It may be well to pause here a moment to show how this mistaken identification of evolution with progress alone, without modification by the more fundamental laws of differentiation, has given rise to misconceptions in the popular and even in the scientific mind. The biologist is continually met with the question, “Do you mean to say that any one of the invertebrates, such, for instance, as a spider, may eventually, in the course of successive generations, become a vertebrate, or that a dog or a monkey is on the highway to become a man?” By no means. There is but one straight and narrow way to the highest in evolution as in all else, and few there be that have found it—in fact, probably two or three only at every step. The animals mentioned above have diverged from that way. In their ancestral history, they have missed the golden opportunity, if they ever had it. It is easy to go on in the way they have chosen, but impossible to get back on the ascending trunk-line. To compare again with the growing tree, only one straight trunk-line leads upward to the terminal bud. A branch once separated must grow its own way, if it grow at all.

Of the same nature is the mistake of some extreme evolutionists, such as Dr. Bastian and Professor Haeckel, and of nearly all anti-evolutionists, viz., that of imagining that the truth of evolution and that of spontaneous generation must stand or fall together. On the contrary, if life did once arise spontaneously from any lower forces, physical or chemical, by natural process, the conditions necessary for so extraordinary a change could hardly be expected to occur but once in the history of the earth. They are, therefore, now, not only unreproducible, but unimaginable. Such golden opportunities do not recur. Evolution goes only onward. Therefore, the impossibility of the derivation of life from non-life now, is no more an argument against such a derivation once, than is the hopelessness of a worm ever becoming a vertebrate now, an argument against the derivative origin of vertebrates. Doubtless if life were now extinguished from the face of the earth, it could not again be rekindled by any natural process known to us; but the same is probably true of every step of evolution. If any class—for example, mammals—were now destroyed, it could not be re-formed from any other class now living. It would be necessary to go back to the time and conditions of the separation of this class from the reptilian stem. Therefore, the falseness of the doctrine of abiogenesis,7 so far from being any argument against evolution, is exactly what a true conception of evolution and knowledge of its laws would lead us to expect.

c. Law of Cyclical Movement.—The movement of evolution has ever been onward and upward, it is true, but not at uniform rate in the whole, and especially in the parts. On the contrary, it has plainly moved in successive cycles. The tide of evolution rose ever higher and higher, without ebb, but it nevertheless came in successive waves, each higher than the preceding and overborne by the succeeding. These successive cycles are the dynasties or reigns of Agassiz, and ages of Dana; the reign of mollusks, the reign of fishes, of reptiles, of mammals, and finally of man. During the early Palæozoic times (Cambrian and Silurian) there were no vertebrates.8 But never in the history of the earth were mollusks of greater size, number, and variety of form than then. They were truly the rulers of these early seas. In the absence of competition of still higher animals, they had things all their own way, and therefore grew into a great monopoly of power. In the later Palæozoic (Devonian) fishes were introduced. They increased rapidly in size, number, and variety; and being of higher organization they quickly usurped the empire of the seas, while the mollusca dwindled in size and importance, and sought safety in a less conspicuous position. In the Mesozoic times, reptiles, introduced a little earlier,9 finding congenial conditions and an unoccupied place above, rapidly increased in number, variety, and size, until sea and land seem to have swarmed with them. Never before or since have reptiles existed in such numbers, in such variety of form, or assumed such huge proportions; nor have they ever since been so highly organized as then. They quickly became rulers in every realm of Nature—rulers of the sea, swimming reptiles; rulers of the land, walking reptiles; and rulers of the air, flying reptiles. In the unequal contest, fishes therefore sought safety in subordination. Meanwhile mammals were introduced in the Mesozoic, but small in size, low in type (marsupials), and by no means able to contest the empire with the great reptiles. But in the Cenozoic (Tertiary) the conditions apparently becoming favorable for their development, they rapidly increased in number, size, variety, and grade of organization, and quickly overpowered the great reptiles, which almost immediately sank into the subordinate position in which we now find them, and thus found comparative safety. Finally, in the Quaternary, appeared man, contending doubtfully for a while, with the great mammals, but soon (in Psychozoic) acquiring mastery through superior intelligence. The huge and dangerous mammals were destroyed and are still being destroyed; the useful animals and plants were preserved and made subservient to his wants; and all things on the face of the earth are being readjusted to the requirements of his rule. In all cases it will be observed that the rulers were such because, by reason of strength, organization, and intelligence, they were fittest to rule. There is always room at the top. To illustrate again by a growing tree: This successive culmination of higher and higher classes may be compared to the flowering and fruiting of successively higher and higher branches. Each uppermost branch, under the genial heat and light of direct sunshine, received in abundance by reason of position, grew rapidly, flowered, and fruited; but quickly dwindled when overshadowed by still higher branches, which, in their turn, monopolized for a time the precious sunshine.

But observe, furthermore: when each ruling class declined in importance, it did not perish, but continued in a subordinate position. Thus, the whole organic kingdom became not only higher and higher in its highest forms, but also more and more complex in its structure and in the interaction of its correlated parts. The whole process and its result is roughly represented in the accompanying diagram, Fig. 1, in which A B represents the course of geological time and the curve, the rise, culmination, and decline of successive dominant classes.

MOLLUSCS—Silurian.
FISHES—Devon. and Carb.
REEPTILES—Mesozoic.
MAMMALS—Tert’y and Quat.
MAN—Present.

Fig. 1.

The Above Three Laws Are Laws of Evolution.

These three laws we have shown are distinctly recognizable in the succession of organic forms in the geological history of the earth. They are, therefore, undoubtedly the general laws of succession. Are they also laws of evolution? Are they also discoverable in embryonic development, the type of evolution? They are, as we now proceed to show:

