, and slender branchlets coated when they first appear with rufous tomentum, light reddish brown or green tinged with red and puberulous during their first summer, turning gray or reddish brown in their first winter, and marked by large lunate or nearly triangular leaf-scars displaying the ends of 3 groups of fibro-vascular bundles; often a shrub, with numerous slender clustered diverging stems. Winter-buds obtuse, ⅛′ long, with ovate apiculate imbricated scales rounded on the back and clothed with thick hoary tomentum, those of the inner ranks becoming at maturity ovate-oblong or obovate, rounded at apex, bright red, and ½′—¾′ long. Bark of the trunk about ⅛′ thick, irregularly fissured, with a dark brown surface broken into thick appressed persistent plate-like scales. Wood light, soft, tough, not strong, white, with thin hardly distinguishable sapwood of about 10 layers of annual growth. A preserve with an agreeable subacid flavor, known as Ogeechee limes, is sometimes made from the fruit in Georgia and South Carolina. The flowers abound in nectar, and are much visited by bees.
Distribution. Deep often inundated river swamps or their borders; South Carolina in the neighborhood of the coast, through the valley of the lower Ogeechee River, Georgia; in northern and in western Florida to the mouth of the Choctawhatchee River (R. H. Harper), and in the valley of the lower Apalachicola River; rare and local.
Leaves oblong-ovate, acute or acuminate and often long-pointed at apex, cuneate, rounded, or subcordate at base, entire or remotely and irregularly angulate-toothed, the teeth often tipped with a long slender mucro, when they unfold light red and coated below and on the petioles with pale tomentum and pubescent above, especially on the broad thick midrib, and at maturity thick and firm, dark green and lustrous on the upper surface, pale and more or less downy-pubescent on the lower surface, 5′—7′ long and 2′—4′ wide, with 10—12 pairs of primary veins forked near the margins and connected by conspicuous cross veins; petioles stout, grooved, hairy, enlarged at base, 1½′—2½′ in length. Flowers appearing in March and April on a long slender hairy peduncle from the axil of an inner scale of the terminal bud; staminate in dense capitate clusters, their peduncle furnished near the middle and occasionally at apex with long linear ciliate bractlets; calyx-tube cup-shaped, obscurely 5-toothed, one third as long as the oblong erect petals rounded at apex and much shorter than the stamens; pistillate solitary, surrounded by 2—4 strap-shaped scarious ciliate bractlets often ½′ long and more or less united below into an involucral cup; calyx-tube oblong and much longer than the ovate minute spreading petals; stamens included, with small mostly fertile anthers; style stout, tapering, reflexed above the middle, and revolute into a close coil. Fruit ripening early in the autumn, on slender drooping stalks 3′—4′ in length, oblong or slightly obovoid, crowned with the pointed remnants of the style, dark purple, marked by conspicuous scattered pale dots, and 1′ long, with thick tough skin and thin acid flesh; stone obovoid, rounded at the narrow apex, pointed at base, flattened, light brown or nearly white, and about 10-ridged, the ridges acute and wing-like, with thin separable margins, and sometimes united by short intermediate ridges.
A tree, 80°—100° high, with a trunk 3°—4° in diameter above the greatly enlarged tapering base, comparatively small spreading branches forming a narrow oblong or pyramidal head, stout pithy branchlets dark red and coated with pale tomentum when they first appear, soon becoming glabrous or nearly so, and in their first winter light or bright red-brown and marked by small scattered pale lenticels and by the conspicuous elevated nearly orbicular leaf-scars displaying the ends of 3 large fibro-vascular bundles, and thick corky roots. Winter-buds: terminal nearly globose, with broad ovate light chestnut-brown scales keeled on the back and rounded and apiculate at apex, those of the inner ranks accrescent and at maturity oblong-ovate or oblong-obovate, rounded at apex, 1′ or more long, and bright yellow; axillary minute, obtuse, nearly imbedded in the bark. Bark of the trunk about ¼′ thick, dark brown, longitudinally furrowed, and roughened on the surface by small scales. Wood light, soft, not strong, close-grained, difficult to split, light brown or often nearly white, with thick sapwood sometimes composed of more than 100 layers of annual growth; used in the manufacture of wooden-ware, broom-handles, and wooden shoes, and largely for fruit and vegetable boxes. The wood of the roots is sometimes employed instead of cork for the floats of nets.
