CHAPTER XIX
THE BASS AND THEIR RELATIVES

The Cardinal-fishes. Apogonidæ.—The Apogonidæ or cardinal-fishes are perch-like fishes, mostly of small size, with two distinct short dorsal fins. They are found in the warm seas, and many of them enter rivers, some even inhabiting hot springs. Many of the shore species are bright red in color, usually with black stripes, bands, or spots. Still others, however, are olive or silvery, and a few in deeper water are violet-black.

Fig. 252.—Cardinal-fish, Apogon retrosella Gill. Mazatlan.

The species of Apogon are especially numerous, and in regions where they are abundant, as in Japan, they are much valued as food. Apogon imberbis, the "king of the mullet," is a common red species of southern Europe. Apogon maculatus is found in the West Indies. Apogon retrosella is the pretty "cardenal" of the west coast of Mexico. Apogon lineatus, semilineatus and other species abound in Japan, and many species occur about the islands of Polynesia. Epigonus telescopium is a deep-sea fish of the Mediterranean and Telescopias and Synagrops are genera of the depths of the Pacific. Paramia with strong canines is allied to Apogon, and similar in color and habit.

Allied to Apogon are several small groups often taken as distinct families. The species of Ambassis (Ambassidæ) are little fishes of the rivers and bays of India and Polynesia, resembling small silvery perch or bass. All these have three anal spines instead of two as in Apogon. Some of these enter rivers and several are recorded from hot springs. Scombrops boops, the mutsu of Japan, is a valued food-fish found in rather deep water. It is remarkable for its very strong teeth, although its flesh is feeble and easily torn. A still larger species in Cuba, Scombrops oculata, known as Escolar chino, resembles a barracuda. These fishes with fragile bodies and very strong teeth are placed by Gill in a separate family (Scombropidæ). Acropoma japonicum is a neat little fish of the Japanese coast, with the vent placed farther forward than in Apogon. It is the type of the Acropomidæ, a small family of the Pacific. Enoplosus armatus is an Australian fish with high back and fins, with a rather stately appearance, type of the Enoplosidæ. In his last catalogue of families of fishes Dr. Gill recognizes Scombropidæ and Acropomidæ as distinct families, but their relationships with Apogon are certainly very close. Many genera allied to Apogon and Ambassis occur in Australian rivers. Several fossils referred to Apogon (Apogon spinosus, etc.) occur in the Eocene of Italy and Germany.

The Anomalopidæ.—The family of Anomalopidæ is a small group of deep-sea fishes of uncertain relationship, but perhaps remotely related to Apogon. Anomalops palpebrata is found in Polynesia and has beneath the eye a large luminous organ unlike anything seen elsewhere among fishes.

The Asineopidæ.—Another family of doubtful relationship is that of Asineopidæ, elsewhere noticed. It is composed of extinct fresh-water fishes found in the Green River shales. In Asineops squamifrons the opercles are unarmed, the teeth villiform, and the dorsal fin undivided, composed of eight or nine spines and twelve to fourteen soft rays. The anal spines, as in Apogon, are two only, and the scales are cycloid.

Fig. 253.—Kuromutsu, Telescopias gilberti Jordan & Snyder. Misaki, Japan.

Fig. 254.Apogon semilineatus Schlegel. Misaki, Japan.

Fig. 255.—Robalo, Oxylabrax undecimalis (Bloch). Florida.

The Robalos:[13] Oxylabracidæ.—The family of Robalos (Oxylabracidæ or Centropomidæ) is closely related to the Serranidæ, differing among other things in having the conspicuous lateral line extended on the caudal fin. These are silvery fishes with elongate bodies, large scales, a pike-like appearance, the first dorsal composed of strong spines and the second spine of the anal especially large. They are found in tropical America only, where they are highly valued as food, the flesh being like that of the striped bass, white, flaky, and of fine flavor. The common robalo, or snook, Oxylabrax (or Centropomus) undecimalis, reaches a weight of fifteen to twenty pounds. It ranges north as far as Texas. In this species the lateral line is black. The smaller species, of which several are described, are known as Robalito or Constantino.

13.  The European zander is the type of Lacépède's genus Centropomus. The name Centropomus has been wrongly transferred to the robalo by most authors.