Differentiation.—In reproduction the new individual appears: 1. As a germ-cell—a single microscopic living cell. 2. Then, by growth and multiplication of cells, it becomes an egg. This may be characterized as an aggregate of similar cells, and therefore is not yet differentiated into tissues and organs. In other words, it is not yet visibly organized; for organization may be defined as the possession of different parts, performing different functions, and all co-operating for one given end, viz., the life and well-being of the organism. 3. Then commences the really characteristic process of development, viz., differentiation or diversification. The cells are at first all alike in form and function, for all are globular in form, and each performs all the functions necessary for life. From this common point now commences development in different directions, which may be compared to a branching and rebranching, with more and more complex results, according as the animal is higher in the scale of organization and advances toward a state of maturity. First, the cell-aggregate (egg) separates into three distinct layers of cells, called ecto-blast, endo-blast, and meso-blast. These by further differentiation form the three fundamental groups of organs and functions, viz., the nervous system, the nutritive system, and the blood system: the first presiding over the exchange of force or influence, by action and reaction with the environment, and between the different parts of the organism; the second presiding over the exchange of matter with the environment, by absorption and elimination; the third presiding over exchanges of matter between different parts of the organism. The first system of functions and organs may be compared to a system of telegraphy, foreign and domestic; the second to foreign commerce; the third to an internal carrying-trade. Following out any one of these groups in higher animals, say the nervous system, it quickly differentiates again into two sub-systems, viz., cerebro-spinal and ganglionic, each having its own distinctive functions, which we can not stop to explain. Then the cerebro-spinal again differentiates into voluntary and reflex systems. All of these have meanwhile separated into sensory and motor centers and fibers. Then, taking only the sensory fibers, these again are differentiated into five special senses, each having a wholly different function. Then, finally, taking any one of these, say the sense of touch or feeling, this again is differentiated into many kinds of fibers, each responding to a different impression, some to heat, others to cold, still others to pressure, etc. We have taken the nervous system; but the same differentiation and redifferentiation takes place in all other systems, and is carried to higher and higher points according to the position in the scale of the animal which is to be formed.

Or, to vary the mode of presentation a little, the cells of the original aggregate, commencing all alike, immediately begin to take on different forms, in order to perform different functions. Some cells take on a certain form and aggregate themselves to form a peculiar tissue which we call muscle, and which does nothing else, can do nothing else, than contract under stimulus. Another group of cells take on another peculiar form and aggregate themselves to form another and very different tissue, viz., nervous tissue, which does nothing and can do nothing but carry influence back and forth between the great external world and the little world of consciousness within. Still another group of cells take still another form and aggregate to form still another tissue, viz., the epithelial, whose only function is to absorb nutritive and eliminate waste matters. Thus, by differentiation of form and limitation of function, or division of labor, the different parts of the organism are bound more and more closely together by mutual dependence, and the whole becomes more and more distinctly individuated, and separation of parts becomes more and more a mutilation, and finally becomes impossible without death. This process, as already said, reaches its highest point only in the later stages of development of the highest animals.

Progress.—The law of progress is, of course, admitted to be a law of ontogeny; but observe here, also, it is true only of the whole and not necessarily of all the parts, except from the point of view of the whole. Thus, for example, starting all from a common form or generalized type, some cells advance to the dignity of brain-cells, whose function is somehow connected with the generation or at least the manifestation of thought, will, and emotion; other cells descend to the position of kidney-cells, whose sole function is the excretion of urine. But here, also, the highest cells are successively higher, and the whole aggregate is successively nobler and more complex. It is again a branching and rebranching, in every direction, some going upward, some downward, some horizontally, anywhere, everywhere, to increase the complexity of relations internal and external, and therefore to elevate the plane of the whole.

Cyclical Movement.—Lastly, the law of cyclical movement is also a law of ontogeny and therefore of evolution. This law, however, is less fundamental than the other two, and is, therefore, less conspicuous in the ontogenic than in the phylogenic series. It is conspicuous only in the later stages of ontogeny, and in other higher kinds of evolution, such as social evolution. For example, in the ontogenic development of the body and mind from childhood to manhood we have plainly successive culminations and declines of higher and higher functions. In bodily development we have culminating first the nutritive functions, then the reproductive and muscular, and last the cerebral. In mental development we have culmination first of the receptive and retentive faculties in childhood, then of imaginative and æsthetic faculties in youth and young manhood; then of the reflective and elaborative faculties—the faculties of productive work in mature manhood; and, finally, the moral and religious sentiments in old age. The first gathers and stores materials; the second vivifies and makes them plastic building materials; the third uses them in actual constructive work—in building the temple of science and philosophy; and the fourth dedicates that temple only to noblest purposes.

Observe here, also, that when each group of faculties culminates and declines, it does not perish, but only becomes subordinate to the next higher dominant group, and the whole psychical organism becomes not only higher and higher in its highest parts, but also more and more complex in its structure and in the interaction of its correlated parts.

Observe, again, the necessity laid upon us by this law—the necessity of continued evolution to the end. Childhood, beautiful childhood, can not remain—it must quickly pass. If, with the decline of its characteristic faculties, the next higher group characteristic of youth do not increase and become dominant, then the glory of life is already past and deterioration begins. Have we not all seen sad examples of this? Youth, glorious youth, must also pass. If the next higher group of reflective and elaborative faculties do not arise and dominate, then progressive deterioration of character commences here—thenceforward the whole nature becomes coarse, as we so often see in young men, or else shrivels and withers, as we so often see in young women. Finally, manhood, strong and self-relying manhood, must also pass. If the moral and religious sentiments have not been slowly growing and gathering strength all along, and do not now assert their dominance over the whole man, then commences the final and saddest decline of all, and old age becomes the pitiable thing we so often see it. But, if the evolution have been normal throughout; if the highest moral and religious nature have been gathering strength through all, and now dominates all, then the psychic evolution rises to the end—then the course of life is like a wave rising and cresting only at the moment of its dissolution, or, like the course of the sun, if not brightest at least most glorious in its setting. And thus—may we not hope?—the glories of the close of a well-spent life become the pledge and harbinger of an eternal to-morrow?

We have thus far illustrated the three laws of succession of organic forms by ontogeny, because this is the type of evolution; but they may be illustrated also by other forms of evolution. Next to the development of the individual, undoubtedly the progress of society furnishes the best illustration of these laws.

Commencing with a condition in which each individual performs all necessary social functions, but very imperfectly; in which each individual is his own shoemaker and tailor, and house-builder and farmer, and therefore all persons are socially alike; as society advances, the constituent members begin to diverge, some taking on one social function and some another, until in the highest stages of social organization this diversification or division and subdivision of labor reaches its highest point, and each member of the aggregate can do perfectly but one thing. Thus, the social organism becomes more and more strongly bound together by mutual dependence, and separation becomes mutilation. I do not mean to say that this extreme is desirable, but only that an approach to this is a natural law of social development. Is not this the law of differentiation?

So also progress is here, as in other forms of evolution—a progress of the whole, but not necessarily of every part. Some members of the social aggregate advance upward to the dignity of statesmen, philosophers, and poets; some advance downward to the position of scavengers and sewer-cleansers.10 But the highest members are progressively higher, and the whole aggregate is progressively grander and more complex in structure and functions.

So, again, the law of cyclical movement is equally conspicuous here. Society everywhere advances, not uniformly, but by successive waves, each higher than the last; each urged by a new and higher social force, and embodying a new and higher phase of civilization. Again: as each phase declines, its characteristic social force is not lost, but becomes incorporated into the next higher phase as a subordinate principle, and thus the social organism as a whole becomes not only higher and higher, but also more and more complex in the mutual relations of its interacting social forces.