Distribution. Deep swamps inundated during a part of every year; coast region of the Atlantic states from southeastern Virginia to northern Florida, through the Gulf states to the valley of the Nueces River, Texas, and through Arkansas and southern and southeastern Missouri to western Kentucky and Tennessee, and to the valley of the lower Wabash River, Illinois; of its greatest size in the Cypress-swamps of western Louisiana and eastern Texas.
Trees or shrubs, with terete branchlets, scaly buds, and alternate or opposite deciduous leaves, without stipules. Flowers perfect or polygamo-diœcious; calyx 4 or 5-toothed, petals 4 or 5; stamens inserted on the margin of the epigynous disk; anthers oblong; introrse, 2-celled, the cells opening longitudinally; ovary 1 or 2-celled; ovule solitary, suspended from the interior angle of the apex of the cell, anatropous; micropyle superior. Fruit drupaceous, 1 or 2-seeded. Seed oblong-ovoid; seed-coat membranaceous; embryo in copious fleshy albumen; cotyledons foliaceous; radicle terete, turned toward the hilum.
The widely distributed Cornel family with ten genera, more numerous in temperate than in tropical regions, has arborescent representatives of the genus Cornus in North America.
Trees and shrubs, with astringent bark, opposite or rarely alternate deciduous leaves conduplicate or involute in the bud. Flowers small, perfect, white, greenish white or yellow; calyx-tube minutely 4-toothed, the teeth valvate in the bud; disk pulvinate, depressed in the centre, or obsolete; petals 4, valvate in the bud, oblong-ovate, inserted on the margin of the disk; stamens 4, alternate with the petals; filaments slender, exserted; ovary 2-celled; style exserted, simple, columnar, crowned with a single capitate or truncate stigma; raphe dorsal. Fruit ovoid or oblong; flesh thin and succulent; nut bony or crustaceous, 2-celled, 2 or sometimes 1-seeded. Seed compressed; embryo straight or slightly incurved.
Cornus with nearly fifty species is widely distributed through the three continents of the northern hemisphere, and south of the equator is represented in Peru by a single species. Of the sixteen or seventeen species of the United States four are arborescent. Cornus is rich in tannic acid, and the bark and occasionally the leaves and unripe fruit are used as tonics, astringents, and febrifuges. Of exotic species, Cornus mas, L., is often planted in the eastern states as an ornamental tree, and its edible fruit is used in Europe in preserves and cordials. The wood of Cornus is hard, close-grained, and durable, and is used in turnery and for charcoal.
The generic name, from cornu, relates to the hardness of the wood produced by plants of this genus.