The Sea-bass: Serranidæ.—The central family of the percoid fishes is that of the Serranidæ, or sea-bass. Of these about 400 species are recorded, carnivorous fishes found in all warm seas, a few ascending the fresh waters. In general, the species are characterized by the presence of twenty-four vertebræ and three anal spines, never more than three. The fresh-water species are all more or less archaic and show traits suggesting the Oxylabracidæ, Percidæ, or Centrarchidæ, all of which are doubtless derived from ancestors of Serranidæ. Among the connecting forms are the perch-like genera Percichthys and Percilia of the rivers of Chile. These species look much like perch, but have three anal spines, the number of vertebræ being thirty-five. Percichthys trucha is the common trucha, or trout, of Chilean waters.

Lateolabrax japonicus, the susuki, or bass, of Japan, is one of the most valued food-fishes of the Orient, similar in quality to the robalo, which it much resembles. This genus and the East Indian Centrogenys waigiensis approach Oxylabrax in appearance and structure. Niphon spinosus, the ara of Japan, is a very large sea-bass, also of this type. Close to these bass, marine and fresh water, are the Chinese genus Siniperca and the Korean genus Coreoperca, several species of which abound in Oriental rivers. In southern Japan is the rare Bryttosus kawamebari, a bass in structure, but very closely resembling the American sunfish, even to the presence of the bright-edged black ear-spot. There is reason to believe that from some such form the Centrarchidæ were derived.

Other bass-like fishes occur in Egypt (Lates), Australia (Percalates, etc.), and southern Africa. Oligorus macquariensis is the great cod of the Australian rivers and Ctenolates ambiguus is the yellow belly, while Percalates colonorum is everywhere the "perch" in Australian rivers. The most important member of these transitional types between perch and sea-bass is the striped bass, or rockfish (Roccus lineatus), of the Atlantic coast of the United States. This large fish, reaching in extreme cases a weight of 112 pounds, lives in shallow waters in the sea and ascends the rivers in spring to spawn. It is olivaceous in color, the sides golden silvery, with narrow black stripes. About 1880 it was introduced by the United States Fish Commission into the Sacramento, where it is now very abundant and a fish of large commercial importance. To the angler the striped bass is always "a gallant fish and a bold biter," and Genio Scott places it first among the game-fishes of America.

The white bass (Roccus chrysops) is very similar to it, but shorter and more compressed, reaching a smaller size. This fish is abundant in the Great Lakes and the upper Mississippi as far south as Arkansas.

The yellow bass (Morone interrupta), a coarser and more brassy fish, replaces it farther south. It is seldom seen above Cincinnati and St. Louis. The white perch (Morone americana) is a little fish of the Atlantic seaboard, entering the sea, but running up all the rivers, remaining contentedly landlocked in ponds. It is one of the most characteristic fishes of the coast from Nova Scotia to Virginia. It is a good pan fish, takes the hook vigorously, and in a modest way deserves the good-will of the angler who cannot stray far into the mountains. Very close to these American bass is the bass, bars, or robalo, of southern Europe, Dicentrarchus labrax, a large olive-colored fish, excellent as food, living in the sea about the mouths of rivers.

The Jewfishes.—In the warm seas are certain bass of immense size, reaching a length of six feet or more, and being robust in form, a weight of 500 or 600 pounds. These are dusky green in color, thick-headed, rough-scaled, with low fins, voracious disposition, and sluggish movements. In almost all parts of the world these great bass are called jewfish, but no reason for this name has ever been suggested. In habit and value the species are much alike, and the jewfish of California, Stereolepis gigas, the prize of the Santa Catalina anglers, may be taken as the type of them all. Closely related to this is the Japanese ishinagi, Megaperca ischinagi, the jewfish, or stone-bass, of Japan. Another Japanese jewfish is the Abura bodzu, or "fat priest," Ebisus sagamius. In the West Indies, as also on the west coast of Mexico, the jewfish, or guasa, is Promicrops itaiara. The black grouper, Garrupa nigrita, is the jewfish of Florida. The European jewfish, more often called wreckfish, or stone-bass, is Polyprion americanus, and the equally large Polyprion oxygeneios is found in Australia, as is also another jewfish, Glaucosoma hebraicum, the last belonging to the Lutianidæ. Largest of all these jewfishes is Promicrops lanceolata of the South Pacific. This huge bass, according to Dr. Boulenger, sometimes reaches a length of twelve feet.