Let us not be misunderstood, however. There is undoubtedly in social evolution something more and higher than we have described, but which does not concern us here, except to guard against misconstruction. There is in society a voluntary progress wholly different from the evolution we have been describing. In true or material evolution natural law works for the betterment of the whole utterly regardless of the elevation of the individual, and the individual contributes to the advance of the whole quite unconsciously while striving only for his own betterment. This unconscious evolution by natural law inherited from the animal kingdom is conspicuous enough in society, especially in its early stages, but we would make a great mistake if we imagined, as some do, that this is all. Besides the unconscious evolution by natural laws, inherited from below, there is a higher evolution, inherited from above, indissolubly connected with man’s spiritual nature—a conscious, voluntary striving of the best members of the social aggregate for the betterment of the whole—a conscious, voluntary striving both of the individual and of society toward a recognized ideal. In the one kind of evolution the fittest are those most in harmony with the environment, and which therefore always survive; in the other, the fittest are those most in harmony with the ideal, and which often do not survive. The laws of this free voluntary progress are little understood. They are of supreme importance, but do not specially concern us here. We will speak of it again in another chapter.

The three laws above mentioned might be illustrated equally well by all other forms of evolution. We have selected only those which are most familiar. They may, therefore, be truly called the laws of evolution. We have shown that they are the laws of succession of organic forms.

III. Change by Means of Resident Forces.—Thus far in our argument I suppose that most well-informed men will raise no objection. It will be admitted, I think, even by those most bitterly opposed to the theory of evolution, that there has been throughout the whole geological history of the earth an onward movement of the organic kingdom to higher and higher levels. It will be admitted, also, that there is a grand and most significant resemblance between the course of development of the organic kingdom and the course of embryonic development—between the laws of succession of organic forms and the laws of ontogenic evolution. But there is another essential element in ontogenic evolution. It is that the forces or causes of evolution are natural; that they reside in the thing developing and in the reacting environment. This we know is true of embryonic development; is it true also of the geologic succession of organic forms? It is true of ontogeny; is it true also of phylogeny? If not, then only by a metaphor can we call the process of change in the organic kingdom throughout geological history an evolution. This is the point of discussion, and not only of discussion, but, alas! of heated and even angry dispute. The field of discussion is thus narrowed to this third point only.

Before stating the two opposite views of the cause of evolution, it is necessary to remind the reader that when the evolutionist speaks of the forces that determine progressive changes in organic forms as resident or inherent, all that he means, or ought to mean, is that they are resident in the same sense as all natural forces are resident; in the same sense that the vital forces of the embryo are resident in the embryo, or that the forces of the development of the solar system according to the nebular or any other cosmogonic hypotheses are resident in that system. In other words, they mean only that they are natural, not supernatural. This does not, of course, touch that deeper, that deepest of all questions, viz., the essential nature and origin of natural forces; how far they are independent and self-existent, and how far they are only modes of divine energy. This is a question of philosophy, not of science. This question is briefly discussed in another place (Part III, Chap. III); it does not immediately concern us here.

The Two Views briefly Contrasted.—As already stated, all will admit a grand resemblance between the stages of embryonic development and those of the development of the organic kingdom. This was first brought out clearly by Louis Agassiz, and is, in fact, the greatest result of his life-work. All admit, also, that the embryonic development is a natural process. Is the development of the organic kingdom also a natural process? All biologists of the present day contend that it is; all the old-school naturalists, with Agassiz at their head, and all anti-evolutionists of every school, contend that it is not. We take Agassiz as the type of this school, because he has most fully elaborated and most distinctly formulated this view. As formulated by him, it has stood in the minds of many as an alternative and substitute for evolution.

According to the evolutionists, all organic forms, whether species, genera, families, orders, classes, etc., are variable, and, if external conditions favor, these variations accumulate in one direction and gradually produce new forms, the intermediate links being usually destroyed or dying out. According to Agassiz, the higher groups, such as genera, families, orders, etc., are indeed variable by the introduction of new species, but species are the ultimate elements of classification, and, like the ultimate elements of chemistry, are unchangeable; and, therefore, the speculations of the evolutionist concerning the transmutation of species are as vain as were the speculations of the alchemists concerning the transmutation of metals—that the origin of man, for example, from any lower species is as impossible as the origin of gold from any baser metal. Both sides admit frequent change of species during geological history, but one regards the change as a change by gradual transmutation of one species into another through successive generations and by natural process, the other as change by substitution of one species for another by direct supernatural creative act. Both admit the gradual development of the organic kingdom as a whole through stages similar to those of embryonic development; but the one regards the whole process as natural, and therefore strictly comparable to embryonic development, the other as requiring frequent special interference of creative energy, and therefore comparable rather to the development of a building under the hand and according to the preconceived plan of an architect—a plan, in this case, conceived in eternity and carried out consistently through infinite time. It is seen that the essential point of difference is this: The one asserts the variability of species (if conditions favor, and time enough is given) without limit; the other asserts the permanency of specific forms, or their variability only within narrow limits. The one asserts the origin of species by “descent with modifications”; the other, the origin of species by “special act of creation.” The one asserts the law of continuity (i. e., that each stage is the natural outcome of the immediately preceding stage) in this, as in every other department of Nature; the other asserts that the law of continuity (i. e., of cause and effect) does not hold in this department; that the links of the chain of changes are discontinuous, the connection between them being intellectual, not physical.

So much for sharp contrasting characterization of the two views, necessary for clear understanding of much that follows. We will have to give them more fully hereafter when we take up the evidences of evolution in Part II.


CHAPTER II.
THE RELATION OF LOUIS AGASSIZ TO THE THEORY OF EVOLUTION.

In order to clear up the conception of evolution, it is necessary to give a brief history of the idea, and especially to explain the relation of Louis Agassiz to that theory. This is the more necessary, because there is a deep and wide-spread misunderstanding on this subject, and thus scant justice has been done our great naturalist, especially by the English and Germans; and also because this relation is an admirable illustration of an important principle in scientific philosophy.

Like all great ideas, we find the first germs of this in Greek philosophy, in the cosmic speculations of Thales and Pythagoras. Next (about 100 B. C.) we find it more clearly expressed by the Roman thinker, Lucretius, in his great philosophic poem entitled “De Rerum Natura.” After a dormancy of nearly eighteen centuries it next emerges with still more clearness in the theological speculations of Swedenborg and the philosophical speculations of Kant. All these we pass over with bare mention, because these thinkers approached the subject from the philosophic rather than the scientific side—in the metaphysical rather than the scientific spirit.

