Classification of Ophidia.–Duméril and Bibron^[176] divided Snakes according to their teeth into Opotérodonts, Aglyphodonts, Solenoglypha, Proteroglypha, and Opisthoglypha.

J. E. Gray^[177] divided Snakes into two sub-orders: Viperina and Colubrinia. Günther^[178] distinguished between Ophidii colubriformes, O. colubriformes venenosi (Elapidae and Hydrophidae) and O. viperiformes. Cope^[179] laid stress upon the modifications of the squamosal, ectopterygoid, and ectopterygoid bones, and also upon the condition of the vestigial limbs. He divided the snakes into Scolecophidia (Typhlopidae), Catodonta, Tortricina, Asinea (the harmless snakes without limb-vestiges), Proteroglypha, and Solenoglypha.

Boulenger^[180] has accepted Cope's principles, and, mainly by combining the Asinea with the Proteroglypha as Colubridae, has produced a logically conceived system, by far the best hitherto proposed. It has been followed in the present work. Boulenger's phylogenetic system stands as follows:–

For ordinary practical purposes this synopsis is useless, being based entirely upon anatomical characters, not all easily ascertained. The following characterisation of families may therefore be preferred:–

Fam. 1. Typhlopidae.–Burrowing snakes which have the whole body covered with uniform cycloid scales, and with the teeth restricted to the small and transversely placed maxillary bones. The pterygoids do not extend backwards to the quadrates, and there are no endopterygoids. The quadrates slant obliquely forwards, and are attached directly to the pro-otics, owing to the absence of squamosal bones. The prefrontals are in lateral contact with the nasals. There are vestiges of the pelvis, reduced to a single bone on each side. The eyes are hidden by shields of the skin.

The Typhlopidae, mainly composed of the genus Typhlops, with about one hundred species, are undoubtedly the last living descendants of formerly cosmopolitan, rather archaic, snakes, which in adaptation to their burrowing life and insectivorous diet have undergone degradation. They are still widely distributed in all tropical and sub-tropical countries, some on the solitary Christmas Island, but not in New Zealand. One species, T. vermicularis, inhabits the Balkan Peninsula and South-West Asia. It is brown above, yellowish below, and reaches a length of about 10 inches. The tail is extremely short and ends in a horny spine. T. braminus is widely distributed in Southern Asia, the Malay Islands, the islands in the Indian Ocean and in Southern Africa.

Fam. 2. Glauconiidae.–In most respects resembling the Typhlopidae, but the maxillaries retain their normal position and are toothless, teeth being restricted to the lower jaw, which is stout and short. The pelvic girdle and the hind-limbs show the least reduction found in any recent Snakes; in the pelvis the ilia, pubes, and ischia can still be distinguished, the last even retaining their symphysis; there are also vestiges of femurs. About thirty species, nearly all belonging to the genus Glauconia, are found in South-Western Asia, Africa, and the warmer parts of America, including the West Indies.

Fam. 3. Ilysiidae.–The scales of the cylindrical body are smooth and small, those on the ventral side are scarcely larger. The tail is extremely short and blunt. The head is very small, not distinct from the neck. The gape of the mouth is very narrow. Teeth are carried by the mandibles, the pterygoids, palatines, maxillaries, and one or two or more by the premaxillae. The endopterygoids are short. An important cranial feature is the short quadrates, which stand rather vertically and are connected with the cranium by the squamosals; these are very small and are firmly wedged in between the upper ends of the quadrates and the pro-otic, lateral, and supra-occipital bones; now forming part of the cranial wall. Vestiges of the pelvis and hind-limbs are very incomplete, and terminate in claw-like spurs, protruding between the scales on either side of the vent. The eyes are very small, and are either free or covered by transparent shields. The few, scarcely half-a-dozen, species are found in South America (Ilysia) and in Ceylon, the Malay Islands, and Indo-China.