Fig. 256.—White Perch, Morone americana Gmelin. (From life by Dr. R. W. Shufeldt; one half natural size.)

Fig. 257.—Florida Jewfish, Promicrops itaiara (Lichtenstein). St. John's River, Fla.

Related to the jewfishes are numerous smaller fishes. One of these, the Spanish-flag of Cuba, Gonioplectrus hispanus, is rose-colored, with golden bands like the flag of Spain itself. Other species referred to Acanthistius and Plectropoma have, like this, hooked spines on the lower border of the preopercle.

Fig. 258.Epinephelus striatus (Bloch), Nassau Grouper: Cherna criolla. Family Serranidæ.

Fig. 259.—John Paw or Speckled Hind, Epinephelus drummond-hayi Goode Pensacola.

Fig. 260.Epinephelus morio (Cuvier & Valenciennes), Red Grouper, or Mero. Family Serranidæ.

The Groupers.—In all warm seas abound species of Epinephelus and related genera, known as sea-bass, groupers, or merous. They are mostly large voracious fishes with small scales, pale flesh of fair quality, and from their abundance they are of large commercial importance. To English-speaking people these fishes are usually known as grouper, a corruption of the Portuguese name garrupa. In the West Indies and about Panama there are very many species, and still others abound in the Mediterranean, in southern Japan, and throughout Polynesia and the West Indies. They have very much in common, but differ in size and color, some being bright red, some gaudily spotted with red or blue, but most of them are merely mottled green or brown. In many cases individuals living near shore are olivaceous, and those of the same species in the depths are bright crimson or scarlet. We name below a few of the most prominent species. Even a bare list of all of them would take many pages. Cephalopholis cruentatus, the red hind of the Florida Keys, is one of the smallest and brightest of all of them. Cephalopholis fulvus, the blue-spotted guativere of the Cubans, is called negro-fish, butter-fish, yellow-fish, or redfish, according to its color, which varies with the depth. It is red, yellow, or olive, with many round blue spots. Epinephelus adscenscionis, the rock-hind, is spotted everywhere with orange. Epinephelus guaza is the merou, or giant-bass, of Europe, a large food-fish of value, rather dull in color. Epinephelus striatus is the Nassau grouper, or Cherna criolla, common in the West Indies. Epinephelus maculosus is the cabrilla of Cuba. Epinephelus drummond-hayi, the speckled hind, umber brown, spotted with lavender, is one of the handsomest of all the groupers. Epinephelus morio, the red grouper, is the commonest of all these fishes in the American markets. In Asia the species are equally numerous, Epinephelus quernus of Hawaii and the red Epinephelus fasciatus of Japan and southward being food-fishes of importance. Epinephelus merra, Epinephelus gilberti, and Epinephelus tauvina are among the more common species of Polynesia. Epinephelus corallicola, a species profusely spotted, abounds in the crevices of coral reefs, while Cepholopholis argus and C. leopardus are showy fishes of the deeper channels. Mycteroperca venenosa, the yellow-finned grouper, is a large and handsome fish of the coast of Cuba, the flesh sometimes poisonous; when red in deep water it is known as the bonaci cardenal. Mycteroperca bonaci; the bonaci arará sells in our markets as black grouper. Mycteroperca microlepis is commonest along our South Atlantic coast, not reaching the West Indies, and Mycteroperca rubra, which is never red, enters the Mediterranean. Mycteroperca falcata is known in the markets as scamp, and Mycteroperca venadorum is a giant species from the Venados Islands, near Mazatlan. Diploprion bifasciatus is a handsome grouper-like fish with two black cross-bands, found in Japan and India. Variola louti, red, with crimson spots and a forked caudal fin, is one of the most showy fishes of the equatorial Pacific.

Fig. 261.—Red Hind, Epinephelus adscensionis (Osbeck). Puerto Rico. (After Evermann.)

Fig. 262.—Yellow-fin Grouper, Mycteroperca venenosa (Linnæus). Havana.

The small fishes called Vaca in Cuba belong to the genus Hypoplectrus. Their extraordinary and unexplained variations in color have been noticed on page 235, Vol. I. The common species—blue, orange, green, plain, striated, checkered, or striped—bears the name of Hypoplectrus unicolor. (Fig. 264).