The first serious attempt at scientific presentation of the subject was by the celebrated naturalist, Lamarck, in a work entitled “Philosophie Zoölogique,” published in 1809. It is not necessary, in this rapid sketch, to give a full account of Lamarck’s views. Suffice it to say that the essential idea of evolution, viz., the indefinite variability and the derivative origin of species, was insisted on with great learning and skill, and illustrated by many examples. With Lamarck, the factors of evolution or causes of change of organic forms were—1. Modification of organs in function and therefore in structure, by a changing environment—external factor; and, 2. Modification of organs by use and disuse—internal factor. In both cases the modifications are inherited and increased from generation to generation, without limit. This second factor seems to have taken, in the mind of Lamarck, the somewhat vague and transcendental form of aspiration or upward striving of the animal toward higher conditions. These are acknowledged to-day as true factors of evolution, but the distinctively Darwinian factor, viz., “divergent variation and natural selection,” was not then thought of. The publication of Lamarck’s views produced a powerful impression, but only for a little while. Pierced by the shafts of ridicule shot by nimble wits of Paris, and crushed beneath the heavy weight of the authority of Cuvier, the greatest naturalist and comparative anatomist of that or perhaps of any time, it fell almost still-born. I believe it was best that it should thus perish. Its birth was premature; it was not fit to live. The world was not yet prepared for a true scientific theory. Nevertheless, the work was not without its effect upon some of the most advanced thinkers of that time; upon Saint-Hilaire and Comte in France, and upon Goethe and Oken in Germany. It was good seed sown and destined to spring up and bear fruit in suitable environment; but not yet.

The next attempt worthy of attention in this rapid sketch is that of Robert Chambers, in a little volume entitled “Vestiges of a Natural History of Creation,” published in 1844. It was essentially a reproduction of Lamarck’s views in a more popular form. It was not a truly scientific work nor written by a scientific man. It was rather an appeal from the too technical court of science to the supposed wider and more unprejudiced court of popular intelligence. It was therefore far more eloquent than accurate; far more specious than profound. It was, indeed, full of false facts and inconsequent reasonings. Nevertheless, it produced a very strong impression on the thinking, popular mind. But it also quickly fell, pierced by keen shafts of ridicule, and crushed beneath the heavy weight of the authority of all the most prominent naturalists of that time, with Agassiz at their head. The question for the time seemed closed. I believe, again, it was best so, for the time was not yet fully ripe.

I know full well that many think with Haeckel that biology was kept back half a century by the baneful authority of Cuvier and Agassiz; but I can not think so. The hypothesis was contrary to the facts of science as then known and understood. It was conceived in the spirit of baseless speculation, rather than of cautious induction; of skillful elaboration rather than of earnest truth-seeking. Its general acceptance would have debauched the true spirit of science. I repeat it: the time was not yet ripe for a scientific theory. The ground must first be cleared and a solid foundation built; an insuperable obstacle to hearty rational acceptance must first be removed, and an inductive basis must be laid.

The Obstacle removed.—The obstacle in the way of the acceptance of the derivative origin of species was the then prevalent notion concerning the nature of life. We must briefly sketch the change which has taken place in the last forty years in our ideas on this subject.

Until about forty years ago, the different forces of Nature, such as gravity, electricity, magnetism, light, heat, chemical affinity, etc., were supposed to be entirely distinct. The realm of Nature was divided up into a number of distinct and independent principalities, each subject to its own sovereign force and ruled by its own petty laws. About that time it began to be evident, and is now universally acknowledged, that all these forces are but different forms of one, universal, omnipresent energy, and are transmutable unto one another back and forth without loss. This is the doctrine of correlation of forces and conservation of energy, one of the grandest ideas of modern times. But one force seemed still to be an exception. Life-force was still believed to be a peculiar, mysterious principle or entity, standing above other forces and subordinating them; not correlated with, not transmutable unto, nor derivable from, other and lower forces, and therefore in some sense supernatural. Now, if this be true of living forces, it is perfectly natural, yea, almost necessary, to believe that living forms are wholly different from other forms in their origin. New forms of dead matter may be derived, but new living forms are underived. Other new forms come by natural process, new organic forms by supernatural process. The conclusion was almost unavoidable. But soon vital force also yielded to the general law of correlation of natural forces. Vital forces are also transmutable into and derivable from physical and chemical forces. Sun-force, falling on the green leaves of plants, is absorbed and converted into vital force, disappears as light to reappear as life. The amount of life-force generated is measured by the amount of light extinguished. The same is true of animal life. As in the steam-engine the locomotive energy is derived from the fuel consumed and measured by its amount, so in the animal body, the animal heat and animal force are derived from and measured by the food and tissue consumed by combustion. Thus, vital force may be regarded as so much force withdrawn from the general fund of chemical and physical forces, to be again refunded without loss at death. This obstacle is, therefore, now removed. If vital force falls in the same category as other natural forces, there is no reason why living forms should not fall into the same category in this regard as other natural forms. If new forms of dead matter are derived from old forms by modification, according to physical laws, there is no reason why new living forms should not also be derived from old forms by modification according to physiological laws. Thus, at last, the obstacle was removed—the ground was cleared.

The Basis laid.—But Science is not content with removal of a priori objections. She must also have positive proofs. The ground must not only be cleared, but a true inductive basis of facts, and especially of laws and methods, must be laid. This was the life-work of Agassiz. Yes, as strange as it may seem to some, it is nevertheless true that the whole inductive basis, upon which was afterward built the modern theory of evolution, was laid by Agassiz, although he himself persistently refused to build upon it any really scientific superstructure. It is plain, then, that all attempts at building previous to Agassiz’s work must, of necessity, have resulted in an unsubstantial structure—an edifice built on sand, which could not and ought not to stand. I must stop here in order to explain somewhat fully this important point, and thus to give due credit to the work of Agassiz.

The title of any scientist to greatness must be determined, not so much by the multitude of new facts he has discovered as by the new laws he has established, and especially by the new methods he has inaugurated or perfected. Now, I think it can be shown that to Agassiz, more than to any other man, is due the credit of having established the laws of succession of living forms in the geological history of the earth—laws upon which must rest any true theory of evolution. Also, that to him, more than to any other man, is due the credit of having perfected the method (method of comparison) by the use of which alone biological science has advanced so rapidly in modern times. This is high praise. I wish to justify it. I begin with the method.