Ilysia (Tortrix) scytale, the Coral-Snake of Tropical South America, is a beautiful coral-red with black rings. On account of its beauty, perfectly harmless nature, and for "cooling purposes," this snake, which grows to nearly a yard in length, is sometimes worn as a necklace by native ladies. All the Ilysiidae lead a partly burrowing life, live chiefly upon worms, insects, and little Typhlopidae, and are viviparous.

Fam. 4. Uropeltidae.–Burrowing snakes of Ceylon and Southern India, with a short and rigid cylindrical body and a very short tail, which ends in a large peculiar shield, often obliquely truncated. The scales of the body are smooth, and are little larger on the belly; the coloration is mostly very beautiful. The eyes are very small.

The Uropeltidae are somewhat intermediate between the Ilysiidae, Glauconiidae, and Boidae. The pterygoids do not reach the quadrates; but ectopterygoids are present; the quadrates are very small and directly attached to the skull, squamosals being absent. Teeth are carried by the mandibles and by the maxillaries, which are normal in their position. There are no vestiges of hind-limbs or of the pelvis. The Uropeltidae, of which about forty species are known, are viviparous, burrow in the ground, and frequent damp localities, preferring mountain-forests. The use of the characteristic tail-shield is not clear; perhaps it assists these rather rigid creatures in digging, by being pressed against the ground.

Uropeltis.–The tail is obliquely truncated, ending in a roundish, flat shield.

U. grandis s. philippinus.–The latter name seems to have misled W. Marshall^[182] into including the Philippine Islands in the range of the family, a mistake which is sure to be propagated. The species, the only one of the genus, is confined to Ceylon; it is blackish above, yellow below, frequently with small yellow spots above and brown spots on the under surface. It grows to about 18 inches in length.

Rhinophis.–The tail-shield is convex and the snout is pointed. Rh. sanguineus of Southern India is black above with a bluish gloss, sometimes with small pale specks; the belly and several of the lateral series of scales are bright red, spotted with black. The tail-shield is black and red.

Fam. 5. Boidae.–Typical Snakes, usually large, and with vestiges of pelvis and hind-limbs, appearing externally as claw-like spurs on each side of the vent. The scales of the upper surface are usually small and smooth, while those of the ventral surface form one broad series on the belly, and one or two rows on the tail. The quadrate is carried by the horizontally elongated squamosal, which rests loosely upon the lateral occipital region. The prefrontal is in contact with the nasal. Teeth are carried by the mandibles, the pterygoids, palatines, maxillaries, and, in the Pythoninae, by the premaxillaries also. For further details see Figs. 155, 156.

The Boidae comprise between sixty and seventy species, which have been grouped into many genera, on unimportant characters, referring to the scales and shields of the head. It is doubtful if they are natural groups, a consideration which detracts much from their value in the study of geographical distribution. Even the two sub-families are not free from this reproach. The range of the family is world-wide, Boidae occurring in all tropical and sub-tropical countries, including islands, except New Zealand. A few species live in South-Eastern Europe (Eryx) and in North-Western America. They mostly prefer wooded districts, especially forests; climbing trees, assisted by the short and partly prehensile tail. Others are semi-aquatic, and a few live in sandy localities. They are all rapacious, and by preference feed on warm-blooded creatures, which they constrict by coils of the body in order to hold, kill, and crush the victim before swallowing it. Exaggerated notions are entertained about their swallowing capacity. It is obvious that a large snake, 20 feet long, half a foot thick, and weighing several hundred pounds, can crush a tiger, a stag, or even a cow; but common sense tells us where to draw the line when it comes to the swallowing of the prey. Small game, although of a bulk apparently far too big for the snake, is so crushed and mangled that it is turned into the shape of a sausage preparatory to the long process of swallowing. The Boidae lay eggs, and some species incubate them, or rather the female coils herself round them for the sake of protection. No appreciable amount of extra warmth is developed. Unfortunately the observations of one of the best cases on record^[183] were conducted so imperfectly that they are of little value.

