Fig. 8. Sketch of typical crinoidal limestone from the Pennsylvanian of north Texas.

Typical invertebrate fossils are foraminifera (principally fusulinids), corals (especially the solitary or “horn” corals), brachiopods, bryozoans (the lacy and branching types are most common), pelecypods, gastropods (exhibiting a variety of coiling), cephalopods (nautiloids and goniatites predominate), and crinoids, which in many areas are found in thick crinoidal limestones (fig. 8). Some typical Pennsylvanian fossils are illustrated in Plates 14, 15, 17, 18, 19, 20, 21, 24, 32, and 35.

Permian

Permian rocks are found in widely separated areas in Texas. The best exposed section of marine Permian rocks is found in the Glass Mountains of Brewester County, and many of these rocks are very fossiliferous. The original shell material of some of the Permian fossils of this area has been replaced by siliceous material which is very well preserved. These silicified fossils are removed from the limestone by solution in acid, and some most remarkable specimens have been recovered in this manner (Pl. 3). Brachiopods are the most common fossils, but corals, bryozoans, and mollusks have also been recovered.

Extensive Permian exposures occur also in the central part of the North-Central Plains region. These rocks were formed from sediments of both marine and continental origin and some of them are fossiliferous. The marine rocks contain a variety of invertebrate fossils including brachiopods, pelecypods, gastropods, and ammonoids. Those rocks representing terrestrial deposits contain vertebrate remains at many localities, and numerous amphibians and primitive reptiles (Pl. 40) have been collected from them.

MESOZOIC ROCKS

Mesozoic rocks occur over a wide area of Texas and include exposures of Triassic, Jurassic, and Cretaceous age. Many of the Upper and Lower Cretaceous outcrops are quite fossiliferous and easily accessible and thus of considerable interest to many amateur collectors.

Triassic

Triassic rocks crop out in parts of the High Plains, the Glass Mountains of Trans-Pecos Texas, and parts of Pecos, Crockett, Upton, Reagan, and Glasscock and other west Texas counties. These are predominantly nonmarine rocks consisting of conglomerates, sandstones, shales, and some gypsum beds.

Triassic fossils are almost exclusively vertebrates, although some poorly preserved plant and invertebrate remains have been reported. Fossil vertebrates of the Texas Triassic include phytosaurs (Pl. 42), crocodiles, amphibians, and fish.

Jurassic

In Texas, surface exposures of Jurassic rocks are known only from Malone Mountain in southwestern Hudspeth County. The rocks there are limestones, shales, sandstones, and conglomerates. Fossils reported from that locality include marine and fresh-water pelecypods, fresh-water gastropods, and ammonites.

Cretaceous

Rocks of Cretaceous age are widely distributed in Texas and represent one of the more important rock systems of the State. Cretaceous outcrops occur in central Texas, north Texas, the Edwards Plateau, parts of the High Plains, the Gulf Coastal Plain, and Trans-Pecos Texas.

As mentioned earlier, the Texas Cretaceous has been divided into the Lower Cretaceous (Comanche series) and Upper Cretaceous (Gulf series). These rocks consist primarily of marls (a type of calcareous clay), shales, chalks, and limestones, but sands and conglomerates also occur. Cretaceous rocks occur on the surface of about 28 percent of Texas, and many of the larger cities of the State are situated on Cretaceous strata.

Many of the Gulf and Comanche formations contain fossils which are of interest both to amateur and professional paleontologists. Because of their wide distribution in and near large population centers, Cretaceous outcrops can be conveniently visited by many amateur fossil collectors. The fossils are usually abundant and varied, and some are well preserved. Although numerous kinds of fossils may be collected, the more common forms are cephalopods, pelecypods, gastropods, and echinoids. Some of the more typical Cretaceous fossils are shown in Plates 16, 21, 25-28, 32, 33, 35, and 36.

Cretaceous fossils are more commonly found in shales and chalky limestones. Fossiliferous outcrops of these rocks can be found along many streams, roads, and highways of central Texas, north Texas, and the Edwards Plateau. Outcrops which have been weathered are more likely to provide good collecting. In general, collecting is poor in areas covered with heavy vegetation or recent stream deposits. Good collecting localities are outcrops which have a fairly steep slope with a covering of weathered rock material and a minimum of vegetation. One should move slowly from the base of the slope upward while searching the ground for any evidence of fossils, and particular attention should be given to any small gullies since these often contain fossils that have been washed out of upper beds in the exposure.

CENOZOIC ROCKS

Cenozoic rocks are widespread in Texas but occur primarily in a broad belt along the Gulf Coastal Plain. In addition, there are exposures of nonmarine Cenozoic strata in the High Plains, North-Central Plains, and Trans-Pecos region. There are also many exposures of Cenozoic igneous rocks in Trans-Pecos Texas.