Scientific methods bear the same relation to intellectual progress that tools, instruments, machines, mechanical contrivances of all sorts, bear to material progress. They are intellectual contrivances—indirect ways of accomplishing results far too hard for bare-handed, unaided intellectual strength. As the civilized man has little or no advantage over the savage in bare-handed strength of muscle, and the enormous superiority of the latter in accomplishing material results is due wholly to the use of mechanical contrivances or machines; even so, in the higher sphere of intellect, the scientist makes no pretension to the possession of greater unaided intellectual strength than belongs to the uncultured man, or even perhaps to the savage. The amazing intellectual results achieved by science are due wholly to the use of intellectual contrivances or scientific methods. As in the lower sphere of material progress the greatest benefactors of the race are the inventors or perfecters of new mechanical contrivances or machines, so also in the higher sphere of intellectual progress the greatest benefactors of the race are the inventors or perfecters of new intellectual contrivances or methods of research.

To illustrate the power of methods, and the necessity of their use, take the case of the method of notation, so characteristic of mathematics, and take it even in its simplest and most familiar form: Nine numeral figures, having each a value of its own, and another dependent upon its position; a few letters, a and b, and x and y, connected by symbols, + and-and =: that is all. And yet, by the use of this simple contrivance, the dullest school-boy accomplishes intellectual results which would defy the utmost efforts of the unaided strength of the greatest genius. And this is only the simplest tool-form of this method. Think of the results accomplished by the use of the more complex machinery of the higher mathematics!

Take next the method of experiment so characteristic of physics and chemistry. The phenomena of the external world are far too complex and far too much affected by disturbing forces and modifying conditions to be understood at once by bare, unaided intellectual insight. They must first be simplified. The physicist, therefore, contrives artificial phenomena under ideal conditions. He removes one complicating condition after another, one disturbing cause and then another, watching meanwhile the result, until finally the necessary condition and the true cause are discovered. On this method rests the whole fabric of the physical and chemical sciences.

But when we rise still higher, viz., into the plane of life, the phenomena of Nature become still more complex and difficult to understand directly; and yet just here, where we are the most powerless without some method, our method of experiment almost wholly fails us. The phenomena of life are not only far more complex than those of dead matter, but the conditions of life are so nicely adjusted, the equilibrium of forces so delicately balanced, that, when we attempt to introduce our clumsy hands in the way of experiment, we are in danger of overthrowing the equilibrium, of destroying the conditions of the experiment, viz., life; and then the whole problem falls immediately into the domain of chemistry. What shall we do? In this dilemma we find that Nature herself has already prepared for us, ready to hand, an elaborate series of simplified conditions equivalent to experiments. The phenomena of life are, indeed, far too complex to be at once understood—the problem of life too hard to be solved—in the higher animals; but, as we go down the animal scale, complicating conditions are removed one by one, the phenomena of life become simpler and simpler, until in the lowest microscopic cell or spherule of living protoplasm we finally reach the simplest possible expression of life. The equation of life is reduced to its simplest terms, and now, if ever, we begin to understand the true value of the unknown quantity. This is the natural history series, or Taxonomic series, already spoken of on page 10. Again, Nature has prepared, and is now preparing daily before our eyes, another series of gradually simplified conditions. Commencing with the mature condition of one of the higher animals—for example, man—and going backward along the line of individual history through the stages of infant embryo, egg and germ, we find again the phenomena of life becoming simpler and simpler, until we again reach the simplest conceivable condition in the single microscopic cell or spherule of living protoplasm. This, as already explained, is the embryonic or Ontogenic series. Again, that there be no excuse for man’s ignorance of the laws of life, Nature has prepared still another series; and this the grandest of all, for it is the cause of both the others. Commencing with the plants and animals of the present epoch, and going back along the track of geological times, through Cenozoic, Mesozoic, Palæozoic, Eozoic, to the very dawn of life—the first syllable of recorded time—and we find again a series of organic forms growing simpler and simpler, until, if we could find the very first, we would undoubtedly again reach the simplest condition in the lowest conceivable forms of life. This, as we have already seen, is the geologic or evolution, or Phylogenic series. We have already explained these three series, only in this connection it suits our purpose to take the terms backward.

Now, it is by comparison of the terms of each of these series going up and down, and watching the first appearance, the growth, and the perfecting of tissues, organs, functions, and by the comparison of the three series with one another term by term—I say it is wholly by comparison of this kind that biology has in recent times become a true inductive science. This is the “method of comparison.” It is the great method of research in all those departments which can not be readily managed by the method of experiment. It has already regenerated biology, and is now applied with like success in sociology under the name of historic method. Yes; anatomy became scientific only through comparative anatomy, physiology through comparative physiology, and embryology through comparative embryology. May we not add, sociology will become truly scientific only through comparative sociology, and psychology through comparative psychology?

Now, while it is true that this method, like all other methods, has been used, from the earliest dawn of thought, in a loose and imperfect way, yet it is only in very recent times that it has been organized, systematized, perfected, as a true scientific method, as a great instrument of research; and the prodigious recent advance of biology is due wholly to this cause. Now, among the great leaders of this modern movement, Agassiz undoubtedly stands in the very first rank. I must try to make this point plain, for it is by no means generally understood.

Cuvier is acknowledged to be the great founder of comparative anatomy. He it was that first perfected the method of comparison, but comparison only in one series—the Taxonomic. Von Baer and Agassiz added to this comparison in the ontogenic series also, and comparison of these two series with each other, and therefore the application of embryology to the classification of animals. If Von Baer was the first announcer, Agassiz was the first great practical worker by this method. Last and most important of all, in its relation to evolution, Agassiz added comparison in the geologic or phylogenic series. The one grand idea underlying Agassiz’s whole life-work was the essential identity of the three series, and therefore the light which they must shed on one another. The two guiding and animating principles of his scientific work were—1. That the embryonic development of one of the higher representatives of any group repeated in a general way the terms of the Taxonomic series in the same group, and therefore that embryology furnished the key to a true classification; and, 2. That the succession of forms and structure in geological times in any group is similar to the succession of forms and structure in the development of the individual in the same group, and thus that embryology furnishes also the key to geological succession. In other words, during his whole life, Agassiz insisted that the laws of embryonic development (ontogeny) are also the laws of geological succession (phylogeny). Surely this is the foundation, the only solid foundation, of a true theory of evolution. It is true that Agassiz, holding as he did the doctrine of permanency of specific types, and therefore rejecting the doctrine of the derivative origin of species, did not admit the causal or natural relation of phylogenic succession to embryonic succession and taxonomic order as we now believe it—it is true that for him the relation between the three series was an intellectual not a physical one—consisted in the preordained plans of the Creator, and not in any genetic connection or inherited property; but evidently the first and greatest step was the discovery of the relation itself, however accounted for. The rest was sure to follow.