Sub-Fam. 1. Pythoninae.–With a pair of supra-orbital bones, intercalated between the prefrontal, frontal, and postfrontal bones. The sub-caudal scales are mostly in two rows. The premaxilla often carries a few small teeth.

The Pythoninae, comprising about twenty species, are restricted to the Palaeotropical and Australian regions, with the sole exception of Loxocemus bicolor in Southern Mexico.

Python, the principal genus, has teeth on the premaxilla. The rostral, each of the anterior upper labials, and some of the lower labial shields, contain a deep, probably sensory, pit. The maxillary and mandibular teeth are long, but decrease from before backwards. The head is distinct from the neck, and is covered with symmetrical shields or with small scales. The scales of the body are small and smooth. The tail is short and prehensile; below with two rows of scales. The pupil of the eye is vertical. The range of the genus extends over the whole of the Palaeotropical and Australian regions, excepting Madagascar and New Zealand.

P. spilotes, the "Carpet Snake" of Australia and New Guinea, is mostly beautifully marked, but is subject to much variation in colour. The more typical specimens are black above, each scale with a yellowish dot, with yellow spots or combinations of dots, more or less arranged in rows. The under parts are yellow. It reaches a length of about two yards, and spends a great part of its time in trees.

P. reticulatus is the commonest species in Indo-China and in the Malay Islands. Four upper labial shields of each side are pitted. The specific name refers to the bold, dark, lozenge-shaped markings upon the lighter yellowish or brown ground. A black line extends over the head from the nose to the neck, and another on each side from the eye to the angle of the mouth. The under parts are mostly yellowish, with small brown spots on the sides.

This is one of the largest species of Python, some specimens being known which measured about 30 feet in length.

As a sample of folk-lore connected with this monstrous snake the following Burmese fable has been recorded by Mason:–^[184]

"According to a Karen legend all the poisonous serpents derive their virulence from the Python, which, though innocuous now, was originally the only one that was venomous. In those days he was perfectly white, but having seduced away a man's wife, Aunt Eu (Eve), he made her, while she was in his den, weave figures on his skin in the forms which are now seen. At that time, if he bit the footstep of a man in the road, such was the virulence of his poison that the man died, how far soever that man might have passed from the bitten track. The Python had not, however, an ocular demonstration of the fact, so he said to the Crow: 'Crow, go and see whether people die or not when I bite the foot-track.' The Crow went to the neighbourhood of a Karen cabin, and found the people, as is their custom at funerals, laughing, singing, dancing, jumping, and beating drums. He therefore returned to the Python, and told him that so far from his efforts producing death, on the contrary they produced joy. The Python was so angry when he heard this that he ascended a tree and spit up all his venom, but other creeping things came and swallowed it, and people die of their malignancy to this day. The tree, therefore, from which the Python spat up his venom became deadly, and its juice is used to this day for the purpose of poisoning arrows. The Python made the other creatures promise not to bite without provocation. The Cobra said: 'If there be transgression so as to dazzle my eyes, to make my tears fall seven times in one day, I will bite.' So said the Tiger (whose bite the Karens esteem as virulent as a serpent's) and others, and they were allowed to retain their poison. But the Water Snake and Frog said they would bite with or without cause as they liked; so the Python drove them into the water, where their poison melted away and their bite became harmless."

P. molurus is the species of India and Ceylon, ranging, however, also into Indo-China. Boulenger quotes W. Elliot^[185] as the authority for the statement that this species grows to the length of 30 feet. Only two pairs of upper labials are pitted. The general colour above is greyish or yellowish brown with a dorsal series of large reddish-brown, black-edged patches, and on the sides of the body with a series of smaller spots with light centres. On the head is a lance-shaped marking; a brown stripe passes from the eye backwards. The under parts are yellowish.