The Serranos.—In all the species known as jewfish and grouper, as also in the Oxylabracidæ and most Centrarchidæ, the maxillary bone is divided by a lengthwise suture which sets off a distinct supplemental maxillary. This bone is wanting in the remaining species of Serranidæ, as it is also in those forms already noticed which are familiarly known as bass. The species without the supplemental maxillary are in general smaller in size, the canines are on the sides of the jaws instead of in front, and there are none of the hinged depressible teeth which are conspicuous in the groupers. The species are abundant in the Atlantic, but scarcely any are found in Polynesia, and few in Japan or India.

Fig. 263.Hypoplectrus unicolor nigricans (Poey). Tortugas, Fla.

Serranus cabrilla is the Cabrilla of the Mediterranean, a well-known and excellent food-fish, the original type of the family of Serranidæ. Serranellus scriba is the serran, a very pretty shore-fish of southern Europe, longer known than any other of the tribe. On the coast of southern California are also species called Cabrillas, fine, large, food-fish, bass-like in form, Paralabrax clathratus, and other less common species. The Cabrillas and their relatives are almost all American, a few straying across to Europe. One of the most important in the number is the black sea-bass, or black will, of our Atlantic coast, Centropristes striatus. This is a common food-and game-fish, dusky in color, gamy, and of fine flesh. The squirrel-fishes (Diplectrum) and the many serranos (Prionodes) of the tropics, small bright-colored fishes of the rocks and reefs, must be passed with a word, as also the small Paracentropristis of the Mediterranean and the fine red creole-fish of the West Indies, Paranthias furcifer. In one species, Anyperodon leucogrammicus of Polynesia, there are no teeth on the palatines.

The barber-fish (Anthias anthias) of southern Europe, bright red and with the lateral line running very high, is the type of a numerous group found at the lowest fishing level in all warm seas. All the species of this group are bright red, very handsome, and excellent as food. Hemianthias vivanus, known only from the spewings of the red snapper (Lutianus aya) at Pensacola, is one of the most brilliant species, red, with golden streaks. The genus Plesiops consists of small fishes almost black in color, with blue spots and other markings, abounding about the coral reefs. In this genus the lateral line is interrupted and there is some indication of affinity with the Opisthognathidæ.

Fig. 264.—Snowy Grouper, Epinephelus niveatus (Cuv. & Val.). Natural size: young. (Photograph by Dr. R. W. Shufeldt.)

Fig. 265.—Soapfish, Rypticus bistrispinus (Mitchill). Virginia.

In the soapfishes (Rypticus) the supplemental maxillary appears again, but in these forms the dorsal fin is reduced to two or three spines and there are none in the anal. Rypticus saponaceus, so called from the smooth or soapy scales, is the best known of the numerous species, which all belong to tropical America. Grammistes, with eight dorsal spines, is a related form in Polynesia, bright yellow, with numerous black stripes. Numerous species referred to the Serranidæ occur in the Eocene and Miocene rocks. Some are related to Epinephelus, others to Roccus and Lates. In the Tertiary lignite of Brazil is a species of Percichthys, Percichthys antiquus, with Properca beaumonti, which seem to be a primitive form of the bass, allied to Dicentrarchus. Prolates heberti of the Cretaceous, one of the earliest of the series, has the caudal rounded and is apparently allied to Lates, as is also the heavily armed Acanus regleysianus of the Oligocene. Smerdis minutus, a small fish from the Oligocene, is also related to Lates, which genus with Roccus and Dicentrarchus must represent the most primitive of existing members of this family. Of both Smerdis and Dicentrarchus (Labrax) numerous species are recorded, mostly from the Miocene of Europe.

Fig. 266.—Flasher, Lobotes surinamensis (Bloch). Virginia.

Fig. 267.—Catalufa, Priacanthus arenatus Cuv. & Val. Wood's Hole, Mass.

The Flashers: Lobotidæ.—The small family of Lobotidæ, flashers, or triple-tails, closely resembles the Serranidæ, but there are no teeth on vomer or palatines. The three species are robust fishes, of a large size, of a dark-green color, the front part of the head very short. They reach a length of about three feet and are good food-fishes. Lobotes surinamensis comes northward from the West Indies as far as Cape Cod. Lobotes pacificus is found about Panama. Lobotes erate, common in India, was taken by the writer at Misaki, Japan.

Fig. 268.—Bigeye, Pseudopriacanthus altus Gill. Young specimen. (From life by Dr. R. W. Shufeldt.)