Leaves ovate to elliptic or rarely slightly obovate, acute and often contracted into a slender point at apex, gradually narrowed at base, remotely and obscurely crenulate-toothed on the somewhat thickened margins, and mostly clustered at the end of the branches, when they unfold pale and pubescent below and puberulous above, and at maturity thick and firm, bright green and covered with minute appressed hairs on the upper surface, pale or sometimes almost white and more or less pubescent on the lower surface, 3′—6′ long and 1½′—2′ wide, with a prominent light-colored midrib deeply impressed above, and 5 or 6 pairs of primary veins connected by obscure reticulate veinlets; in the autumn turning bright scarlet on the upper surface, remaining pale on the lower surface; petioles grooved, ½′—¾′ in length. Flowers: head of flower-buds appearing during the summer between the upper pair of lateral leaf-buds, inclosed by 4 involucral scales remaining light brown and more or less covered with pale hairs during the winter, and borne on a stout club-shaped puberulous peduncle ¼′ long or less during the winter and becoming 1′—1½′ in length; involucral scales beginning to unfold, enlarge and grow white in early spring and when the flowers open in March at the south to May at the north, when the leaves are nearly fully grown, forming a flat corolla-like cup 3′—4′ in diameter, becoming at maturity obovoid, 1′—1½′ wide, gradually narrowed below the middle and notched at the rounded apex, reticulate-veined, pure white, pink, or rarely bright red, deciduous after the fading of the flowers; flowers in dense many-flowered cymose heads, in the axils of broad-ovate nearly triangular minutely apiculate glabrous light green deciduous bracts, ⅛′ in diameter; calyx terete, slightly urceolate, puberulous, obtusely 4-lobed, light green; corolla-lobes strap-shaped, rounded or acute at apex, slightly thickened on the margins, puberulous on the outer surface, reflexed after anthesis, green tipped with yellow; disk large and orange-colored; style crowned with a truncate stigma. Fruit ripening in October, ovoid, crowned with the remnants of the narrow persistent calyx and with the style, bright scarlet or rarely yellow (f. xanthocarpa Rehd.), lustrous, ½′ long and ¼′ broad, with thin mealy flesh, and a smooth thick-walled slightly grooved stone acute at the ends, and 1 or 2-seeded; seeds oblong, pale brown.
A bushy tree, rarely 40° high, with a short trunk 12′-18′ in diameter, slender spreading or upright branches, and divergent branchlets turning upward near the end, pale green or green tinged with red when they first appear, glabrous or slightly puberulous, bright red or yellow-green during their first winter and nearly surrounded by the narrow ring-like leaf-scars, later becoming light brown or gray tinged with red; frequently toward the northern limits of its range a much-branched shrub. Winter-buds formed in midsummer; the terminal covered by 2 opposite acute pointed scales rounded on the back and joined below for half their length, and accompanied by 2 pairs of lateral buds, each covered by a single scale, those of the lower pair shedding their scales in the autumn and remaining undeveloped. Bark of the trunk ⅛′—¼′ thick, with a dark red-brown surface divided into quadrangular or many-sided plate-like scales. Wood heavy, hard, strong, close-grained, brown sometimes changing to shades of green and red, with lighter colored sapwood of 30—40 layers of annual growth; largely used in turnery, for the bearings of machinery, the hubs of small wheels, barrel-hoops, the handles of tools, and occasionally for engravers’ blocks.
Distribution. Usually under the shade of taller trees in rich well-drained soil; southern Maine to southern Ontario, southern Michigan, southeastern Kansas and eastern Oklahoma, and southward to central Florida and the valley of the Brazos River, Texas; on the mountains of northern Mexico; comparatively rare at the north; one of the commonest and most generally distributed inhabitants of the deciduous-leaved forests of the middle and southern states, ranging from the coast nearly to the summits of the high Alleghany Mountains. Trees with rose-colored or with pink involucral scales occasionally occur (var. rubra André). A variety with pendulous branches is known in gardens (var. pendula Dipp.); the var. xanthocarpa near Oyster Bay, Nassau County, Long Island, New York, and at Saluda, Polk County, North Carolina.
Often planted as an ornament of parks and gardens in the eastern states.