P. sebae and P. regius are African species. The former has two pairs of upper labials pitted, the latter four pairs. P. sebae is generally pale brown above with dark brown, black-edged cross-bars, which are usually connected by a sinuous dark stripe along each side of the back. The upper surface of the tail has a light stripe between two black stripes. The belly is spotted and dotted with dark brown. P. sebae ranges over the whole of Tropical and Southern Africa, perhaps with the exception of Eastern Africa. P. regius of West Africa is beautifully marked, and may be recognised by the dark brown, black-edged band along the back, sending down triangular or Y-shaped processes on the sides, which are pale brown. This dorsal band encloses a light streak on the neck and another on the tail. The belly is yellowish.

These African Pythons grow to a length of about 15 feet, but specimens so large as this are not often met with. The negroes of certain parts of the coast of Guinea are said to worship them and to keep them in special temples, where they are regularly attended to. Their food consists chiefly of small Mammals, notably rats, and of Birds. A couple of these snakes paired in the Zoological Gardens of London in the month of June. The female laid nearly one hundred eggs in the following January, and incubated them until April, when the embryos were found to be still unripe.

Sub-Fam. 2. Boinae.–Without supra-orbital bones. The premaxilla is toothless. The subcaudal scales form mostly a single row.

The Boinae comprise between forty and fifty species. Most of them are American, but the genus Eryx inhabits North Africa, Greece, and South-Western Asia; the genus Enygrus inhabits New Guinea and many of the Pacific Islands, for instance New Britain (Neu Pommern), the Solomon, Loyalty and Fiji Islands, and the New Hebrides. Casarea dussumieri is found on Round Island near Mauritius; and two species of Boa and one of Corallus represent the Boidae in Madagascar, while all the others live in Central and South America.

Boa.–The maxillary and mandibular teeth gradually decrease in size. The scales of the upper parts of the body and tail are smooth and very small. The rostral shield is enlarged. The nostrils are placed between two or three nasals, and these are separated from those of the other side by small scales. The tail is short and prehensile. The pupil is vertical.

B. constrictor, of South America, has the head covered with small scales, one of the pre-oculars being enlarged. The eye is separated from the labials by several series of tiny scales. The general colour is a delicate "pale brown above, with fifteen to twenty dark brown cross-bars widening on each side, and, if connected by a dark dorso-lateral streak, enclosing large elongate oval spots.... On each side is a series of large dark brown spots with light centres, most of which alternate with the cross-bars. On the tail the markings become much larger, brick-red, edged with black, and separated by narrow, yellowish interspaces." Under parts yellowish with black dots. Boa constrictor, a name applied in popular parlance to many species, reaches a length of more than 10 feet; the largest specimen in the British Museum measures exactly 11 feet. A few other species inhabit Central America and the West Indies. B. dumerili and B. madagascariensis, both of Madagascar, cannot be separated from the genus Boa.

A. D. Bartlett^[186] has described the following incident:–

"In the evening of 5th October 1892 two pigeons were put into the cage in which two fine specimens of Boa constrictor had been living on friendly terms since the beginning of the year. The larger snake seized one of the pigeons and the keeper left the house. The next morning only one of the snakes, the larger specimen, was visible, and from its enormously extended body it was evident that it had swallowed its companion, which was about 9 feet in length. It had no longer the power of curling itself round, but remained extended nearly to its full length in a straight line, and appeared to be at least three times its normal circumference. It was almost painful to see the distended skin, which had separated the scales all over the middle of the body. By 2nd November, twenty-eight days later, the snake had not only digested its companion but had regained its appetite as well as its normal size, and it immediately swallowed a pigeon put into its den."