The Bigeyes: Priacanthidæ.—The Catalufas or bigeyes (Priacanthidæ) are handsome fishes of the tropics, with short, flattened bodies, rough scales, large eyes, and bright-red coloration. The mouth is very oblique, and the anal fin about as large as the dorsal. The commonest species is Priacanthus cruentatus, widely diffused through the Pacific and also in the West Indies. This is the noted Aweoweo of the Hawaiians, which used to come into the bays in myriads at the period of death of royalty. It is still abundant, even after Hawaiian royalty has passed away.

Pseudopriacanthus altus is a short, very deep-bodied, and very rough fish, scarlet in color, occasionally taken along our coast, driven northward by the Gulf Stream. The young fishes are quite unlike the adult in appearance. Numerous other species of Priacanthus occur in the Indies and Polynesia.

The Pentacerotidæ.—Another family with strong spines and rough scales is the group of Pentacerotidæ. Histiopterus typus, the Matodai, is found in Japan, and is remarkable for its very deep body and very high spines. Equally remarkable is the Tengudai, Histiopterus acutirostris, also Japanese. Anoplus banjos is a third Japanese species, more common than the others, and largely taken in the Inland Sea. All these are eccentric variations from the perch-like type.

The Snappers: Lutianidæ.—Scarcely less numerous and varied than the sea-bass is the great family of Lutianidæ, known in America as snappers or pargos. In these fishes the maxillary slips along its edge into a sheath formed by the broad preorbital. In the Serranidæ there is no such sheath. In the Lutianidæ there is no supplemental maxillary, teeth are present on the vomer and palatines, and in the jaws there are distinct canines. These fishes of the warm seas are all carnivorous, voracious, gamy, excellent as food though seldom of fine grain, the flesh being white and not flaky. About 250 species are known, and in all warm seas they are abundant.

Fig. 269.—Gray Snapper, Lutianus griseus L. Puerto Rico. (After Evermann.)

To the great genus Lutianus most of the species belong. These are the snappers of our markets and the pargos of the Spanish-speaking fishermen. The shore species are green in color, mostly banded, spotted, or streaked. In deeper water bright-red species are found. One of these, Lutianus aya, the red snapper or pargo guachinango of the Gulf of Mexico, is, economically speaking, the most important of all these fishes in the United States. It is a large, rather coarse fish, bright red in color, and it is taken on long lines on rocky reefs chiefly about Pensacola and Tampa in Florida, although similar fisheries exist on the shores of Yucatan and Brazil.

Fig. 270.Lutianus apodus (Walbaum), Schoolmaster or Cají. Family Lutianidæ.

Fig. 271.Hoplopagrus guntheri Gill. Mazatlan.

Fig. 272.—Lane Snapper or Biajaiba, Lutianus synagris (Linnæus). Key West.

A related species is the Lutianus analis, the mutton snapper or pargo criollo of the West Indies. This is one of the staple fishes of the Havana market, always in demand for banquets and festivals, because its flesh is never unwholesome. The mangrove snapper, or gray-snapper, Lutianus griseus, called in Cuba, Caballerote, is the commonest species on our coasts. The common name arises from the fact that the young hide in the mangrove bushes of Florida and Cuba, whence they sally out in pursuit of sardines and other small fishes. It is a very wary fish, to be sought with care, hence the name "lawyer," sometimes heard in Florida. The cubero (Lutianus cyanopterus) is a very large snapper, often rejected as unwholesome, being said to cause the disease known as ciguatera. Certain snappers in Polynesia have a similar reputation. The large red mumea, Lutianus bohar, is regarded as always poisonous in Samoa—the most dangerous fish of the islands. L. leioglossus is also held under suspicion on Tutuila, though other fishes of this type are regarded as always safe. Other common snappers of Florida and Cuba are the dog snapper or jocu (Lutianus jocu), the schoolmaster or cají (Lutianus apodus), the black-fin snapper or sese de lo alto (Lutianus buccanella), the silk snapper or pargo de lo alto (Lutianus vivanus), the abundant lane snapper or biajaiba (Lutianus synagris), and the mahogany snapper or ojanco (Lutianus mahogani). Numerous other species occur on both coasts of tropical America, and a vastly larger assemblage is found in the East Indies, some of them ranging northward to Japan.