Leaves ovate or slightly obovate, acute and often contracted into a short point at the apex, cuneate at base, faintly crenulate-serrate, and generally clustered toward the end of the branches, when they unfold coated below with pale tomentum and puberulous above, and at maturity thin, bright green and slightly puberulous, with short appressed hairs on the upper surface, and woolly pubescent on the lower surface, 4′—5′ long and 1½′—3′ wide, with a prominent midrib impressed above, and about 5 pairs of slender primary veins connected by remote reticulate veinlets; in the autumn turning bright orange and scarlet before falling; petioles stout, grooved, pubescent, ½′—⅔′ in length, with a large clasping base. Flowers: head of flower-buds appearing during the summer between the upper pair of lateral leaf-buds, surrounded at base but not inclosed by the involucral scales during the winter, hemispheric, ½′ in diameter, usually nodding on a stout hairy peduncle ¾′—1′ long; involucral scales becoming when the flowers open 1½′—3′ long and 1½′—2′ wide, white or white tinged with pink, oblong to obovate or nearly orbicular, and acute, acuminate, or obtuse, entire and thickened at apex, puberulous on the outer surface, gradually narrowed below the middle and conspicuously 8-ribbed, the spreading ribs united by reticulate veinlets; flowers in dense cymose heads from the axils of minute acuminate scarious deciduous bracts; calyx terete, slightly urceolate, puberulous on the outer surface, yellow-green, or light purple, with dark red-purple lobes; petals strap-shaped, rounded at apex, spreading, somewhat puberulous on the outer surface, with thickened slightly inflexed margins, yellow-green; style crowned with a truncate stigma. Fruit ripening in October, in dense spherical heads of 30—40 drupes surrounded at base by a ring of abortive pendulous ovaries, ½′ long, ovoid, much flattened, crowned with the broad persistent calyx, bright red or orange-colored, with thin mealy flesh, and a thick-walled 1 or 2-seeded stone obtuse at the ends and scarcely grooved; seeds oblong, compressed, with a very thin pale papery coat.
A tree, 40°—60°, or exceptionally 100° high, with a trunk 1°—2° in diameter, small spreading branches forming an oblong conic or ultimately round-topped head, and slender light green branchlets coated while young with pale hairs, becoming glabrous or puberulous, dark reddish purple or sometimes green during their first winter and conspicuously marked by the elevated lunate leaf-scars, ultimately becoming light brown or brown tinged with red. Winter-buds formed in July; the terminal acute, ⅓′ long, covered by 2 narrow-ovate acute long-pointed puberulous light green opposite scales, accompanied by 2 pairs of lateral buds, each covered by a single scale, those of the lower pair shedding their scales in the autumn and remaining undeveloped, those of the upper pair clothed with pale hairs, especially toward the apex, their scales thickening, turning dark purple, lengthening in the spring with the inclosed shoot, finally becoming scarious and developing into small leaves, and in falling marking the base of the branchlets with ring-like scars. Bark of the trunk about ¼′ thick, brown tinged with red, and divided on the surface into small thin appressed scales. Wood heavy, exceedingly hard, strong, close-grained, light brown tinged with red, with lighter colored sapwood of 30—40 layers of annual growth; used in cabinet-making, for mauls and the handles of tools.
Distribution. Usually in moist well-drained soil under the shade of coniferous forests; valley of the lower Fraser River and Vancouver Island, British Columbia, southward through western Washington and Oregon, on the coast ranges of California to the San Bernardino Mountains, and on the western slopes of the Sierra Nevada; southward up to altitudes of 4000°—5000°, of its largest size near the shores of Puget Sound and in the Redwood-forests of northern California.
Leaves ovate or oblong, gradually or abruptly contracted at apex into a long slender point, gradually narrowed or rounded and cuneate at base, and slightly thickened on the undulate margins, coated with lustrous silvery tomentum when they unfold, and nearly fully grown when the flowers open from the middle of May in Texas to the middle of July at the north, and then dark green and roughened above by short rigid white hairs, and pale, often glaucous or rough-pubescent below, and at maturity thin, scabrous on the upper surface, pubescent or puberulous on the lower surface, 3′—4′ long and 1½′—2′ wide, with a thin midrib, and 4—6 pairs of slender primary veins parallel with their sides; petioles stout, grooved, pubescent, usually about ½′ in length. Flowers cream color, on slender pedicels, in loose broad or narrow often panicled pubescent cymes, on peduncles frequently 1′ in length; calyx oblong, cup-shaped, obscurely toothed, covered with fine silky white hairs; corolla-lobes narrow-oblong, acute, about ⅛′ long, and reflexed after the flowers open; style thickened at apex into a prominent stigma. Fruit ripening from the end of August until the end of October, in loose spreading red-stemmed clusters, subglobose, white, tipped with the remnants of the style, about ¼′ in diameter, with thin dry, bitter flesh, and a full and rounded stone broader than high, somewhat oblique, slightly grooved on the edge, and 1 or 2-seeded; seeds nearly ¼′ long, with a pale brown coat.