This peculiar case is not one of ordinary cannibalism. It is rather an unintentional accident. When two snakes happen to get hold of the same animal (in the present case a pigeon) and begin to swallow it, the action of swallowing becomes almost mechanical, the snakes continuing to push their jaws over the prey–which in the case of a bird or mammal they cannot taste, nor can they see it–so long as they feel something in the mouth. After the original prey has been mastered, it is the turn of the opposite snake's head, and if the weaker snake does not give way it is swallowed by its stronger mate. Grass-Snakes will swallow several frogs if these are tied together in a string, and other snakes do the same with mice. There are instances on record in which a Python swallowed its blanket, which, being absolutely indigestible, caused its death.

Casarea, the "Round-Island Snake," differs from Boa chiefly by the rough and strongly keeled scales, and by the relatively much longer tail.

Eunectes murinus, the "Anaconda," is an aquatic Boa. It differs from this genus mainly by the inner of the three nasal shields being in contact with that of the other side (see Fig. 159), and by the absence of the little scales between the eye and the labials; the snout is, moreover, covered with shields instead of small scales. The pupil of the eye is normally vertical, but it had contracted into a round pinhole in the dead but still fresh specimen from which the figure was drawn. The general colour is dark olive-brown, with large oval black spots arranged in two more or less alternating rows along the back, and with smaller black, white-eyed spots along the sides. The under parts are whitish, spotted with black. The upper parts of this and of many other dark-coloured species of Boidae are often shiny, with an iridescent lustre.

The Anaconda combines an arboreal with an aquatic life, a kind of existence eminently in harmony with the well-watered, dense forests of Tropical South America, which are the home of this, the largest of all modern Snakes. It is said to attain a length of as much as 33 feet. There is no inherent impossibility in such statements, but the giant specimens seem to have a knack of keeping out of the naturalist's way.

The Anaconda feeds chiefly upon Birds and Mammals, which it catches either on land, mostly during the night-time, or in the water. For the latter purpose it lies submerged in the rivers or floats about leisurely, only the head being above the surface, and anything suitable is attacked. In other localities the snake, if so inclined, establishes itself upon the branches of a tree which overhangs the water, or the track of the game. These aquatic Snakes seem to be viviparous.

Eryx has the head not distinct from the neck and covered entirely with small scales. Those of the body are likewise small, and are either smooth or keeled. The tail is very short. The anterior maxillary and mandibular teeth are longer than the posterior teeth. These snakes, most of which are less than 3 feet in length, inhabit the sandy districts of North Africa, Arabia, and South-Western Asia, extending into Central Asia. One species, E. jaculus, extends into Greece and the Ionian Islands. Like the other species it is an ugly creature, pale grey or yellowish above, with darker patches and spots. The under parts are whitish. The scales are smooth on the front half of the body, becoming keeled further back and on the tail. Total length under 2 feet. The pupil is vertical.

According to Zander^[187] and Werner^[188] this snake lives in sandy localities, digging itself into the sand, or covering the body lightly with sand and leaving only the eyes and nostrils free. The whole body is very flabby, and presses itself into any irregularity of the ground over which the snake creeps. Some specimens live on lizards, others prefer mice. The prey is caught by the head, and further secured by several turns of the body of the captor, whose tail is then turned forwards, round the head of the victim, so as to form a kind of knot.

Not less striking than their agility is their jealousy, which is so strong that a snake will occasionally leave the mouse which it has just strangled in order to seize another snake's mouse. Sometimes several snakes fight for the same mouse, coiled together into one inextricable lump so that the mouse itself is quite invisible. The snakes poke their heads about in search of the hidden prey, and every attempt of one of the snakes to free itself, causes the others to squeeze it firmer and firmer, thinking apparently that the motion was caused by the lost prey.

Occasionally one of Werner's captives caught several mice in succession. With these it crawled into a corner, dropped the mice, and then proceeded quietly to swallow one after another. After a fortnight the whole repast was digested, and the snake was ready for more.