Rocks of Cenozoic age occur in more than one-third of Texas and consist of conglomerates, sands, clays, and some limestone and lignite beds.

Tertiary

Extensive exposures of Tertiary rocks trend northeast-southwest in a broad band across the Gulf Coastal Plain area. These strata, consisting of sands, clays, and poorly consolidated limestones, are underlain by Cretaceous rocks.

Invertebrate fossils are common in certain Tertiary formations and pelecypods, gastropods, and corals are the predominant forms. In general, however, fossiliferous exposures are of local occurrence and most of the Tertiary formations are unfossiliferous. Those Tertiary invertebrates that are present, however, are often well preserved and represent many interesting types (Pls. 16, 22, 23, 29, 30, 31).

Tertiary invertebrate fossils are commonly found in sands, clays, and marls. Many of these sands and marls have a green color which is due to the presence of glauconite (a green mineral containing iron and closely related to the micas). At certain localities on the Gulf Coastal Plain the glauconite marls and sands of the Weches and Crockett formations contain large numbers of well-preserved clams, snails, and corals. Fossiliferous exposures of Tertiary rocks are sometimes found in road cuts, but better exposures may be found along the banks of rivers and creeks. Certain bluffs along the Brazos, Sabine, and Trinity rivers are well-known Tertiary fossil collecting localities. Many of these better localities are listed in some of the Bureau of Economic Geology bulletins included in the bibliography of this publication (pp. 109-110).

Quaternary

Quaternary deposits of Pleistocene age (geologic time scale, Pl. 1) are found in many parts of Texas and consist of sands, clays, and gravels.

These rocks are distributed along the Gulf Coast in a belt from 50 to 100 miles wide. They occur also as stream terraces in the Edwards Plateau and North-Central Plains regions. In addition, Quaternary sands and gravels are widely distributed over the surface of much of Trans-Pecos Texas. There are also fossiliferous Pleistocene strata in the High Plains region.

Invertebrate fossils are rare in Pleistocene rocks, but some fresh-water and terrestrial mollusks occur. Vertebrate remains, however, are abundant in many localities, and large numbers of horses, camels, mammoths, and other mammals (Pls. 46-49) have been collected. Fossil bones and teeth (figs. 25, 26, p. 104) are commonly found in the gravels and sands of many of the river terraces of the State.

MAIN TYPES OF FOSSILS

The beginning fossil collector is usually amazed by the many different plants and animals that have left some trace of their existence. In order to understand these different types of prehistoric life, it is necessary to know something about the organisms that are living today.

This handbook discusses the more important groups of plants and animals which have left some sort of paleontological record, and each major group begins with a discussion of the more simple organisms and continues through the more advanced forms. Because scientific workers do not always agree on exactly the same classification, the system adopted in this handbook contains the latest ideas of several workers. It is simple enough to understand, yet complete enough to help one know and classify his fossils. It should be noted that this classification may differ in some respects from that of certain older paleontological publications. Therefore, it has seemed advisable to list other names for some of the groups that are discussed.

In some instances, the brief descriptions and illustrations of each group will enable the collector to make a preliminary identification of his fossils. For more detailed information about each group, the reader should refer to “Books About Fossils” (pp. 108-110).

This part of the handbook begins with a brief summary of the major groups of the plant kingdom, followed by a discussion of the characteristics and relative paleontological importance of the various invertebrate animals. Emphasis is placed on the invertebrates because this type of fossil is most commonly collected by the amateur. Finally, there is a general review of the vertebrates.

PLANT FOSSILS

Plant fossils are usually fragmental and poorly preserved, and this tends to discourage most amateurs from an active interest in paleobotany. However, in spite of these problems, much is known of the evolution of plants, and plant fossils provide much information about life of the past. In addition, certain plants are of considerable value as indicators of ancient climatic conditions, and their remains have played a large part in the formation of vast coal deposits.

Classification of the Plant Kingdom

In the following classification only the larger taxonomic groups are discussed. Notice that the term division has been used in place of the term phylum as used in the animal kingdom. This usage is now preferred by many botanists and paleobotanists.

DIVISION THALLOPHYTA

Thallophytes are simple plants without roots, stems, or leaves. They include the fungi, algae, and diatoms (Pl. 12). Diatoms are microscopic fossils that are found in many of the rocks of Texas, and they are quite abundant in Recent sediments as well. Certain of the Paleozoic limestones of central Texas contain banded spherical masses of algae called “algal biscuits.” Although not particularly useful fossils, thallophytes have a long geologic history and are known in rocks ranging from Precambrian to Recent in age.