Fig. 273.—Yellow-tail Snapper, Ocyurus chrysurus (Linnæus). Key West.

Fig. 274.—Cachucho, Etelis oculatus (Linnæus). Havana.

Hoplopagrus guntheri is a large snapper of the west coast of Mexico, having very large molar teeth in its jaws besides slit-like nostrils and other notable peculiarities. From the standpoint of structure this species, with its eccentric characters—is especially interesting. The yellow-tail snapper or rabirubia (Ocyurus chrysurus) is a handsome and common fish of the West Indies, with long, deeply forked tail, which makes it a swifter fish than the others. Another red species is the diamond snapper or cagon de lo alto, Rhomboplites aurorubens. All these true snappers have the soft fins more or less scaly. In certain species that swim more freely in deep waters, these fins are naked. Among them is the Arnillo, Apsilus dentatus, a pretty brown fish of the West Indies, and its analogue in Hawaii, Apsilus brighami, red, with golden cross-bands. Aprion virescens, the Uku of Hawaii, is a large fish of a greenish color and elongate body, widely diffused throughout Polynesia and one of the best of food-fishes. A related species is the red voraz (Aprion macrophthalmus) of the West Indies.

Most beautiful of all the group are the species of Etelis, with the dorsal fin deeply divided and the head flattened above. These live in rather deep water about rocky reefs and are fiery red in color. Best known is the Cuban species, Etelis oculatus, the cachucho of the markets. Equally abundant and equally beautiful is Etelis carbunculus of Polynesia, Etelis evurus of Hawaii, and other species of the Pacific islands.

Fig. 275.Xenocys jessiæ Jordan & Bollman. Family Lutianidæ. Galapagos Islands.

Verilus sordidus, the black escolar of Cuba, has the form of Etelis, but the flesh is very soft and the color violet-black, indicating its life in very deep water. Numerous small silvery snappers living near the shore along the coast of western Mexico belong to the genera called Xenichthys, Xenistius, and Xenocys. Xenistius californiensis is the commonest of these species, Xenocys jessiæ, the largest in size, with black lines like a striped bass. To the genus Dentex belongs a large snapper-like fish of the Mediterranean, Dentex dentex. Very many related species occur in the old world, the prettily colored Nemipterus virgatus, the Itoyori of Japan being one of the best known. Another interesting fish is Aphareus furcatus, a handsome, swift fish of the open seas occasionally taken in Japan and the East Indies. Glaucosoma burgeri is a large snapper of Japan, and a related species, Glaucosoma hebraicum, is one of the "jewfishes" of Australia. Numerous fossil forms referred to Dentex occur in the Eocene of Monte Bolca, as also a fish called Ctenodentex lackeniensis from the Eocene of Belgium.

Fig. 276.Aphareus furcatus (Lacépède). Odawara, Japan. Family Lutianidæ.

Fig. 277.—Grunt, Hæmulon plumieri (Bloch). Charleston, S. C.

The Grunts: Hæmulidæ.—The large family of Hæmulidæ, known in America as grunters or roncos, is represented with the snappers in all tropical seas. The common names (Spanish, roncar, to grunt or snore) refer to the noise made either with their large pharyngeal teeth or with the complex air-bladder. These fishes differ from the Lutianidæ mainly in the feebler detention, there being no canines and no teeth on the vomer. Most of the American species belong to the genus Hæmulon or red-mouth grunts, so called from the dash of scarlet at the corner of the mouth. Hæmulon plumieri, the common grunt, or ronco arará, is the most abundant species, known by the narrow blue stripes across the head. In the yellow grunt, ronco amarillo (Hæmulon sciurus), these stripes cross the whole body. In the margate-fish, or Jallao (Hæmulon album), the largest of the grunts, there are no stripes at all. Another common grunt is the black spotted sailor's choice, Ronco prieto (Hæmulon parra), very abundant from Florida southward. Numerous other grunts and "Tom Tates" are found on both shores of Mexico, all the species of Hæmulon being confined to America. Anisotremus includes numerous deep-bodied species with smaller mouth, also all American. Anisotremus surinamensis, the pompon, abundant from Louisiana southward is the commonest species. Anisotremus virginicus, the porkfish or Catalineta, beautifully striped with black and golden, is very common in the West Indies. Plectorhynchus of Polynesia and the coasts of Asia contains numerous large species closely resembling Anisotremus, but lacking the groove at the chin characteristic of Anisotremus and Hæmulon. Some of these are striped or spotted with black in very gaudy fashion. Pomadasis, a genus equally abundant in Asia and America, contains silvery species of the sandy shores, with the body more elongate and the spines generally stronger. Pomadasis crocro is the commonest West Indian species, Pomadasis hasta the best known of the Asiatic forms. Gnathodentex aurolineatus with golden stripes is common in Polynesia.