Fam. 6. Xenopeltidae.–The single species, Xenopeltis unicolor, of South-Eastern Asia, including the Malay Islands, has been raised to the dignity of family-rank on account of the following combination of characters. The prefrontal bones are still in contact with the nasals as in the previous families, but the coronoid bones of the mandibles are absent as in the remaining families. The whole suspensorial apparatus and the lower jaw itself are peculiar. The dentary bone is movably attached to the end of the much-elongated articular bone, the movability being enhanced by the absence of the coronoid element.^[189] The quadrate is short and thick, and is carried by the short and broad squamosal, which lies flat against the skull, resembling in this respect that of some of the Ilysiidae. Boulenger rightly considers Xenopeltis to be in various ways intermediate between this family, the Boidae and the Colubridae. The head is small and not distinct from the neck. The eyes are small and have a vertical pupil. The body is cylindrical, covered above with smooth black or brown and highly iridescent scales, hence the generic name. The ventral scales are white and transversely enlarged as in the majority of snakes. The tail is short, but not stunted, measuring about 4 inches in full-grown specimens of a total length of 3 feet.

Fam. 7. Colubridae.–This family comprises those snakes (about nine-tenths of all recent species) which combine the following characters:–ectopterygoids are present: the squamosals are loosely attached to the skull, and carry the quadrates, which are not reached by the pterygoids: the prefrontals are not in contact with the nasals: the maxillaries are horizontal and form the greater portion of the upper jaws: the mandibles lack the coronoid process or element: both jaws are toothed.

The best arrangement of this enormous cosmopolitan family with terrestrial, arboreal, and aquatic forms, is that by Boulenger, who, adopting Duméril's terms, has divided them into three parallel series.

A. Aglypha.–All the teeth are solid and not grooved.

B. Opisthoglypha.–One or more of the posterior maxillary teeth are grooved.

C. Proteroglypha.–The anterior maxillary teeth are grooved or "perforated."

The Aglypha are harmless, non-poisonous. Most of the Opisthoglypha are poisonous, although few of them are dangerously so. The Proteroglypha, which comprise the "Cobras" and their allies, are deadly poisonous.

Series A. AGLYPHA.

Sub-Fam. 1. Acrochordinae.–The postfrontal bones, besides bordering the orbits posteriorly, are extended forwards so as to form the upper border of the orbits, separating the latter from the frontals. The few genera and species of this sub-family are mostly aquatic, inhabiting rivers, or estuaries with brackish water, and they have been known to swim far out into the sea. The body is covered with small, frequently granular scales; in the typically aquatic forms the body is slightly compressed laterally, and the ventral scales are scarcely larger than the others. Most of these ugly snakes inhabit the rivers of coasts of South-Eastern Asia and Papuasia; one, Stoliczkaia, is found in the Khasia Hills of North-Eastern India; another, Nothopsis, lives far from its supposed allies, on the Isthmus of Darien, Central America.

Acrochordus javanicus has no ventral shields. The head is flat, covered with small granules, with the eyes and nostrils on the upper surface. The general colour is dull olive-brown, lighter and spotted beneath. The food consists of fishes. Total length up to 4 feet.

Chersydrus granulatus ranges from the coast of Madras to New Guinea. The body and tail are compressed, and form a ventral fold, covered with tiny scales like the rest of the body. General colour grey above, yellow below.

Sub-Fam. 2. Colubrinae.–The postfrontal bones are restricted to the posterior border of the orbits. The maxillary and dentary bones carry teeth on their whole length. The scales are usually imbricating. This sub-family contains the overwhelming majority of snakes, about 1000 species, all of them harmless so far as poison is concerned. None of them reach a great size, species of 6 or 7 feet in length being rare, e.g. Zamenis mucosus, but a few species of the Indian genus Zaocys s. Coryphodon grow to 10 feet. Most of the Colubrine snakes are oviparous, but some, e.g. Coronella, are viviparous. Some are aquatic, or semi-aquatic, others are absolutely arboreal, others again prefer dry, sandy, or rocky localities, according to their food. The distribution of the sub-family is cosmopolitan, finding its natural limits only in the permanently frozen under-ground, a condition which makes hibernation impossible. Most of them love warmth and like to bask, although many are not fond of the broiling sun. In the temperate regions they hibernate. As a rule they are intelligent and some of them become even affectionate.