DIVISION BRYOPHYTA

The bryophytes are simple rootless plants and include the mosses and liverworts. Although more complex, the bryophytes resemble the algae in some respects. They are uncommon fossils, but undoubted bryophytes (liverworts) have been reported from rocks as old as Mississippian.

DIVISION TRACHEOPHYTA

This division has been divided into four subdivisions, among which are many of the more common living and fossil plants. Such important plants as the ferns, evergreens, hardwood trees, and the flowering plants are all tracheophytes. Among the more common and abundant fossil tracheophytes are the ferns, cycads, and Gingko, in addition to such important “coal plants” as the scale trees, club mosses, and scouring rushes (Pls. 12, 13). The latter commonly occur in many of the world’s great coal deposits, and their remains make up a large part of the coal. Plant fossils of this type may be collected in the dumps around some of the abandoned coal mines in north-central Texas and from other Pennsylvanian rocks in north and Trans-Pecos Texas.

Plate 11
GEOLOGIC RANGE OF THE MAJOR GROUPS OF PLANTS AND ANIMALS
The bands give some indication of the geologic range and relative abundance of the major groups of plants and animals. An increase in the width of the range band corresponds to a relative increase in numbers during the corresponding portion of geologic time.

PRECAMBRIAN
PALEOZOIC
CAMBRIAN
ORDOVICIAN
SILURIAN
DEVONIAN
MISSISSIPPIAN
PENNSYLVANIAN
PERMIAN
MESOZOIC
JURASSIC
CRETACEOUS
CENOZOIC
Thallophyta
Bryophyta
Tracheophyta
Protozoa
Porifera
Coelenterata
Bryozoa
Brachiopada
Mollusca
Annelida
Arthropoda
Echinodermata
Chordata

Plate 12
FOSSIL PLANTS

THALLOPHYTES

DIATOMS × 900
ALGAE × 400

TRACHEOPHYTES

LEPIDODENDRON × ½
SIGILLARIA × ½

Plate 13
FOSSIL PLANTS
TRACHEOPHYTES

NEUROPTERIS × ½
PSILOPHYTON × ⅓
CALAMITES × ½
AMELANCHIER × ½
CORDAITES × ¼
GINGKO × ½

Fairly well-preserved plant remains may also be collected from the Woodbine group of the Upper Cretaceous in north Texas, and fossil wood, most of it silicified, has been reported from rocks of almost all ages and in almost every section of the State. In addition, some of the carbonaceous clays and shales of east Texas contain large assemblages of plant leaves, which in some places are well preserved.

It is also possible to find the fossilized remains of seeds, spores, and pollen. Because of their small size, these minute remains are not destroyed by the drill bit and can be brought out of deep wells without being damaged, and for this reason they are a valuable tool for the micropaleontologist.

ANIMAL FOSSILS

The fossilized remains of animals are very common in many of the sedimentary rocks of Texas. These remains are of many different kinds and represent the fossils of such diverse organisms as the shell of a tiny one-celled animal or the bones or tusk of a huge elephant. The fossils most commonly found, however, are the remains of invertebrate animals such as clams, snails, and corals, and it is this type of fossil that attracts the interest of most amateur collectors.

It is not always easy to tell whether certain organisms are plants or animals, and because of this some scientists have suggested that these “in-betweens” be placed in a separate kingdom—the Protista. The protistans are primarily unicellular organisms and are represented by such forms as bacteria, algae, diatoms, and the protozoans (see below). But in this publication, only the plant and animal kingdoms are recognized.

Phylum Protozoa

This phylum is composed of simple one-celled animals many of which have no shell or external body covering. Some, however, have external hard parts that can become fossilized, and these forms are quite useful microfossils.

CLASS SARCODINA.—

This class contains a group of one-celled animals which may secrete an exoskeleton (external protective covering) of chitin, silica, or calcium carbonate. Included in this class are foraminiferans (commonly called forams) and radiolarians.

Order Foraminifera.—

Members of this order secrete tiny chambered shells which are very useful microfossils. The forams are predominantly marine organisms and have shells composed of chitin, silica, or calcium carbonate. In addition, some forms construct a shell of sand grains or some other material which is cemented together by a sticky substance that is secreted by the animal.

Forams are very abundant in the rocks of Texas and particularly so in rocks of Mesozoic and Cenozoic age. The most numerous and easily observed Paleozoic foraminiferans are the fusulinids (fig. 9a), and their small spindle-shaped remains are very abundant in many of the Pennsylvanian limestones of north-central and Trans-Pecos Texas. Some typical Texas forams are illustrated in figure 9.

Order Radiolaria.—

The radiolarians (fig. 10) have delicate spine-covered shells composed of silica, and their remains are very abundant in certain recent marine sediments. They may also be found as fossils and have been reported from Devonian and Permian rocks in Trans-Pecos Texas, and probable radiolarians have been reported from still younger beds.