A tree, sometimes nearly 50° high, with a short trunk 8′—10′ in diameter, thin erect wand-like branches forming a narrow irregular rather open head, and slender branchlets marked by numerous small pale lenticels, light green and puberulous when they first appear, pale red, lustrous, and puberulous during their first winter, light reddish brown in their second year, and ultimately light gray-brown or gray; usually shrubby. Winter-buds acute, compressed, pubescent, sessile, or stalked, about ⅛′ long, with 2 pairs of opposite scales, the terminal bud nearly twice as large as the compressed lateral buds. Bark of the trunk about ⅛′ thick, and divided by shallow fissures into narrow interrupted ridges broken into small closely appressed dark red-brown scales. Wood close-grained, hard, pale brown, with thick cream-colored sapwood.
Distribution. Southwestern Ontario (Point Pelee and Pelee Island), southward through Ohio, Kentucky, Tennessee and Mississippi to western Florida (Gadsden and Levy Counties) and westward to southeastern South Dakota, southeastern Nebraska, central Kansas, northwestern Oklahoma (near Alva, Woods County) and western Texas (Kerr, Menard and Brown Counties); probably only arborescent on the rich bottom-lands of southern Arkansas and eastern Texas.
Leaves mostly alternate, clustered at the end of the branches, rarely opposite, oval or ovate, gradually contracted at apex into a long slender point, cuneate or occasionally somewhat rounded at base, obscurely crenulate-toothed on the slightly thickened and incurved margins, coated when they unfold on the lower surface with dense silvery white tomentum, and faintly tinged with red and pilose above, and at maturity thin, bright yellow-green, glabrous or sparsely pubescent on the upper surface, pale or sometimes nearly white and covered with appressed hairs on the lower surface, 3′—5′ long and 2½′—3½′ wide, with a broad orange-colored midrib slightly impressed above, and about 6 pairs of primary veins parallel with their sides; in the autumn turning yellow or yellow and scarlet; petioles slender, pubescent, grooved, 1½′—2′ in length, with an enlarged clasping base. Flowers cream color, opening from the beginning of May to the end of June on slender jointed pedicels ⅛′—¼′ long, in terminal flat puberulous many-flowered cymes 1½′—2½′ wide, mostly on lateral branchlets; calyx cup-shaped, obscurely toothed; corolla-lobes narrow, oblong, rounded at apex, ⅛′ long, reflexed after anthesis; style enlarged into a prominent stigma. Fruit in loose spreading red-stemmed clusters, ripening in October, subglobose, dark blue-black, or rarely yellow (f. ochrocarpa Rehd.), ⅓′ in diameter, tipped with the remnants of the style rising from the bottom of a small depression, with thin and bitter flesh; and an obovoid nutlet, pointed at base, gradually longitudinally many-grooved, thick-walled, and 1 or 2-seeded; seeds lunate, ¼′ long, with a thin membranaceous pale coat.
A flat-topped tree, rarely 25°—30° high, with a short trunk 6′—8′ in diameter, long slender alternate diverging horizontal branches, and numerous short upright slender branchlets pale orange-green or reddish brown when they first appear, mostly light green or sometimes brown tinged with green during their first winter, later turning darker green and marked by pale lunate leaf-scars and small scattered pale lenticels; often a shrub, with numerous stems. Bark of the trunk about ⅛′ thick, dark reddish brown, and smooth or divided by shallow longitudinal fissures into narrow ridges irregularly broken transversely. Wood heavy, hard, close-grained, brown tinged with red, with thick lighter colored sapwood of 20—30 layers of annual growth.
Distribution. Rich woodlands, the margins of the forest, and the borders of streams and swamps, in moist well-drained soil, New Brunswick and Nova Scotia, westward along the valley of the St. Lawrence River to the northern shores of Lake Superior and to Minnesota, and southward through the northern states and along the Appalachian Mountains to North Carolina, up to altitudes of 3500°—4000°; in northern Alabama, southwestern Georgia, and western Florida (River Junction, Gadsden County, T. G. Harbison).