Tropidonotus.–The teeth form closely set series on the whole length of the maxillaries, palatines, pterygoids, and the greater portion of the dentaries. The premaxilla is toothless. The teeth of the maxillaries gradually increase in length, the posterior teeth being the longest. The pupil is round. There is a pair of internasal shields. The scales covering the body have each an apical, sensory pit, are mostly keeled, and are arranged in longitudinal series. The ventral shields are broad; the sub-caudals form two rows. This genus, with more than seventy species, has a wide range, practically over the whole world with the exception of New Zealand and the southern half of Australia.

T. natrix, the common Grass-Snake, has a divided, or double, anal shield. The strongly keeled scales of the body form nineteen rows. There are normally seven upper labials, the third and fourth of which border the eye. The usual colour of the Grass-Snake is olive-grey or brown above, with black spots and narrow cross-bands. The labials are white or yellowish, with black sutures. The belly is checkered black and white, more or less suffused with grey. There are several colour-varieties. The typical or northern form has a white, yellow, or orange collar, bordered behind by a black collar; the pale collar is sometimes faint or absent. The second variety, rather common in Spain and Portugal, although not the only form in the Peninsula, has no collar whatever, and these specimens are sometimes almost uniformly grey-green above. The third variety, common in South-Eastern Europe and in Asia Minor, has a well-marked collar and a yellowish streak along each side of the back. But there are also almost black specimens.

The usual length of an adult female Grass-Snake is about 3 feet, but very exceptional cases of more than 6 feet are on record; the males are smaller and more slenderly built. The range extends over the whole of Middle Europe, Algeria, West and Central Asia. It does not, however, occur in Ireland or Scotland. Its northern limit is the southern part of Sweden.

The Grass-Snake prefers moist, grassy localities, with the neighbourhood of water, chiefly on account of the food, which consists entirely of fishes and Amphibia, notably of frogs; tree-frogs are preferred to anything else; toads are occasionally eaten, but mice are never taken.

The Grass-Snake can climb trees or rather shrubs and is an accomplished swimmer, often spending much of its time in water for fishing purposes. The fish is caught by the belly and then generally swallowed on land. The Grass-Snakes appear in the spring and disappear in the autumn to hibernate in the ground. They pair, in England, in the month of May or June, usually on warm and sunny mornings. The eggs are laid from July to the end of August, mostly in rich vegetable soil, in heaps of weeds or in manure-heaps. Young snakes lay fewer eggs than old specimens, which sometimes produce more than three dozen at a time. The eggs are soft, whitish yellow, about one inch long, and soon stick together, so that the whole clump can be taken up at once. As a rule the new-laid eggs do not contain any visible sign of the embryo, but it often happens that the snake has to delay oviposition, and then the embryos are more or less advanced. This is especially the case with recently caught specimens. The young are hatched in the late summer or in the autumn, and seem to live at first upon soft insects and worms. Curiously enough they are easily drowned when they fall into the water, even in a shallow tank. My tame snakes have often laid eggs between the stones in the greenhouse; the young throve well upon unknown food, but most of them met their fate in the water. When they are a few weeks old they are strong enough to take baby-frogs.

The Grass-Snake becomes very tame, learns to distinguish between different people, allows itself to be handled without hissing or without voiding the obnoxiously smelling contents of its cloaca and anal glands, will in time take the offered food from the hand, and will even crawl up the arm or sleeve and coil itself up contentedly. One of the finest specimens, quite green, without a trace of a collar, and with brownish-red eyes, I caught in the Guadiana, where it had been fishing in midstream. It swam towards the bank, dived, and hid itself at the bottom between rocks. This snake, a female, became very tame. It never hibernated, shed its skin regularly every few months, and grew within nine years from 35 inches to 42 inches in length.