But more. Not only did Agassiz establish the essential identity of the geologic and embryonic succession, the general similarity of the two series, phylogenic and ontogenic, but he also announced and enforced all the formal laws of geologic succession (i. e., of evolution), as we now know them. These, as already stated and illustrated, are the law of differentiation, the law of progress of the whole, and the law of cyclical movement, although he did not formulate them in these words. No true inductive evidence of evolution was possible without the knowledge of these laws, and for this knowledge we are mainly indebted to Agassiz. He well knew also that they were the laws of embryonic development and therefore of evolution; but he avoided the word evolution, as implying the derivative origin of species, and used instead the word development, though it is hard to see in what the words differ. Thus, it is evident that Agassiz laid the whole foundation of evolution, solid and broad, but refused to build any scientific structure on it; he refused to recognize the legitimate, the scientifically necessary outcome of his own work. Nevertheless, without his work a scientific theory of evolution would have been impossible. Without Agassiz (or his equivalent), there would have been no Darwin.

There is something to us supremely grand in this refusal of Agassiz to accept the theory of evolution. The opportunity to become the leader of modern thought, the foremost man of the century, was in his hands, and he refused, because his religious, or, perhaps better, his philosophic intuitions, forbade. To Agassiz, and, indeed, to all men of that time, to many, alas! even now, evolution is materialism. But materialism is Atheism. Will some one say, the genuine Truth-seeker follows where she seems to lead whatever be the consequences? Yes; whatever be the consequences to one’s self, to one’s opinions, prejudices, theories, philosophies, but not to still more certain truth. Now, to Agassiz, as to all genuine thinkers, the existence of God, like our own existence, is more certain than any scientific theory, than anything can possibly be made by proof. From his standpoint, therefore, he was right in rejecting evolution as conflicting with still more certain truth. The mistake which he made was in imagining that there was any such conflict at all. But this was the universal mistake of the age. A lesser man would have seen less clearly the higher truth and accepted the lower. A greater man would have risen above the age, and seen that there was no conflict, and so accepted both. All thinking men are coming to this conclusion now, but none had done so then.

Now, then, at last, the obstacle of supernaturalism in the realm of Nature having been removed by the establishment of the doctrine of correlation of natural forces, and the extension of this doctrine to embrace also life-force; and now also a broad and firm basis of carefully-observed facts and well-established laws of succession of organic forms having been laid by Agassiz, when again, for the third time, the doctrine of origin of species “by derivation with modifications” was brought forward by Darwin in a far more perfect form, with more abundant illustrative materials, and with a new and most potent factor of modification—viz., divergent variations and natural selection—it found the scientific world already fully prepared, and anxiously waiting. I say anxiously waiting—for the supposed supernatural origin of species had been the one exception to the otherwise universal law of cause and effect, or the law of continuity. It was therefore in open contradiction to the whole drift of scientific thought for five hundred years. Is it any wonder, then, that the derivative origin of species was welcomed with joy by the scientific world? For five hundred years, scientific thought, like a rising tide which knows no ebb, had tended thitherward with ever-increasing pressure, but kept back by the one supposed fact of the supernatural origin of species. Darwin lifted the gate, and the in-rushing tide flooded the whole domain of thought.

What, then, is the place of Agassiz in biological science? What is the relation of Agassiz to Darwin—of Agassizian development to Darwinian evolution? I answer, it is the relation of formal science to physical or causal science. Agassiz advanced biology to the formal stage; Darwin carried it forward, to some extent at least, to the physical stage. All true inductive sciences in their complete development pass through these two stages. Science in the one stage treats of the laws of phenomena; in the other, of the causes or explanation of these laws. The former must precede the latter, and form its foundation; the latter must follow the former, and constitute its completion. The change from the one to the other is always attended with prodigious impulse to science.

To illustrate: Until Kepler, astronomy was little more than an accumulation of disconnected facts concerning celestial motions—abundant materials, but no science; piles of brick and stone, but no building. Kepler reduced this chaos to beautiful order and musical harmony by the discovery of the three great laws which bear his name, and therefore he has been justly called the legislator of the heavens—the lawgiver of space. But, had he been asked the cause of these beautiful laws, he could only have answered, “The first cause—the direct will of the Deity.” A good answer and a true, but not scientific; because it places the question beyond the domain of science, which deals only with second or physical causes. But Newton comes forward and gives a physical cause. He shows that all these beautiful laws are the necessary result of gravitation; and thus astronomy becomes a physical science. So, until Agassiz, the facts of geological succession of organic forms were in a state of lawless confusion. Agassiz by establishing the three great laws of succession, which ought to bear his name, reduced this chaos to order and beauty; and, therefore, he might justly be called the legislator of geological history—the lawgiver of time. But, when asked the cause of these laws, he could only answer, and did indeed answer, “The plans of the Creator.” A noble answer and true, but not scientific. Darwin now comes forward and gives, partly at least, the cause of these laws. He shows that all these beautiful laws are explained by the doctrine of “origin of species by derivation with modifications”; that these laws are not ultimate, but derivative from more fundamental laws of life; and thus biology is advanced one step, at least, toward the causal stage. Newton and Darwin substituted second causes for first cause—natural for supernatural. They each in his own department broke the bonds of supernaturalism in the domain of Nature.

One more important reflection: There are two, and only two, fundamental conditions of material existence—space and time. There are, therefore, two, and only two, cosmoses—space-cosmos and time-cosmos. These have been redeemed from confusion and reduced to law and order and beauty—changed from chaos to cosmos—by science. For this result we are chiefly indebted, in the one case, to Kepler and Newton; in the other, to Agassiz and Darwin. The universal law, in the one cosmos, is the law of gravitation; in the other, the law of evolution. Traced by analysis to its deepest roots of philosophic truth, the one law may be called the divine mode of sustentation; the other, the divine process of creation.

Or again: we have all heard of the “music of the spheres”—a beautiful and significant name used by the old thinkers for the divine order of the universe—a music heard not by human ear, but only by the attentive human spirit. Harmonic relation apprehended by reason we call Law, and its embodiment Science; the same apprehended by the imagination and æsthetic sense, we call Beauty, and its embodiment Art, music. Now, in music there are two kinds of harmony, simultaneous and consecutive—chordal harmony and melody. These must be combined to produce the grandest effect. So in cosmic order, too, there are two kinds of harmonic relation—the co-existent in space and the consecutive in time. The law of gravitation expresses the universal harmonic inter-relation of objects co-existent in space, the law of evolution, the universal harmonic relation of forms successive in time. Of the divine spheral music, the one is the chordal harmony, the other the consecutive harmony or melody. Combined they form the divine chorus which “the morning stars sang together.”


PART II.
EVIDENCES OF THE TRUTH OF EVOLUTION.


CHAPTER I.
GENERAL EVIDENCES OF EVOLUTION AS A UNIVERSAL LAW.