Occasionally cultivated as an ornamental plant in the eastern states.
Section 2. Gamopetalæ. Corolla of united petals (divided in Elliottia in Ericaceæ 0 in some species of Fraxinus in Oleaceæ).
A. Ovary superior (inferior in Vaccinium in Ericaceæ, partly inferior in Symplocaceæ, partly superior in Styraceæ).
Trees or shrubs, with scaly buds, and alternate simple leaves, without stipules. Flowers perfect, regular; calyx 4—5-lobed; corolla hypogynous, 5-lobed (of 4 petals in Elliottia), the lobes imbricated in the bud; stamens hypogynous, mostly free from the corolla, as many, or twice as many as its lobes; anthers introrse, 2-celled, opening by terminal pores, often appendaged; ovary 4—10-celled (inferior in Vaccinium); styles terminal, simple, stigma terminal; ovules numerous, anatropous or amphitropous; raphe ventral; micropyle superior. Fruit capsular, drupaceous, or baccate. Seeds with fleshy or horny albumen, embryo small; cotyledons small and short.
The Heath family with seventy-one genera is widely distributed over the temperate and tropical parts of the earth’s surface. Of the twenty-one genera found in the United States seven have arborescent representatives.
A glabrous tree or shrub, with slender terete branchlets, scaly buds, and fibrous roots. Leaves petiolate, oblong or oblong-obovate, acute at the ends or occasionally rounded at apex, entire, thin, dark green and glabrous above, pale and villose below, particularly on the thin yellow midrib and obscure forked veins; deciduous; petioles slender and flattened, with an abruptly enlarged base nearly covering the small axillary buds. Flowers perfect, on slender elongated pedicels, in erect terminal elongated racemose panicles, with minute acute scarious caducous bracts and bractlets; calyx short, tubular, puberulous, dark red-brown, 4-toothed, the broad apiculate teeth erose on the margins and imbricated in the bud; petals 4, imbricated in the bud, spatulate-linear, sessile; stamens 8, hypogynous, shorter than the petals, filaments broad, flattened; anthers oblong-ovoid, the cells callous-mucronate, free at the apex of the spreading lobes, opening from above downward; disk much thickened, fleshy; ovary sessile, subglobose, 4-lobed, 4-celled, concave at apex; style elongated, slender, gradually enlarged and club-shaped above and incurved at apex; stigma 3—5-lobed, smaller than the thickened end of the style; ovules numerous in each cell, attached on the inner angle of a tumid placenta, ascending, anatropous. Fruit unknown.
Elliottia with a single species is confined to the southern United States.
The genus is named in honor of Stephen Elliott (1771—1830), the distinguished botanist of South Carolina.
Leaves 3′—4′ long, 1′—1½′ wide; petioles ⅓′—½′ in length. Flowers about ½′ long, opening from the middle to the end of June, in clusters 7′—10′ in length.
A tree, 15°—20° high, with a trunk 4′—5′ in diameter, short ascending branches forming a pyramidal head, and erect branchlets light red-brown and pilose when they first appear, bright orange-brown, lustrous, and nearly glabrous during their first winter, and roughened by slightly raised oblong-obovate leaf-scars with conspicuous central fibro-vascular bundle-scars, becoming light brown slightly tinged with red during their second season and dark gray-brown the following year; or more frequently shrubby. Winter-buds: terminal broad-ovoid, acute, about ⅛′ long, with much thickened bright chestnut-brown shining scales conspicuously white-pubescent near the margins toward the apex; lateral buds smaller, ovoid, compressed, rounded or short-pointed at apex. Bark thin, smooth, pale gray.
Distribution. Sandy woods in a few isolated stations in the valley of the Savannah River, near Augusta, Richmond County, and in Burke and Bullock Counties, Georgia.