The Grass-Snake is perfectly harmless: although hissing, and striking out furiously with its head, it never bites, not even when it is severely handled. Its only defence consists of the awful contents of the cloaca and the anal glands, the secretion of which smells of concentrated essence of garlic mixed with other indescribable odours. The wildest specimens I have ever met with inhabited a swamp with a little stream to the north of Oporto close to the coast. To my utter surprise some of them actually made for me, swimming along rapidly with the head erect, about 6 inches above the water, and darting forwards with widely opened jaws, but they did not bite. These and other kinds of allied snakes require to drink much and often. Occasionally they drink milk when this is offered them, but that they suck the udders of cows or the breasts of women is an idle fable.

T. viperinus.–The scales are strongly keeled and form twenty-one to twenty-three longitudinal rows. The third and fourth labials border the eye. The anal shield is divided. The eyes and nostrils are directed upwards instead of sidewards, in adaptation to the essentially aquatic habits of this species, which lives upon fishes and Amphibia. The general colour is grey to reddish brown, with a black zigzag band along the back and a lateral series of black, yellow-eyed spots. The belly is yellow or red, checkered with black.

The Viperine Snake bears a general resemblance to the common viper. It inhabits France, Italy, Spain and Portugal, and Morocco. Very large specimens attain a length of nearly 3 feet, but the ordinary size of adults is 2 feet. This snake spends most of its time in the water, but it is often found on land, basking on the top of a low wall or on a low shrub. It is exceedingly common in Spain and Portugal, where it inhabits almost every ditch, any standing water or slow river. In the Alemtejo, when during the rainless and hot summer the small rivers have nearly dried up, these snakes collect in great quantities in the remaining stagnant and muddy pools, and as the stock of suitable fish gets exhausted, are often reduced to a deplorably emaciated condition. By the month of August they have become so thoroughly aquatic that they cannot be kept alive in dry surroundings for twenty-four hours. Those which I collected generally died, apparently from some kind of cutaneous suffocation, during the night following their capture. Taken under other conditions they are very easily kept and tamed.

I once caught a Viperine Snake in a ditch whilst it was swallowing an eel of nearly its own length. Both were separated, and then put into a small bag together with other creatures, and no more attention was paid to them for several hours. When I opened the bag again, the snake, undisturbed by my incessant walking about, was again busily engaged in trying to get outside that same eel!

T. sirtalis (Fig. 160) is one of the almost endless varieties of what is now known by the name T. ordinatus, of North and Central America.

T. tesselatus is closely allied to T. viperinus, which it represents in South Germany, Italy, South-Eastern Europe, and Asia; but the scales form only nineteen rows, and the fourth, or fourth and fifth labials, border the eye. The usual colour is olive-grey with dark little spots, and with a dark chevron-shaped band behind the occiput. The lower parts are yellow or red checkered with black, hence the specific name.

Zamenis.–The maxillary teeth are not closely packed; they increase slightly in size backwards, and the last two are often a little larger and separated from the rest by a diastema. The mandibular teeth rather decrease in size from before backwards, inversely with the upper teeth. The scales are smooth with apical pits; the sub-caudals form two rows. The eye is large, and has a round pupil. The range of this genus, with about thirty species, extends over the whole of the Periarctic region.

Z. (Ptyas) mucosus (Fig. 161), the Rat Snake of India, extending from Transcaspia to Java, is a very common species, often seen in menageries. Its general colour is brown above, often with black cross-bands on the hinder part of the body and tail. The under parts are yellowish. The fourth and fifth labials border the eye. The scales on the body form only seventeen rows. Another feature of this species is the prominent ridge of the back-bone, not only in half-starved but in well-conditioned specimens. The Rat Snake grows to a length of more than 7 feet, and is as ill-tempered as most species of this genus.