Let us again remind the reader that evolution means, first of all, continuity. The law of evolution, although it doubtless means much more, means, first of all, a law of continuity, or causal relation throughout Nature. It means that, alike in every department of Nature, each state or condition grew naturally out of the immediately preceding. In a word, it means that, in the course of Nature, nothing appears suddenly and without natural cause, but, on the contrary, everything is the natural and usually the gradual outcome of a previous condition. This is now admitted by every one in regard to nearly everything: evolutionists apply it to the whole course of Nature. I said this is now admitted by every one in regard to nearly everything; but this has not always been so. The world has come to its present position on this subject only by a very gradual process. Let us then trace rapidly the history of the gradual change, for it will prepare us for much that follows.

There was a time (and that not many decades ago) when all things, the origin of which transcends our ordinary experience, were supposed to have originated suddenly and without natural process—to have been made at once, out of hand. There was a time when, for example, mountains were supposed to have been made at once, with all their diversified forms, of beetling cliffs and thundering waterfalls, or gentle slopes and smiling valleys, just as we now find them. But now we know that they have become so only by a very gradual process, and are still changing under our very eyes. In a word, they have been formed by a process of evolution. We know now the date of mountain-births; we trace their growth, maturity, decay, and death; and find even, as it were, the fossil bones of extinct mountains in the crumpled strata of their former places. There was a time when continents and seas, gulfs, bays, and rivers, were supposed to have originated at once, substantially as we now see them. Now, we know that they have been changing throughout all geological time, and are still changing. Not, however, change back and forth in any direction indifferently and without goal, but gradual change from less perfect to more perfect condition, with more and more complex inter-relations—i. e., by a process of evolution. We are able now, though still imperfectly, to trace some of the stages of this evolution. There was a time when rocks and soils were supposed to have been always rocks and soils; when soils were regarded as an original clothing made on purpose to hide the rocky nakedness of the new-born earth. God clothed the earth so, and there an end. Now we know that rocks rot down to soils; soils are carried down and deposited as sediments; and sediments re-consolidate as rocks—the same materials being worked over and over again, passing through all these stages many times in the history of the earth. In a word, there was a time when it was thought that the earth with substantially its present form, configuration, and climate, was made at once out of hand, as a fit habitation for man and animals. Now we know that it has been changing, preparing, becoming what it is by a slow process, through a lapse of time so vast that the mind sinks exhausted in the attempt to grasp it. It has become what it now is by a process of evolution. The same change of view has taken place concerning the origin of all the heavenly bodies. We may, therefore, confidently generalize—we may assert without fear of contradiction that all inorganic forms, without exception, have originated by a process of evolution.

The proof of all this we owe to geology—a science born of the present century. This science establishes the law of universal continuity of events, through infinite time, as astronomy does that of universal inter-relation of objects through infinite space. How great the change these two sciences have made in the realm of human thought! Until the birth of modern astronomy the intellectual space-horizon of the human mind was bounded substantially by the dimensions of our earth; sun, moon, and stars, being but inconsiderable bodies circulating at a little distance about the earth, and for our behoof. Astronomy was then but the geometry of the curious lines traced by these wandering fires on the concave blackboard of heaven. With the first glance through a telescope the phases of Venus and the satellites of Jupiter, revealed clearly to the mind the existence of other worlds besides and like our own. In that moment the idea of infinite space, full of worlds like our own, was for the first time completely realized, and became thenceforward the heritage of man. In that moment the intellectual horizon of man was infinitely extended. So also until the birth of geology, about the beginning of the present century, the intellectual time-horizon of the human mind was bounded by six thousand years. The discovery about that time of vertebrate remains, all wholly different from those now inhabiting the earth, revealed the existence of other time-faunas, besides our own and the idea of infinite time, of which the life of humanity is but an epoch, was born in the mind of man; and again the intellectual horizon of man was infinitely extended. These two are the grandest ideas, and their introduction the grandest epochs, in the intellectual history of man. We have long ago accepted and readjusted our mental furniture to the requirements of the one, but the necessary readjustment to the other is not yet complete.

All inorganic forms, then, it is admitted, have come by evolution. But how is it with organic or living forms? Let us see.

Every one knows, because it is within the limits of ordinary experience, that every individual organism now originates and gradually becomes what we see it, by a natural process—that is, by evolution. If, then, there be any exception, it must be only the first of each kind. But what kind? There are many kinds of kinds; classes, orders, families, genera, species, varieties. Now, many of these kinds can be shown to have become what we see them by a gradual process similar, at least, to evolution. Take for example, classes. The class of fishes and the class of reptiles are now widely distinct and have little in common except a vertebrate structure; but, as already shown, page 12, this extreme difference has not always existed. On the contrary, the earliest representatives of these two classes so merged into one another that each seemed either. From this common stock the two classes were gradually separated, each going its own way and becoming more and more widely distinct even to the present day. There can be no doubt, therefore, that these two classes, as we now know them, have become what they are by a gradual process. Again: In the whole realm of Nature there is not a class more distinctly separate from every other and without intermediate links than birds. But this has not always been so. They have gradually become so. The earliest birds were so reptilian in structure and appearance that if we could see them now we would be in doubt whether we should call them birds or reptiles. Birds have gradually separated themselves from the reptilian stem, becoming more and more bird-like from age to age, until now, at last, the two classes are wholly separated and the intermediate links destroyed. So far as external characters are concerned, birds may be said to have finally and wholly released themselves from entangling alliance with any other class.

Classes, then, it will be admitted, have undoubtedly become what we now know them by a very gradual process following laws identical (as we have already seen, page 19) with the laws of evolution. Shall we try orders? Of the class Mammalia there are two well-recognized and widely-distinct orders, viz., the Carnivores and the Herbivores. We all know how widely diverse these are in form, in structure, in habits, and in food. Has it always been so? Have these been made so at once? By no means. They have gradually become so. The earliest mammals were neither the one nor the other distinctively. They were omnivores, completely intermediate in food, habits, form, and structure. From this common stock the two orders have gradually separated, the carnivores becoming more and more adapted to one mode of life and the herbivores to another, by a process following the laws of evolution, as already explained. Shall we try families and genera? Marsh and Huxley have shown us how completely the horse family (Equidæ) and the horse-genus (Equus) illustrate the process of gradual becoming and the law of evolution. Under their guidance, we see that the earliest traceable ancestor of the horse family, before it was distinctively a horse family at all, had on the fore-foot five toes in the Lower Eocene, four toes in the Upper Eocene, and three toes in the Miocene; then we see the two side-toes shortening up more and more in the Pliocene and becoming rudimentary splints, leaving only one toe in the Quaternary and present epochs. Thus, the side-splints in the foot of the modern horse tell the story of its three-toed ancestry. Similar gradual changes are clearly traceable in size, shape, structure of limbs, of teeth, and of brain. In all respects the members of the horse family have become more and more horse-like in the course of time.