Fig. 278.—Porkfish, Anisotremus virginicus (Linnæus). Key West.

The pigfishes, Orthopristis, have the spines feebler and the anal fin more elongate. Of the many species, American and Mediterranean, Orthopristis chrysopterus is most familiar, ranging northward to Long Island, and excellent as a pan fish. Parapristipoma trilineatum, the Isaki of Japan, is equally abundant and very similar to it. Many related species belong to the Asiatic genera, Terapon, Scolopsis, Cæsio, etc., sometimes placed in a distinct family as Teraponidæ. Terapon servus enters the streams of Polynesia, and is a very common fish of the river mouths, taken in Samoa by the boys. Terapon theraps is found throughout the East Indies. Terapon richardsoni is the Australian silver perch. Cæsio contains numerous small species, elongate and brightly colored, largely blue and golden. Scolopsis, having a spine on the preorbital, contains numerous species in the East Indies and Polynesia. These are often handsomely colored. Among them is the taiva, Scolopsis trilineatus of Samoa, gray with white streaks and markings of delicate pattern. A fossil species in the Italian Eocene related to Pomadasis is Pomadasis furcatus. Another, perhaps allied to Terapon, is called Pelates quindecimalis.

Fig. 279.—The Red Tai of Japan, Pagrus major Schlegel. Family Sparidæ. (After Kishinouye.)

The Porgies: Sparidæ.—The great family of Sparidæ or porgies is also closely related to the Hæmulidæ. The most tangible difference rests in the teeth, which are stronger, and some of those along the side of the jaw are transformed into large blunt molars, fitted for grinding small crabs and shells. The name porgy, in Spanish pargo, comes from the Latin Pagrus and Greek πάγρος, the name from time immemorial of the red porgy of the Mediterranean, Pagrus pagrus. In this species the front teeth are canine-like, the side teeth molar. It is a fine food-fish, very handsome, being crimson with blue spots, and in the Mediterranean it is much esteemed. It also breeds sparingly on our south Atlantic and Gulf coasts.

Fig. 280.—Ebisu, the Fish-god of Japan, bearing a Red Tai. (Sketch by Kako Morita.)

Fig. 281.—Scup, Stenotomus chrysops (Linnæus). Wood's Hole, Mass.

Fig. 282.Calamus bajonado (Bloch & Schneider), Jolt-head Porgy. Pez de Pluma. Family Sparidæ.

Fig. 283.—Little-head Porgy, Calamus proridens Jordan & Gilbert. Key West.

Very similar to the porgy is the famous red tai or akadai of Japan (Pagrus major), a fish so highly esteemed as to be, with the rising sun and the chrysanthemum, a sort of national emblem. In all prints and images the fish-god Ebisu (Fig. 280), beloved of the Japanese people, appears with a red tai under his arm. This species, everywhere abundant, is crimson in color, and the flesh is always tender and excellent. A similar species is the well-known and abundant "schnapper" of Australia, Pagrus unicolor. Another but smaller tai or porgy, crimson, sprinkled with blue spots, Pagrus cardinalis, occurs in Japan in great abundance, as also two species similar in character but without red, known as Kurodai or black tai. These are Sparus latus and Sparus berda. The gilt-head of the Mediterranean, Sparus aurata, is very similar to these Japanese species. Sparus sarba in Australia is the tarwhine, and Sparus australis the black bream. The numerous species of Pagellus abound in the Mediterranean. These are smaller in size than the species of Pagrus, red in color and with feebler teeth. Monotaxis grandoculis, known as the "mu," is a widely diffused and valuable food-fish of the Pacific islands, greenish in color, with pale cross-bands. Very closely related is also the American scup or fair maid (Stenotomus chrysops), one of our commonest pan fishes. In this genus and in Calamus the second interhæmal spine is very greatly enlarged, its concave end formed like a quill-pen and including the posterior end of the large air-bladder. This arrangement presumably assists in hearing. Of the penfishes, or pez de pluma, numerous species abound in tropical America, where they are valued as food. Of these the bajonado or jolt-head porgy (Calamus bajonado) is largest, most common and dullest in color. Calamus calamus is the saucer-eye porgy, and Calamus proridens, the little-head porgy. Calamus leucosteus is called white-bone porgy, and the small Calamus arctifrons the grass-porgy.