Fig. 9. Typical Texas Foraminifera (all greatly enlarged). (a) Fusulina (Pennsylvanian). (b) Robulus. (c) Globigerina. (d) Frondicularia. (b-d, Cretaceous).

Fig. 10. Typical radiolarians (greatly enlarged). (a) Actinomma (Recent). (b) Porodiscus (Eocene).

Phylum Porifera

These are sponges and are the simplest of the many-celled animals. Living sponges secrete a skeleton which may be composed of chitin, silica, or calcium carbonate. These substances are commonly found in the form of spicules—tiny hard parts that are used to help support the soft tissues of the animal. These spicules take on a variety of shapes (Pl. 14) and are occasionally found as microfossils in some marine sediments.

Although sponges are not particularly common fossils, their remains occur in some parts of the State. Sponges have been collected from Paleozoic and Mesozoic formations of north and Trans-Pecos Texas, and their spicules have been reported from well cuttings.

Phylum Coelenterata

The coelenterates are multicelled animals which, though more complex than the sponges, are rather primitive animals. The living animal is characterized by a sac-like body cavity, a definite mouth, and tentacles which bear stinging cells. Some forms, for example, the jellyfishes, have an umbrella-shaped body and are single free-moving organisms. Others, like the colonial corals, are composed of many individuals living together in a colony.

Most zoologists and paleontologists recognize three classes of coelenterates: (1) the Hydrozoa, containing the small animals known as hydroids, (2) the Scyphozoa, which includes the jellyfish, and (3) the Anthozoa, which includes the corals and sea anemones. Because of their extreme fragility and lack of hard parts, hydrozoans and scyphozoans are not commonly found as fossils. They do, however, have a long geologic history and may be preserved when unusual conditions of fossilization occur. The anthozoans, especially the corals, are by far the most important class geologically, and these forms have left a very good paleontological record.

CLASS ANTHOZOA.—

This class is composed of a group of exclusively marine organisms and includes the corals and sea anemones. The coral animal, or polyp, secretes a cup-shaped calcareous (limy) exoskeleton. This skeleton, called a corallite, is usually divided by radial partitions called septa. The polyp lives in the calyx, which is the central bowl-shaped depression in the top of the corallite (fig. 11a).

Solitary corals form an individual corallite for each polyp, and because of their shape these may be given such names as “horn corals” (Lophophyllidium, Pl. 15) or “button corals” (Micrabacia, Pl. 16). Colonial or compound corals (Pl. 15) live together in colonies, which are formed of many individual skeletons attached to each other (fig. 11b), and the compound mass of coral skeletons formed in this manner is called a corallum. Fossil corals commonly occur in many marine limestones and in places constitute a large portion of the rock.

Fig. 11. Morphology and principal parts of corals. (a) Solitary or “horn” coral. (b) Colonial or compound coral.

a
Columella
Septum
Corallite
b
Calyx
Septum
Corallum

The class Anthozoa has been divided into several subclasses, but only one, the Zoantharia, is of paleontological importance.

Subclass Zoantharia.—

Most corals and all sea anemones belong to this subclass. Zoantharians are either colonial or solitary and, because most of them possess a hard preservable exoskeleton, they are the most important group of anthozoans geologically. The various orders of the subclass Zoantharia are discussed below.

Order Rugosa.—

These are corals in which the septa are arranged in cycles of four. Both solitary and colonial forms occur, and they are found only in rocks of Paleozoic age. Rugose corals are abundant in many of the Paleozoic formations of Texas, and two of the more typical forms (Lophophyllidium and Caninia) are illustrated in Plate 15. Members of this order have been placed in the subclass Tetracoralla of older classifications.

Order Scleractinia.—

The scleractinians are solitary or colonial corals in which the septa grow in multiples of six, and they are the most important and abundant of the modern corals. These corals were the dominant reef builders of Mesozoic and Cenozoic seas, and their remains are common in many of the marine formations of the State. Plate 16 illustrates some typical scleractinian corals from the rocks of Texas. This order has also been referred to as subclass Hexacoralla, and its members have been called hexacorals.

Order Tabulata.—

These are corals that are now extinct but are known from fossils in both Paleozoic and Mesozoic rocks. Tabulate corals are characterized by horizontal partitions called tabulae, and septa are absent or poorly developed. The tabulates were the most abundant reef-building corals during Paleozoic time and are well known as fossils. Because of certain similarities with other anthozoans, some paleontologists have treated the Tabulata as a distinct subclass rather than as an order of the Zoantharia.

Tabulate corals are not uncommon in many of the Paleozoic rocks of Texas, and two of these (Cladochonus and Striatopora) are illustrated in Plate 15.

Phylum Bryozoa