Trees or shrubs, with scaly bark, terete branchlets, terminal buds formed in summer, and fibrous roots. Leaves usually clustered at the end of the branches, revolute and entire on the margin, persistent or deciduous. Flowers in terminal umbellate corymbs from buds with numerous caducous scales; calyx 5-parted or toothed, persistent under the fruit, corolla 5—10-lobed, deciduous; stamens 5 or 10, rarely more, more or less unequal, ultimately spreading; filaments subulate-filiform, pilose at the base; disk thick and fleshy, crenately lobed; ovary 5—10-celled; style slender, crowned with a capitate stigma and persistent on the fruit; ovules numerous in each cell, attached in many series to an axile 2-lipped placenta projected from the inner angle of the cell, anatropous. Fruit a woody many-seeded capsule. Seed scobiform; seed-coat loose, reticulate, produced at the ends beyond the nucleus into a short often laciniate appendage; embryo minute, cylindric, axile in fleshy albumen; cotyledons oblong, shorter than the radicle turned toward the hilum.
Rhododendron with some four or five hundred species occurs in eastern Thibet, on the Himalayas, in southwestern China, the Malay peninsula and Archipelago, New Guinea, northern China and Corea, Japan, the mountains of central Europe, on the Caucasus, and in eastern and western North America, the largest number of species being found in southwestern China and on the Himalayas. Of the twenty-three or twenty-four North American species one only is arborescent.
Rhododendron possesses astringent narcotic properties. It produces hard close-grained compact wood sometimes used in turnery and for fuel. Many of the species are cultivated in gardens for the beauty of their large and conspicuous flowers.
The generic name is from ῥόδον and δένδρον, the Rose-tree.
Leaves revolute in the bud, ovate-lanceolate or obovate-lanceolate, acute or short-pointed at apex, and narrowed, cuneate or rounded at base, when they unfold covered with a thick pale or ferrugineous tomentum of gland-tipped hairs, and at maturity glabrous, thick and coriaceous, dark green and lustrous on the upper surface, usually pale or whitish on the lower surface, 4′—12′ long and 1½′—2½′ wide, with a broad pale midrib and obscure reticulate veinlets; persistent for two or three years; petioles stout, ridged above, rounded below, 1′—1½′ in length. Flowers: inflorescence-buds surrounded at first by several loose narrow leaf-like scales, and when fully grown in September cone-shaped, 1½′ long and ½′ broad, with many imbricated ovate scales rounded and contracted at apex into a long slender point, opening late in June after the shoots of the year from buds in the axils of upper leaves have reached their full length; flowers on slender pink pedicels covered with glandular white hairs and furnished at base with two linear scarious bractlets, from the axils of the scales of the inner ranks of the inflorescence-bud, in 16—24-flowered umbellate clusters 4′—5′ in diameter, with accrescent scarious resinous puberulous bracts, those of the outer ranks becoming 1′ long and ⅓′ wide, and shorter than the lanceolate bracts of the inner ranks contracted into a long slender point; calyx light green and puberulous, with rounded remote lobes; corolla prominently 5-angled or ridged in the bud, campanulate, gibbous on the posterior side, puberulous in the throat, light rose color, purplish, or white, 1′ long, cleft to the middle into 5 oval rounded lobes, with conspicuous central veins, the upper lobe marked on the inner face by a cluster of yellow-green spots, and furnished on the outer surface at the bottom of each sinus with a conspicuous dark red gland; stamens 8—12, white, inserted on the bright green disk; filaments enlarged and flattened at base, slightly bent inward above the middle, and bearded with stiff white hairs, the 4 or 5 short ones at the back of the flower for more than half their length and the others only near the base; ovary ovoid, green, coated with short glandular pale hairs, crowned with a long slender glabrous white declining style club-shaped and inflexed at apex, and terminating in a 5-rayed scarlet stigma. Fruit dark red-brown, ovoid, ½′ long, glandular-hispid, ripening and shedding its seeds in the autumn, the clusters of open capsules remaining on the branches until the following summer; seeds oblong, flattened, the coat prolonged at the ends into scarious fringed appendages.