This subject will be taken up and more fully illustrated, under the head of special evidences, in a subsequent chapter. We here touch it only sufficiently to illustrate this universal law of gradual becoming.

We have taken only a few examples, but the same is undoubtedly true of all Taxonomic groups above species. Passing over these last for the moment, we take next races and varieties. These smaller groups are admitted by all to be formed by a natural process, because not only can we make them artificially, but all the intermediate links may be found in Nature. So we have only species remaining. Yes; species are imagined by the old-school naturalist and by the anti-evolutionist of to-day as the ultimate elements of Taxonomy. This, then, is the last ditch upon which the defense of supernaturalism in the realm of Nature is made. “Other groups,” they say, “may have gradually become what they now are by the successive introduction of specific forms according to a preordained plan which is well expressed by the formal laws of evolution. But species are without transition forms. They come in suddenly, remain unchanged while they continue, and finally pass out suddenly, so far as specific characters are concerned. New species come in their places by direct act of creation—by substitution, not by transmutation.” This, then, is the last intrenchment. Can we give any good evidence of gradual formation of species? I believe we can.

First, then, it is admitted that we can easily make varieties and races artificially. We will not now describe the process; we are all familiar with the results, viz., the varieties of domestic animals and of useful and ornamental plants; the extremely different breeds of horses, cattle, sheep, dogs, pigeons, etc.; of wheat, cabbages, turnips; of roses, dahlias, etc., etc. No one will doubt that the extreme varieties of any of these, say greyhound and pug, if wild, would be called distinct species, or even distinct genera. We do not call them so, for two reasons: first, because we see them made; and, second, because we find all intermediate links between them; and the usual definition of species is that they can not be made, and they have no intermediate links. Thus, then, the question is narrowed down to wild species. They say: “We take our stand on these” (surely a very narrow ground for so broad a philosophy). “We defy you to show gradual formation with intermediate links.”

Now, in fact, by diligent search such intermediate links between well-recognized species have been found in some cases, especially in birds, on account of their great power of dispersal. Certain forms have long been known from widely-separated regions, and universally regarded as distinct species, as distinct as any. Then, by minute examinations of intermediate regions, a complete series of intermediate forms has been picked up. This has occurred not only in one case but in many cases, and not in birds only but in many other classes—examples increase with our increasing knowledge.11 The only answer to such evidence is that these are not true species. Now, see the fallacy lurking here! They define species as ultimate elements of taxonomy, as distinct and without intermediate links, and then require us to find such intermediate links; and, finally, when with infinite pains some such links are found, they say: “Oh! I see; we were mistaken; they are only varieties!!” It is true that naturalists, when intermediate links are found, usually put all together as one species, but this they do purely for the sake of clearness of definition and description. It is freely admitted by the evolutionist that species are now usually distinct and without intermediate links, these having been destroyed in the struggle for life. This will be fully explained in another chapter. It is also freely admitted that although intermediate links must have existed at one time, their remains are rarely found. The reason of this will also be explained hereafter. Nevertheless, in some cases, as already seen, we do find them still existing. Now, we add that in some cases, where they no longer exist, we find them in the form of fossil remains. The most remarkable example of this is found in the gradual changes in the forms of Planorbis in the fresh-water deposits of Steinheim, as shown by the admirable researches of Hyatt.12 We shall discuss these also more fully in another place. Now, if there be any such links at all, however rare, then every objection to the derivative origin of species is removed.

Perhaps it may be well to make bare mention of another kind of evidence, viz., the actual change of species under the eyes, by the action of change of environment. The different species of the genus Artemia (a low form of crustacean) live in brine-pools. By concentrating the brine of such a pool, one species (A. salina) has been observed to change in successive generations into another (A. Muhlhausenii), and the latter back again to the former by slow freshening.13 Again: The siredon and the amblystoma have always, until recently, been regarded as not only distinct species, but distinct genera of amphibians. Siredon was supposed to be a permanent gill-breather, while amblystoma becomes by metamorphosis a pure air-breather. Now, however, it is known that the former may change into the latter. But the most curious part of the life-history of these animals, is that if water be abundant the siredon reproduces freely, and remains indefinitely a gill-breather; but if the water dries up it changes into the lung-breathing amblystoma. We do not give this as examples of change of species, for the change is in the individual life, and therefore in the nature of metamorphosis, but as evidence of the power of physical conditions in modifying the development of organic forms and therefore of the manner in which gill-breathers were probably transformed into air-breathers.

To sum up: 1. All inorganic forms, without exception, have become what we find them by a natural process—i. e., by evolution. 2. All organic or living forms within the limits of observation, i. e., every living thing, has become what we now see, by a gradual, natural process—i. e., by evolution. 3. All taxonomic groups, except species, have undoubtedly become what we now see them by a gradual process, following the laws of evolution, and therefore presumably by a natural process of evolution. 4. By artificial means, breeds, races, etc., very similar, at least in many respects, to species, are seen to arise by a gradual natural process—i. e., by evolution. 5. In some instances, at least, natural species are observed to pass into one another by intermediate links in such wise that we are forced to conclude that they have been formed by a natural process.

May we not, then, safely generalize, and make the law universal? Is not this a sufficient ground for confident induction? Even though some facts are still inexplicable, is that a sufficient reason for withholding assent to a theory which explains so much? In all induction we first establish a law provisionally from the observation of a comparatively few facts, and then extend it over a multitude of facts not included in the original induction. If it explains these also, the law is verified. The law of gravitation was first based on the observation of a few facts, and then verified by its explanation of nearly all the facts of celestial motion. There are some outstanding facts of celestial motion still unexplained, but we do not, therefore, doubt the law of gravitation. The same principle applied in biology ought to establish the law of evolution, for it also explains all the facts of biology as no other law can. But inductive evidence differs from other kinds of evidence in one respect, which, in fact, constitutes its strength to the scientific, but its weakness to the popular mind. It is a kind of circumstantial evidence, but its force does not consist in a few strong circumstances easily appreciated, such as strike the popular mind, and force conviction, but rather in a multitude of small circumstances, each by itself insignificant, but all together pointing to one conclusion and demanding one explanation. Such evidence is, indeed, overwhelming, but only to the mind that masters it. The evidence for the law of gravitation is literally the whole science of astronomy. So also the evidence for the law of evolution is the whole science of biology. Neither of these laws can be proved in a debating society, but only by a course of study. In the one case the law has been universally accepted—not, however, on evidence, for there are few indeed who appreciate the evidence, but on the authority of scientific unanimity. In the other case there has not yet been time enough for the already established unanimity to have its full effect.