The Chopa spina, or pinfish, Lagodon rhomboides, is a little porgy with notched incisors, exceedingly common on our South Atlantic coast.

In some of the porgies the front teeth instead of being canine-like are compressed and truncate, almost exactly like human incisors. These species are known as sheepshead, or sargos.

Fig. 284.Diplodus holbrooki Bean. Pensacola.

Diplodus sargus and Diplodus annularis are common sargos of the Mediterranean, silvery, with a black blotch on the back of the tail. Diplodus argenteus of the West Indies and Diplodus holbrooki of the Carolina coast are very close to these.

The sheepshead, Archosargus probatocephalus, is much the most valuable fish of this group. The broad body is crossed by about seven black cross-bands. It is common from Cape Cod to Texas in sandy bays, reaching rarely a weight of fifteen pounds. Its flesh is most excellent, rich and tender. The sheepshead is a quiet bottom-fish, but takes the hook readily and with some spirit. Close to the sheepshead is a smaller species known as Salema (Archosargus unimaculatus), with blue and golden stripes and a black spot at the shoulder. It abounds in the West Indies.

Fig. 285.Archosargus unimaculatus (Bloch), Salema, Striped Sheepshead. Family Sparidæ.

On the coast of Japan and throughout Polynesia are numerous species of Lethrinus and related genera, formed and colored like snappers, but with molar teeth and the cheek without scales. A common species in Japan is Lethrinus richardsoni.

Fossil species of Diplodus, Sparus, Pagrus, and Pagellus occur in the Italian Eocene, as also certain extinct genera, Sparnodus and Trigonodon, of similar type. Sparnodus macrophthalmus is abundant in the Eocene of Monte Bolca.

The Picarels: Mænidæ.—The Mænidæ, or Picarels, are elongate, gracefully formed fishes, remarkable for the extreme protractility of the upper jaw. Spicara smaris and several other small species are found in the Mediterranean. Emmelichthys contains species of larger size occurring in the West Indies and various parts of the Pacific, chiefly red and very graceful in form and color. Emmelichthys vittatus, the boga, is occasionally taken in Cuba, Erythrichthys schlegeli is found in Japan and Hawaii.

Fig. 286.—Mojarra, Xystæma cinereum (Walbaum). Key West.

The Mojarras: Gerridæ.—The Gerridæ, or Mojarras, have the mouth equally protractile, but the form of the body is different, being broad, compressed, and covered with large silvery scales. In some species the dorsal spines and the third anal spine are very strong, and in some the second interhæmal is quill-shaped, including the end of the air-bladder, as in Calamus. Most of the species, including all the peculiar ones, are American. The smallest, Eucinostomus, have the quill-shaped interhæmal and the dorsal and anal spines are very weak. The commonest species is the silver jenny, or mojarra de Ley, Eucinostomus gula, which ranges from Cape Cod to Rio Janeiro, in the surf along sandy shores. Equally common is Eucinostomus californiensis of the Pacific Coast of Mexico, while Eucinostomus harengulus of the West Indies is also very abundant. Ulæma lefroyi has but two anal spines and the interhæmal very small. It is common through the West Indies. Xystæma, with the interhæmal spear-shaped and normally formed, is found in Asia and Polynesia more abundantly than in America, although one species, Xystæma cinereum, the broad shad, or Mojarra blanca, is common on both shores of tropical America. Xystæma gigas is found in Polynesia, X. oyena in Japan, and X. filamentosum in Formosa and India. Xystæma massalongoi is also fossil in the Miocene of Austria. The species of Gerres have very strong dorsal and anal spines and the back much elevated. Gerres plumieri, the striped mojarra, Gerres brasiliensis, the patao, Gerres olisthostomus, the Irish pampano, and Gerres rhombeus are some of the numerous species found on the Florida coast and in the West Indies. The family of Leiognathidæ, already noticed (page 287), should stand next to the Gerridæ.