A bushy tree, 30°—40° high, with a short crooked often prostrate trunk occasionally 10′—12′ in diameter, stout contorted branches forming a round head, and branchlets green tinged with red and covered with dark red or slightly ferrugineous glandular-hispid hairs when they first appear, dark green and glabrous in their first winter, gradually turning bright red-brown in their second year, and ultimately gray tinged with red, the thin bark separating on branches four or five years old into persistent scales; more often a broad shrub, with many divergent twisted stems 10°—12° high. Winter-buds: leaf-buds conic, dark green, axillary, or terminal on barren shoots, with many closely imbricated scales, those of the inner ranks accrescent, increasing in length from the outer to the inner, and at maturity 1½′ long, ¼′ wide, gradually narrowed at base, and terminating at apex in a long slender point, light green, glabrous, closely held against the shoot by a resinous exudation from the glandular hairs, and in falling marking the branchlet with numerous conspicuous narrow remote scars persistent for three or four years. Bark of the trunk about 1/16′ thick, light red-brown, broken on the surface into small thin appressed scales. Wood heavy, hard, strong, rather brittle, close-grained, light clear brown, with thin lighter colored sapwood; occasionally made into the handles of tools and used as a substitute for boxwood in engraving. A decoction of the leaves is occasionally employed in domestic practice in the treatment of rheumatism.
Distribution. Nova Scotia, Mt. Chocorua, New Hampshire, and southward in New England and eastern New York and along the Appalachian Mountains to northern Georgia and westward to the northern shores of Lake Erie and to southeastern Ohio (Hocking and Fairfield Counties); rare at the north and an inhabitant of deep cold swamps in a few isolated stations; more abundant on the mountains of western Pennsylvania, becoming exceedingly common farther south and occupying the steep banks of streams up to altitudes of 3000°; of its largest size on the high mountains of eastern Tennessee and the Carolinas, and here often forming thickets hundreds of acres in extent.
Often cultivated as an ornament of parks and gardens in the United States, and in Europe, and one of the parents of a number of distinct and beautiful hybrids.
Trees or shrubs, with scaly bark, terete branchlets without a terminal bud, minute axillary leaf-buds, elongated axillary inflorescence-buds covered by imbricated scales, and fibrous roots. Leaves ovate-oblong or linear, short-petiolate, with flat entire margins, coriaceous, persistent or deciduous in one species. Flowers on slender pedicels bibracteolate at the base, from the axils of foliaceous coriaceous ovate or acute persistent bracts, in axillary umbels; calyx 5, rarely 6-parted, the divisions imbricated in the bud, persistent under the fruit; corolla 5, rarely 6-lobed, rose-colored, purple, or white, saucer-shaped, with a short tube and 10 pouches just below the 5 or 6-parted limb, the lobes ovate, acute, before anthesis prominently 10 or 12-ribbed from the pouches to the acute apex of the bud, the salient keel of the ribs running to the point of the lobes and to the sinuses; stamens 10, shorter than the corolla; filaments filiform; anthers oblong, each cell opening by a short apical oblong longitudinal pore, at first free in the bud, the filaments then erect, later received in the pouches of the corolla, the filaments becoming bent back by its enlargement and expansion, straightening elastically and incurving on the release of the anthers, and in straightening discharging the pollen-grains; disk prominently 10-lobed; ovary subglobose, 5-celled; style filiform, exserted, crowned with a capitate stigma; ovules numerous in each cell, inserted on a 2-lipped placenta, pendulous or spreading from near the top of the thin columella, few-ranked, anatropous. Fruit a woody many-seeded globose slightly 5-lobed 5-celled capsule, tardily septicidally 5-valved, the valves crustaceous, ultimately opening down the middle by a narrow slit and separating from the persistent placenta-bearing axis. Seeds oblong or subglobose, minute; seed-coat crustaceous or membranaceous; embryo in fleshy albumen, terete, near the hilum; radicle erect, rather shorter than the oblong cotyledons.
Kalmia with six species is North American and Cuban, one species occasionally becoming under favorable conditions a small tree.
The generic name is in honor of the Swedish traveler and botanist, Peter Kalm (1715—1779).